Fiche d'espèce de Copépode
Calanoida ( Ordre )
    Clausocalanoidea ( Superfamille )
        Clausocalanidae ( Famille )
            Clausocalanus ( Genre )
Clausocalanus lividus  Frost & Fleminger, 1968   (F,M)
Syn.: Clausocalanus arcuicornis sp.2 : Fleminger,1964 a; 1967 a (tab. I)
Ref.:
Frost & Fleminger, 1968 (p.35, figs.F,M, Rem.); Carli & Crisafi, 1969 (p.277, figs.F,M); Razouls, 1972 (p.94, Annexe: p.31, 33, figs.F, Rem.); Williams & Wallace, 1975 (p.175, fig.F); Dawson & Knatz, 1980 (p.7, 8, figs.F,M); Alvarez-Marques, 1981 (p.157, figs.F, Rem.); Gardner & Szabo, 1982 (p.182, figs.F,M); Bradford-Grieve, 1994 (p.118, figs.F,M, fig.101); Chihara & Murano, 1997 (p.776, Pl.90,93: F,M); Bradford-Grieve & al., 1999 (p.878, 916, figs.F,M); Bucklin & al., 2003 (p.335, tab.2, fig.1, Biomol); Blanco-Bercial & Alvarez-marqués, 2007 (p.73, fig.2, Biomol.); Avancini & al., 2006 (p.78, Pl. 47, figs.F,M, Rem.); Vives & Shmeleva, 2007 (p.619, figs.F,M, Rem.); Blanco-Bercial & al., 2011 (p.108, fig.3, phylogeographic biomol. analysis)
Espèce Clausocalanus lividus - Planche 1 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull; Scripps Inst. Oceanogr. Univ. California, 1968, 12. [Pl.14, p.128-129; Pl.15, p.130-131; Pl.16, p.132-133].

Female: 1 a, habitus (right lateral view); 1 b, idem (dorsal view); taken from different specimens.2 a, b, frontal region (right lateral view); 2, c-f, rostrum (right lateral view); 2 g, rostrum (anterior view); 2, h-j, Th. 4-5 (posterior part) and genital segment (right lateral view); from different specimens; 3 a, Th.4-5 (posterior part) and genital segment (right lateral view); 3 b, genital segment (ventral view); 3 c, Th.4-5 (posterior part) and urosome with spermatophore attached (right lateral view); 3 d, B2 of P2; 3 e, B2 of P3; 3 f, P5; from different specimens.


Espèce Clausocalanus lividus - Planche 2 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull; Scripps Inst. Oceanogr. Univ. California, 1968, 12. [Pl.17, p.134-135; Pl.18, p.136-137].

Male: 1 a, habitus (right lateral view); 1 b, idem (dorsal view); 1 c, frontal region (right lateral view); 1 d, urosome 4 (posterior part), urosome 5 and caudal rami (armature incomplete) (dorsal view); from different specimens. 2 a, Th.4-5 (posterior part) and urosome (right lateral view); 2 b, urosome (armature of caudal rami incomplete) (dorsal view); 2 c, B2 of P2; 2 d, B2 of P3; 2 e, P5 (right lateral view); 2 f, right P5 (posterior ); 2 g, right P5 (left lateral); from different specimens.
Nota: 2nd segment of P5 more than 4.4 times as long as wide. Urosomal segment 2 more than 1.35 times as long as 2nd segment of P5.


Espèce Clausocalanus lividus - Planche 3 de figures morphologiquesissued from : J.M. Bradford-Grieve in The Marine Fauna of New Zealand: Pelagic Calanoid Copepoda. National Institute of Water and Atmospheric Research (NIWA). New Zealand Oceanographic Institute Memoir, 102, 1994. [p.118, Fig.67].
Female: A, habitus (lateral right side); B, genital somite (lateral right side); C, rostrum (lateral view); D, basipod of P3.

Male: F, habitus (lateral left side); G, basipod 2 of P3; H, P5.


Espèce Clausocalanus lividus - Planche 4 de figures morphologiquesissued from : C. Razouls in Thèse Doc. ès Sciences, Univ. P & M. Curie, Paris , Tome annexe, 1972, Fig.32 (unpublished).
Female (from Banyuls): A, habitus (lateral left side); B, basipod 2 of P2; C, basipod 2 of P3; D, P5; E, anal segment and right furcal ramus (dorsal).


