Species Card of Copepod
Calanoida ( Order )
    Clausocalanoidea ( Superfamily )
        Aetideidae ( Family )
            Gaetanus ( Genus )
Gaetanus kruppii  Giesbrecht, 1903   (F,M)
Syn.: Gaetanus major Wolfenden, 1903 (in Fowler, 1903 (p.125); 1904 (p.114, figs.F, Rem.M); 1908 (p.31); Farran, 1908 b (p.36); Wolfenden, 1911 (p.231, Rem.); Farran, 1926 (p.249, Rem.); Lysholm & Nordgaard, 1921 (p.14);
? G. antarcticus Brady, 1918 (p.19, figs.F);
Gaetanus microcanthus C.B. Wilson, 1950 (p.231, figs.F,M); Björnberg, 1973 (p.323, 386);
no G. kruppii : Sewell, 1947 (p.62, figs.M)
Ref.:
Giesbrecht, 1903 (p.202, figs.F,M); Wolfenden, 1908 (p.32, Rem.); A. Scott, 1909 (p.48, figs.F, Rem.F,M); With, 1915 (p.97, figs.F,M); Sars, 1925 (p.61, figs.F,M); Sewell, 1929 (p.102); Rose, 1929 (p.19); Wilson, 1932 a (p.52, figs.F,M); Rose, 1933 a (p.99, figs.F,M); Jespersen, 1934 (p.62); 1940 (p.22); Lysholm & al., 1945 (p.17); Sewell, 1947 (p.58, 62, 64, Rem.F, non M, non figs.M, Rem.); C.B. Wilson, 1950 (p.231); Vervoort, 1952 e (n°46, p.3, figs.F,M); Tanaka, 1957 b (p.169, figs.F,M); Vervoort, 1963 b (p.126, Rem.); Mazza, 1965 a (p.285, figs.juv., F,M); 1966 (p.54: figs.F,M, p.55: Rem.); 1967 (p.123, 139, 367, 387, figs.juv., F,M); Owre & Foyo, 1967 (p.44, figs.F,M); Mazza, 1968 (p.533, figs.); Tanaka, 1969 (p.261: Rem., fig.4i: F); Tanaka & Omori, 1970 (p.135, figs.M, Rem.F); Razouls, 1972 (p.94, Annexe: p.43, figs.F); Park, 1975 a (p.24, figs.M); 1978 (p.136, figs.F); Bradford & Jillett, 1980 (p.48, figs.F,M, Rem.M, as kruppi); Markhaseva, 1996 (p.196, figs.F,M); Chihara & Murano, 1997 (p.687, Pl.43,44: F,M); Bradford-Grieve & al., 1999 (p.879, 922, figs.F,M); Yamaguchi & al., 2005 (p.835, Tabl. VIII); Vives & Shmeleva, 2007 (p.577, figs.F,M, Rem.); Park & Ferrari, 2009 (p.143, Appendix 1)
Species Gaetanus kruppii - Plate 1 of morphological figuresissued from : Tanaka O. in Publ. Seto Mar. Biol. Lab., 1957, 6 (2). [Fig.41, p.170].
Female: habitus (dorsal aspect); b, head (lateral aspect); c, last thoracic segment and urosome (lateral aspect); d, exopodite of P1.
Male: e, head (lateral aspect); f, last thoracic segment (lateral aspect); g, P5.


Species Gaetanus kruppii - Plate 2 of morphological figuresissued from : T. Park in Antarctic Res. Ser. Washington, 1978, 27. [p.137, Fig.21].
Female: A, habitus (dorsal view); B, idem (right lateral side); C, forehead (dorsal view); D, idem (right lateral side); E, posterior part of metasome and urosome (rignt lateral side); F, idem (dorsal view); G, A2.


Species Gaetanus kruppii - Plate 3 of morphological figuresissued from : T. Park in Antarctic Res. Ser. Washington, 1978, 27. [p.138, Fig.22].
Female: A, Mxp; B, P1; C, P2; D, P3; E, medial part of coxa of P4 (posterior). P1-3: legs (anterior view).


Species Gaetanus kruppii - Plate 4 of morphological figuresissued from : E.L. Markhaseva in Trudy Zool. Inst. RAN, St. Petersburg, 1996, 268. [p.200, Fig.157].
Female (from different specimens of Izu-Bonin Trench and SE Pacif.). Ce: forehead (lateral and dorsal).


Species Gaetanus kruppii - Plate 5 of morphological figuresissued from : E.L. Markhaseva in Trudy Zool. Inst. RAN, St. Petersburg, 1996, 268. [p.201, Fig.158].
Male (from SE Pacif.)


