Species Card of Copepod
Calanoida ( Order )
    Arietelloidea ( Superfamily )
        Augaptilidae ( Family )
            Euaugaptilus ( Genus )
Euaugaptilus hecticus  (Giesbrecht, 1889)   (F,M)
Syn.: Augaptilus hecticus Giesbrecht,1889; 1892 (p.400, 414, 770, figs.F,M); Giesbrecht & Schmeil, 1898 (p.122, Rem. F,M); I.C. Thompson, 1903 a (p.29); A. Scott, 1909 (p.136, Rem.); Wolfenden, 1911 (p.339); Rose, 1924 d (p.481); Lysholm & al., 1945 (p.37, Rem.)
Ref.:
Farran, 1926 (p.288); 1929 (p.209, 269); Sewell, 1932 (p.323); Rose, 1933 a (p.231, figs.F,M); C.B. Wilson, 1950 (p.205, figs.F); Grice, 1962 (p.226, figs.F); Tanaka, 1964 b (p.71, figs.F,M); Owre & Foyo, 1967 (p.86, figs.F,M); Matthews, 1972 (p.32); Tanaka & Omori, 1974 (p.200, 217, 199: fig.F); Björnberg & al., 1981 (p.653, figs.F,M); Ali-Khan & Ali-Khan, 1984 (p.304, figs.F,M); Mauchline, 1998 (p.68: F); Bradford-Grieve & al., 1999 (p.882, 940, figs.F,M); Bradford-Grieve,1999 b (p.51, figs.F,M, Rem., figs.171, 190); Conway & al., 2003 (p.83, figs F,M, Rem.); Vives & Shmeleva, 2007 (p.199, figs.F,M, Rem.); Matsuura & al., 2010 (p.2098, Table 4, fig.10: Mx2, Mxp)
Species Euaugaptilus hecticus - Plate 1 of morphological figuresissued from : O. Tanaka in Publs Seto Mar. Biol. Lab., 1964, XII (1). [p.71, Fig.209].
Female: a, habitus (dorsal); b, Mx1; c, P5, d, P5 from immature specimen.
Nota: The urosome segments and furca are in the proportional lengths as 54:13:11:22 = 100. A1 exceeds the end of the furca by last 5 or 6 segments

Male: e, habitus (dorsal); f, right P5; g, left P5. Nota: The urosome segments and furca are in the proportional lengths as 20:17:14:11:16:22 = 100. The furcal rami 3 times as long as wide. The left A1 exceeds the end of the furca by distal 3 segments. Grasping A1 on the right.


Species Euaugaptilus hecticus - Plate 2 of morphological figuresissued from : J.M. Bradford-Grieve in The Marine Fauna of New Zealand: Pelagic Calanoid Copepoda. National Institute of Water and Atmospheric Research (NIWA). NIWA Biodiversity Memoir, 111, 1999. [p.52, Fig.24].
Female: A, habitus (dorsal); B, Mx1; C, P5.


Species Euaugaptilus hecticus - Plate 3 of morphological figuresissued from : O. Tanaka & M. Omori in Publs Seto Mar. Biol. Lab., 1974, XXI (3/4). [p.199, Fig.2,a].
Female: a, Md (mandibular teeth).


Species Euaugaptilus hecticus - Plate 4 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.39, Fig.45]. As Augaptilus hecticus.
Female: 45, urosome (ventral).


Species Euaugaptilus hecticus - Plate 5 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.27, Fig.30]. As Augaptilus hecticus.
Female: 30, P5 (anterior view).


Species Euaugaptilus hecticus - Plate 6 of morphological figuresissued from : C.B. Wilson in U.S. National Museum. Bull.100, 14 (4), 1950. [Pl.21, Fig.299].
Female: 299, P5.


Species Euaugaptilus hecticus - Plate 7 of morphological figuresissued from : S. Ali-Khan & J. Ali-Khan in Crustaceana, 1984, 47 (3). [p.304, Figs.7-12].
Female (from 24°09'N, 64°27'E): 7, habitus (dorsal); 8, Md (mandibular blade); 9, Mx1; 10, P5.