Espèce Clausocalanus lividus - Planche 5 de figures morphologiquesissued from : C. Razouls in Thèse Doc. ès Sciences, Univ. P & M. Curie, Paris , Tome annexe, 1972, Fig.33 (unpublished).
Female (from Banyuls): A-E, rostrum from different specimens (lateral view); F, P5.


Espèce Clausocalanus lividus - Planche 6 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.30, Table 2a].
Clausocalanus lividus females: Measurements and ratios *.


Espèce Clausocalanus lividus - Planche 7 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.31, Table 2b].
Clausocalanus lividus males: Measurements and ratios *.
r = sample range; m = sample mean; n = number of specimens measured; s = sample standard deviation.


Espèce Clausocalanus lividus - Planche 8 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.102, P1.1, b, c].
Female: b, urosome with spermatophore attached (right lateral); c, basipodite 2 of P3.
cm = cementing material; am = accessory material associated with spermatophore neck; sn = spermatophore neck; ss = spermatophore sac; D = proximal width of basipodite 2 spiniform process 3; L = distance between processes 2 and 3.


Espèce Clausocalanus lividus - Planche 9 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.27, Fig.10].
Urosomal segment 2:segment 2 of P5 length ratio (ordinate) plotted against urosomal segment 2 length tabscissa) for males of Clausocalanus lividus and C. mastigophorus.


Espèce Clausocalanus lividus - Planche 10 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.18, Fig.1].
Female caudal ramus length-to-width ratio (ordinate) plotted against caudal ramus length (abscissa) for Clausocalanus mastigophorus and C. lividus.


Espèce Clausocalanus lividus - Planche 11 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.131, Pl.15, c-f, g].
Female: c-f, rostrum (right lateral); g, rostrum (anterior).


Espèce Clausocalanus lividus - Planche 12 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.133, Pl.16, d, e, f].
Female: d, basipodite 2 of P2; e, basipodite 2 of P3; f, P5.


Espèce Clausocalanus lividus - Planche 13 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.137, Pl.18, a, b, e, f-g].
Male: a, posterior part of last thoracic segment and urosome (right lateral); b, urosome (dorsal); e, P5 (right lateral); f-g, right P5 (left lateral).


Espèce Clausocalanus lividus - Planche 14 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.137, Pl.18, c, d].
Male: c, basipodite 2 of P2; d, basipodite 2 of P3.


Espèce Clausocalanus lividus - Planche 15 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.133, Pl.16, a, b].
Female: a, posterior part of last thoracic segment and genital segment (right lateral); b, genital segment (ventral; see legend of structure of seminal receptacle in C. arcuicornis).


Espèce Clausocalanus lividus - Planche 16 de figures morphologiquesissued from : C. Razouls in Th. Doc. Etat Fac. Sc. Paris VI, 1972, Annexe. [Fig.32].
Female (from Banyuls, G. of Lion): A, habitus (lateral); B, basipod 2 of P2; C, basipod 2 of P3; D, P5; E, anal segment and left caudal ramus (dorsal).


Espèce Clausocalanus lividus - Planche 17 de figures morphologiquesissued from : C. Razouls in Th. Doc. Etat Fac. Sc. Paris VI, 1972, Annexe. [Fig.34].
Female: A-E, various forms of rostrum; F, P5.


Espèce Clausocalanus lividus - Planche 18 de figures morphologiquesissued from : C. Razouls in Th. Doc. Etat Fac. Sc. Paris VI, 1972, Annexe. [Table XXXI] (unpublished).
Sizes and ratios in Female (from Banyuls, G. of Lion): Lt= Bodyl length; C= Cephalolome length; P= prosome length; U= urosome length; U1= 1st urosomal segment; U2= 2nd urosomal length; U3= 3rd urosomal segment; F= caudal ramus length; n= number of specimens; moyenne= mean; Cv= coefficient of variation (percentage).


Espèce Clausocalanus lividus - Planche 19 de figures morphologiquesIssued from F. Alvarez-Marques in Rev. Fac. Cienc. Univ. Oviedo (Ser. Biologia), (1979-80), 20-21, 1981. [p.161, Pl. I, Figs.3-5].
Female (from Gijon, NW Spain): 3, habitus (lateral); 4, urosome and spermatheca (lateral); 2, P5.