Species Gaetanus kruppii - Plate 6 of morphological figuresissued from : T. Park in Bull. Mar. Sci., 1975, 25 (1). [p.24, Fig.8].
Male: A, B, forehead (showing a variability of the cephalic process of rostrum); C, posterior part of metasome and urosome (lateral); D, A2; F, fifth pair of legs (posterior); F, last two exopodal segments of left fifth leg (medial).
Note: In habitus, the male closely resembles that of G. pileatus. See: left fifth leg ( length of the endopod and characteristic third exopodal segment) and presence of setae on the second exopodal segment of A2


Species Gaetanus kruppii - Plate 7 of morphological figuresissued from : J.M. Bradford & J.B. Jillett in Mem. N.Z. Oceanogr. Inst., 86, 1980. [p.50, Fig.32]. As .
Female: A, habitus (dorsal); B, idem (lateral right side); C, basipod 1 of Mxp; D, exopod of P1; E, inner border of basipod 1 of P4.
Nota: The south-west specimens have long posterior metasomal points and a metasome with a length less than three times its width.

Male: F, forehead (lateral); G, posterior of thorax and 1st urosomal segment (dorsal); H, P5 (r = right; l = left).
Nota: Right P5 endopod not reaching beyond left leg endopod, left leg endopod about two thirds length of left leg exopod joint 1 and may be two-jointed.


Species Gaetanus kruppii - Plate 8 of morphological figuresIssued from: G.O. Sars in Résult. Camp. Scient. Prince Albert I, 69, pls.1-127 (1924). [Pl.XVIII, figs.5-8].
Female: 5, habitus (dorsal); 6, idem (lateral left side).

Male: 7, forehead (lateral); 8, P5.


Species Gaetanus kruppii - Plate 9 of morphological figuresissued from : R.B.S. Sewell in The John Murray Expedition, 1933-34, Scientific Reports, VIII (1), 1947. [p.63, Fig.10, B].
Female (from Arabian Sea): B, A2.


Species Gaetanus kruppii - Plate 10 of morphological figuresissued from : A. Scott in Siboga-Expedition, 1909, XIX a. [Plate IX, Figs.9-15].
Female (from Indonesia-Malaysia): 9, habitus (dorsal); 10, forehead (lateral); 11, last thoracic and genital segments (left side); 12, rostrum; 13, Mxp (basal portion); 14, P1; 15, P4 (basal portion).


Species Gaetanus kruppii - Plate 11 of morphological figuresissued from : A. Scott in Siboga-Expedition, 1909, XIX a. [Plate X, Figs.1-9].
Male (from Indonesia-Malaysia): 1, habitus (dorsal); 2, forehead (lateral); 3, last thoracic and genital segments (left side); 4, rostrum; 5, Mxp; 6, P1; 7, P2; 8, P4 (basal portion); 9, P5.


Species Gaetanus kruppii - Plate 12 of morphological figuresissued from : C. Razouls in Th. Doc. Etat Fac. Sc. Paris VI, 1972, Annexe. [Fig.43, A, B].
Female (from Banyuls, G. of Lion): A, thoracic segment 5 and urosome (lateral); B, forehead (lateral).


Species Gaetanus kruppii - Plate 13 of morphological figuresissued from : C.B. Wilson in Bull. U.S. natn. Mus., 1950, 100, 14 (4). [Pl. 11, Figs.114-116]. As Gaetanus microcanthus.
Female (from 11°30'S, 87°19'W): 114-115, habitus (dorsal and lateral, respectively).

Male: 116, P5.


Species Gaetanus kruppii - Plate 14 of morphological figuresissued from : C.B. Wilson in Bull. U.S. natn. Mus., 1950, 100, 14 (4). [Pl. 19, Fig.246]. As Gaetanus microcanthus.
Female: 246, basipod of P4.


Species Gaetanus kruppii - Plate 15 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.44, Fig.243].
Female (from 23°38'N, 83°06'W): 243, habitus (lateral).


Species Gaetanus kruppii - Plate 16 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.26, Figs.119].
Female: forehead (lateral).


Species Gaetanus kruppii - Plate 17 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.29, Fig.134].
Male: 134, P5.