Male: 11, last thoracic segmen and urosome (dorsal); 12, P5.


Species Euaugaptilus hecticus - Plate 8 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. - Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.29 , Fig.18 ]. As Augaptilus hecticus.
Male: 18, P5.


Species Euaugaptilus hecticus - Plate 9 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. - Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.28 , Fig.9 ]. As Augaptilus hecticus.
Male: 9, A2 (anterior view).


Species Euaugaptilus hecticus - Plate 10 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. - Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.28 , Fig.16 ]. As Augaptilus hecticus.
Male: 16, Md.


Species Euaugaptilus hecticus - Plate 11 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. - Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.28 , Figs.33, 37 ]. As Augaptilus hecticus.
Male: 33, Mxp (posterior view); 37, Mx1 (posterior view).


Species Euaugaptilus hecticus - Plate 12 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. - Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.28 , Fig.5 ]. As Augaptilus hecticus.
Female: 5, A1 (distal segments):


Species Euaugaptilus hecticus - Plate 13 of morphological figuresissued from : G.D. Grice in Fish. Bull. Fish and Wildl. Ser., 1962, 61. [p.225, Pl.26, Figs.15-17].
Female (from equatorial Pacific): 15, urosome (dorsal); 16, Md (biting edge); 17, P5.


Species Euaugaptilus hecticus - Plate 14 of morphological figuresissued from : H. Matsuura, S. Nishida & J. Nishikawa in Deep-Sea Res. II, 2010, 57. [p.2106, Fig.10, c-d].
Female (from Sulu Sea): c, SEM images of button setae on Mx2 and Mxp ; d, mandible blade.
Nota : Blade is like a curved needle with 2 spiniform teeth (Type-1).


Species Euaugaptilus hecticus - Plate 15 of morphological figuresissued from : J.M. Bradford-Grieve, E.L. Markhaseva & C.E.F. Rocha & B. Abiahy in South Atlantic Zooplankton, Edit. D. Boltovskoy. 1999. Vol.2. Copepoda. [p.1038, Fig. 7.251].
Female: Mx1.

Nota: Exopod of Mx1 with 3 + 2 setae; Outer lobe 1 with 3 S; inner lobe with 4 setae.

Compl. Ref.:
Sewell, 1948 (p.349, 504, 509, 516, 522, 547, 558); Björnberg, 1963 (p.54, Rem.); Grice, 1963 a (p.496); V.N. Greze, 1963 a (tabl.2); Shmeleva, 1964 a (p.1068); Grice & Hulsemann, 1965 (p.224); Mazza, 1966 (p.72); Pavlova, 1966 (p.44); Fleminger, 1967 a (tabl.1); Delalo, 1968 (p.138); Park, 1970 (p.477); Timonin, 1971 (p.281, trophic group); Deevey, 1971 (p.224); Roe, 1972 (p.277, tabl.1, tabl.2); Deevey & Brooks, 1977 (p.156, tab.2, Station "S"); Carter, 1977 (1978) (p.36); Vaissière & Séguin, 1980 (p.23, tab.2); Vives, 1982 (p.294); Kovalev & Shmeleva, 1982 (p.85); Scotto di Carlo & al., 1984 (1043); Cummings, 1984 (p.163, Table 2); Guangshan & Honglin, 1984 (p.118, tab.); Brinton & al., 1986 (p.228, Table 1); Madhupratap & Haridas, 1986 (p.105, tab.1); Lozano Soldevilla & al., 1988 (p.60); Suarez & al., 1990 (tab.2); Scotto di Carlo & al., 1991 (p.271); Suarez & Gasca, 1991 (tab.2); Suarez, 1992 (App.1); Shih & Young,1995 (p.68); Hure & Krsinic, 1998 (p.102); Suarez-Morales & Gasca, 1998 a (p108); Lopez-Salgado & al., 2000 (tab.1); Weikert & al., 2001 (p.227, tab.4, Rem.); Holmes, 2001 (p.11); Vukanic, 2003 (p.139, tab.1); Hsiao & al., 2004 (p.325, tab.1); Lo & al., 2004 (p.89, tab.1); Kasmi, 2004 (p.227); Dur & al., 2007 (p.197, Table IV); Fernandes, 2008 (p.465, Tabl.2); Williamson & McGowan, 2010 (p.273, Table 3, Pacific central gyres: N and S); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical/tropical); Matsuura & al., 2010 (p.2098, Table 3, figs.7, 8, vertical distribution); Mazzocchi & Di Capua, 2010 (p.424); Medellin-Mora & Navas S., 2010 (p.265, Tab. 2); Hsiao S.H. & al., 2011 (p.475, Appendix I); Tutasi & al., 2011 (p.791, Table 2, abundance distribution vs La Niña event); in CalCOFI regional list (MDO, Nov. 2013; M. Ohman, comm. pers.); Bonecker & a., 2014 (p.445, Table II: frequency, horizontal & vertical distributions); Benedetti & al., 2016 (p.159, Table I, fig.1, functional characters); Benedetti & al., 2018 (p.1, Fig.2: ecological functional group); Belmonte, 2018 (p.273, Table I: Italian zones)
NZ: 15