Ref. compl.:
Björnberg, 1973 (p.310, 385); Carter, 1977 (1978) (p.35); Kovalev & Schmeleva, 1982 (p.83); Scotto di Carlo & Ianora, 1983 (p.150); Scotto di Carlo & al., 1984 (p.1041); Tremblay & Anderson, 1984 (p.6: Rem.); Regner, 1985 (p.11, Rem.: p.27); Brenning, 1985 a (p.28, Table 2); Greze & al., 1985 (p.7); Hure & Krsinic, 1998 (p.100); Pancucci-Papadopoulou & al., 1990 (p.199); Ianora & al., 1990 (p.249, fig., parsitism effects); Scotto di Carlo & al., 1991 (p.270); Seguin & al., 1993 (p.23); Saiz & Calbet, 1999 (p.599, reproduction); Moraitou-Apostolopoulou & al., 2000 (tab.I, fig.6); Fragopoulu & al., 2001 (p.49, tab.1); Mackas & Galbraith, 2002 a (p.423, Table 2); Vukanic, 2003 (139, tab.1); Hsiao & al., 2004 (p.325, tab.1); Lan & al., 2004 (p.332, tab.1); Peralba & Mazzocchi, 2004 (p.645, figs.3,6); Lo & al., 2004 (p.89, tab.1); Vukanic & Vukanic, 2004 (p.9, tab. 2, 3); Fernandez & al., 2004 (p.501, tab.5); Isari & al., 2006 (p.241, tab.II); Hooff & Peterson 2006 (p.2610); Valdés & al., 2007 (p.103: tab.1); Dur & al., 2007 (p.197, Table IV); Khelifi-Touhami & al., 2007 (p.327, Table 1); Cabal & al., 2008 (289, Table 1); Galbraith, 2009 (pers. comm.); Lan & al., 2009 (p.1, Table 2); Brugnano & al., 2010 (fig.8); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical/tropical, Fig.4, 6, 7, Tabe 4); Mazzocchi & Di Capua, 2010 (p.425); Hsiao S.H. & al., 2011 (p.475, Appendix I); Mazzocchi & al., 2011 (p.1163, Table II, long-term time-series 1984-2006); Uysal & Shmeleva, 2012 (p.909, Table I); Miloslavic & al., 2012 (p.165, Table 2, transect distribution); Brugnano & al., 2012 (p.207, Table 3); in CalCOFI regional list (MDO, Nov. 2013; M. Ohman, comm. pers.); Hirai & al., 2013 (p.1, Table I, molecular marker); Peijnenburg & Goetze, 2013 (p.2765, genetic data); Fernandez de Puelles & al., 2014 (p.82, Table 3, seasonal abundance)
NZ: 16

Carte de distribution de Clausocalanus lividus par zones géographiques
Espèce Clausocalanus lividus - Carte de distribution 3issued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [p.36, Chart 2, a].
Occurrence of C. lividus in samples examined; closed circles represent samples examined; open circles represent samples in which adults were found; bars through open circles represent samples from which specimens were removed for measurements.
Espèce Clausocalanus lividus - Carte de distribution 4issued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [p.37, Chart 2, b].
Occurrence of C. lividus in samples examined; closed circles represent samples examined; open circles represent samples in which adults were found; bars through open circles represent samples from which specimens were removed for measurements.
Loc:
China Seas (East China Sea, South China Sea), Taiwan Strait, Taiwan (SW, E, NW, N: Mienhua Canyon), Japan, off SE Japan, Pacif. (subtropical & temperate N & S), Gulf of Alaska (58°N, 147°W), Vancouver Is., Oregon (off Newport), California, New Zealand, Tahiti, Chile, Indian S, E South Africa (Natal), off S Cape Verde Is., G. of Mexico, Florida, New-York, Gulf Stream (41° N,58° W), off S Newfoundland, off W Ireland, Bay of Biscay, Gijon (NW Spain), Cantabrian Sea, Baie Ibéro-marocaine, Medit. (Alboran Sea, Gulf of Annaba, Baleares, Banyuls, Ligurian Sea, Tyrrhenian Sea, Naples, Strait of Messina, Adriatic Sea, Ionian Sea, Aegean Sea, Lebanon Basin)
N: 59
Lg.:
(30) F: 1,77-1,26; M: 1,45-1,13; (374) F: 1,75-1,262; (784) F: 1,98-1,66; (1138): F: 1,79-1,22; M: 1,30-1,09; {F: 1,22-1,98; M: 1,09-1,45}
Rem.: épipélagique.
Voir en annexe (in Razouls, 1972) la comparaison entre les femelles de cette espèce et Clausocalnus mastigophorus, notamment la variabilité du rostre (figs.)
Voir aussi les remarques en anglais
Dernière mise à jour : 20/09/2014

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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2014. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Disponible sur http://copepodes.obs-banyuls.fr 
[Accédé le 29 novembre 2014]

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