Compl. Ref.:
Wilson, 1942 a (p.187); Massuti Alzamora, 1942 (p.110); Sewell, 1948 (p.330, 500, 508, 520, 529, 531, 534, 539, 545, 556, 566); C.B. Wilson, 1950 (p.231); Fagetti, 1962 (p.18); V.N. Greze, 1963 a (tabl.2); Grice, 1963 a (p.495); Gaudy, 1963 (p.22, Rem.); De Decker & Mombeck, 1964 (p.12); Mazza, 1966 (p.70); Grice & Hulsemann, 1967 (p.15); 1968 (tab.2); Vinogradov, 1968 (1970) (p.268); Evans, 1968 (p.13); Roe, 1972 (p.277, tabl.1, tabl.2); Björnberg, 1973 (p.323, 386); Vives & al., 1975 (tab.II); Gaudy, 1975 (p.109, fig.1, Table I, 4, respiration); Deevey & Brooks, 1977 (p.256, tab.: as kruppi<*em>, Station "S"); Vaissière & Séguin, 1980 (p.23, tab.2); Kovalev & Schmeleva, 1982 (p.83); Vives, 1982 (p.291); Scotto di Carlo & Ianora, 1983 (p.150); Scotto di Carlo & al., 1984 (p.1042); Lozano Soldevilla & al., 1988 (p.58); Pancucci-Papadopoulou & al., 1990 (p.199); Scotto di Carlo & al., 1991 (p.270); Suarez Gasca, 1991 (tab.2); Suarez, 1992 (App.1); Suarez-Morales & Gasca, 1998 a (p108); Lapernat, 2000 (tabl. 3, 4); Razouls & al., 2000 (p.343, Appendix); Holmes, 2001 (p.48); Sameoto & al., 2002 (p.12); Fernandes, 2008 (p.465, Tabl.2); Galbraith, 2009 (pers. comm.); Licandro & Icardi, 2009 (p.17, Table 4); Hidalgo & al., 2010 (p.2089, Table 2); Mazzocchi & Di Capua, 2010 (p.423); Medellin-Mora & Navas S., 2010 (p.265, Tab. 2)
NZ: 21

Distribution map of Gaetanus kruppii by geographical zones
Loc:
sub-Antarct. (Pacif. SE), Afr. S (E), off Tristan da Cunha E, G. de Guinée, off I. St. Paul E, off Is. du Cap Vert NW, off Maroc-Mauritanie, Canaries, off Madère, off Portugal, Açores, Brésil, Caribbean Colombia, G. du Mexique, Floride, Mer des Sargasses, Station "S" (32°10'N, 64°30'W), off Woods Hole, Nouvelle-Ecosse S, off Irlande S & W, G. de Gascogne, Mer du Nord, Détr. de Davis S, Islande, Féroé, Baie Ibéro-marocaine, Médit. (Mer d'Alboran, Banyuls, Marseille, Ligurian Sea, Tyrrhenian Sea, Strait of Messina, Malte, Ionienne, bassin Levantin), G. d'Aden, Mer Arabe, off Ceylan SW, Madagascar (Nosy Bé), Indien, Bay of Bengal, Indonésie-Malaisie, Philippines, Japon (Izu), Fosse des Mariannes, Kouriles-Kamtchatka, Mer de Béring, Vancouver Is., Nouvelle-Zélande, Pacif. (tropical & central N), Is. Marquises, Hawaii, Californie, off Galapagos, off Pérou, off Is. Juan Fernandez, Chili (., N)
N: 70
Lg.:
(1) F: 5,2; M: 4,1; (2): F: 4-3,6; M: 3,7; (5) F: 5,7; (7) F: 5,4; M: 5,04; (10) F: 5,6-5; (14) F: 5,6-4,7; M: 4,4-4,2; (20) F: 5,66-5,16 (32) F: 4-3,6; M: 3,7; (37) F: 5,7-4,7; M: 5,2-4,5; (38) F: 4,8-4,45; (40) F: 5,25-4,83; M: 4,91-4,32; (45) F: 4,5-3,6; M: 5-3,7; (56) F: 4,95; M: 4,87; (113) F: 5,3-4,7; M: 4,9-4,5; (199) 5,28-4,1; M: 4,67; (201) F: 5,4-5; M: 4,6; (207) F: 4,56-3,7; M: 4,04; (260) F: 4,56-3,7; M: 4,04; (866) M: 3,7-5,6; {F: 3,60-5,70; M: 3,70-5,60}
Rem.: bathypelagic. Sargasso Sea: 500-1500 m (Deevey & Brooks, 1977, station "S");
Sampling depth (sub-Antarct.) : 0-800-2000 m.
Vervoort (1963 b, p.127) follows Sewell (1947, p.62) that G. clarus Esterly (1906, p.57) is the male of G. kruppii, and have no doubt that G. major Wolfenden (1904) is conspecific with G. kruppi, inclining not to attach too much importance to the fact that in Wolenden's desciptions asserted that the lamella is present on the coxa of Mxp; the thin lamella is easily ruptured during dissection or is easily folds under the coxa and becomes more or less invisible.Furthermore, the greatly increased interest in Atlantic deep water copepods would almost certainty have turned up a species of Gaetanus of the G. kruppii type without a lamella if Wolfenden's G. major were a species distinct from G. kruppii.
R. Stephen: Data sheets of NIO, Kochi, India (on line).
Last update : 15/03/2013
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