Distribution map of Euaugaptilus hecticus by geographical zones
Species Euaugaptilus hecticus - Distribution map 3issued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. 1967. [p.86, Table 38].
Vertical distribution of Euaugaptilus hecticus at the ''40-Mile station'' in the Florida Current (± 25°35'N, 79°27'W).
SL 53: 18 V 1958. A: during midday; B;: during midnight.
Species Euaugaptilus hecticus - Distribution map 4issued from : H. Matsuura, S. Nishida & J. Nishikawa in Deep-Sea Res. II, 2010, 57. [p.2104, Fig.7].
Vertical distribution of Euaugaptilus hecticus in the Sulu Sea, compared with those in the Celebes Sea.
Loc:
South Africa (E), G. of Guinea, off S Cape Verde Is., off SW Ascension Is., S & central Brazil , off Amazon, Venezuela, Caribbean Colombia, Caribbean Sea, G. of Mexico, Florida, Sargasso Sea, off Bermuda (Station "S"), off Canary Is., off Morocco-Mauritania, off Madère, Portugal, Bay of Biscay, off W Ireland , Medit. (Algiers Bay, W Basin, Monaco, Tyrrhenian Sea, Adriatic Sea, Ionian Sea, S Aegean Sea, bassin Lebanon Basin), Red Sea, Mer Arabian Sea, Laccadive Is., Natal, Madagascar, Bay of Bengal, Indonesia-Malaysia, Celebes and Sulu Seas, Philippines, China Seas (East China Sea, South China Sea), Taiwan (S, E, N: Mienhua Canyon), Japan (Izu), Pacif. (W equatorial), New Zealand (off North Island N), Pacif. (equatorial), Galapagos-Ecuador, Pacific (central gyres: N and S), California, Gulf of California, N Easter Is.
N: 73
Lg.:
(35) M: 2,28; (38) F: 2,85; M: 2,4; (46) F: 2,75-2,45; M: 2,4; (68) F: 2,66; (69) F: 2,63; M: 2,28; (101) F: 2,7-2,28; (199) F: 2,74-2,36; M: 2,43-2,28; (237) F: 2,8; (315) F: 1,8-1,6; M: 1,2; (909) F: 2,5; 2,71; (991) F: 2,45-2,75; M: 2,28-2,40; {F: 1,60-2,85; M:1,20-2,43}
Rem.: epi-mesopelagic. Known to occur at depths between 30 and 700 m (Matthews, 1972). Sargasso Sea: 0-1000 m (Deevey & Brooks, 1977, Station "S").
Timonin (1971, p.282) considers the trophic interrelations in the equatorial and tropical Indian Ocean, and divides the plankters into 6 trophic groups from the litterature and the results of studies of mouth-parts structure and intestine content. This species is a piercing and sucking carnivorous.
After Benedetti & al. (2018, p.1, Fig.2) this species belonging to the functional group 2 corresponding to small ambush feeding carnivorous.

See in DVP Conway & al., 2003 (version 1).

In ''affinis"" Group after Matthews (1972, p.64) {See remarks in Euaugaptilus genus}
Last update : 06/08/2021
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