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Calanoida ( Ordre )
    Clausocalanoidea ( Superfamille )
        Clausocalanidae ( Famille )
            Clausocalanus ( Genre )
Clausocalanus pergens  Farran, 1926   (F,M)
Syn.: no Clausocalanus pergens : Mori, 1937 (1964) (p.35); Tanaka, 1960 (Pl.XII, figs.F, M)
Ref.:
Farran, 1926 (p.239, Descr.F, figs.F); 1929 (p.208, 224); Tanaka, 1956 cp.383, Rem.F); Vervoort, 1957 (p.38, Rem.); ? Park, 1968 (p.541, figs.F); Frost & Fleminger, 1968 (p.66, figs.F,M, Rem.); Carli & Crisafi, 1969 (p.280, figs.F); Corral Estrada, 1970 (p.142); Razouls, 1972 (p.94, Annexe: p.36, figs.F); Bradford, 1972 (p.36, figs.F); Williams & Wallace, 1975 (p.175, fig.F, Rem.); Dawson & Knatz, 1980 (p.7, figs.F,M); Björnberg & al., 1981 (p.629, figs.F,M); Alvarez-Marques, 1981 (p.157, figs.F, Rem.); Bradford-Grieve, 1994 (p.124, figs.F,M, fig.101); Chihara & Murano, 1997 (p.777, Pl.91,92: F,M); Bradford-Grieve & al., 1999 (p.878, 916, figs.F,M); Bucklin & al., 2003 (p.335, tab.2, fig.1, Biomol); Blanco-Bercial & Alvarez-marqués, 2007 (p.73, fig.2, Biomol.); Avancini & al., 2006 (p.82, Pl. 51, figs.F,M, Rem.); Vives & Shmeleva, 2007 (p.625, figs.F,M, Rem.)
Espèce Clausocalanus pergens - Planche 1 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [Pl. 51, p.202-203; Pl.52, p.204-205; Pl.53, p.206-207].

Female: 1 a, habitus (right lateral view); 1 b, idem (dorsal view); 1 c, rostrum (right lateral); 1 a-c taken from different specimens; 2 a , Th.4-5 (posterior part) and genital segment (right lateral); 2 b, genital segment (ventral view); 2 c-d, B2 of P2; 2 e-f, B2 of P3; 2 g, St of P2; 2 h, St of P3; 2 i, Re3 and St of P3; 2 j, Re3 and St of P4; 2 a-b taken from different specimens; 2 c, e from a third; 2 d-f from a fourth; 2 g-h from a fifth; 2 i-j from a sixth; 3 a, Th.4-5 (posterior part) and urosome with spermatophore attached (right lateral view); 3 b, Th.2 (posterior part), Th.3, Th.4-5 and urosome (right lateral view); 3 c-d, P5; 3 a-d taken from different specimens.
Nota: A1 segment 2 less than 0.95 times as long as segment 24. Exopodal segment 3 of P3 less than 1.20 times as long as terminal seta of P3. Exopodal segment 3 of P4 less than 1.30 times long as terminal seta of P4..


Espèce Clausocalanus pergens - Planche 2 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [Pl. 54, p.208-209; Pl.55, p.210-211].

Male: 1 a, habitus (right lateral view); 1 b, idem (dorsal view); 1 c, frontal region (right lateral view); 1 d, B2 of P2; 1 e-f, B2 of P3; 1 a-b from different specimens; 1 c from a third; 1 d-e from a fourth; 1 ffrom a fifth; 2 a, urosome (armature of caudal rami incomplete) (dorsal view); 2 b, Th.4-5 (posterior part) and urosome (left lateral view); 2 c, P5 (right posterolateral); 2 d, right P5; 2 a-b from different specimens; 2 c-d from a third.
Nota: Total body length less than 0.80 mm.


Espèce Clausocalanus pergens - Planche 3 de figures morphologiquesissued from : C. Razouls in Thèse Doc. ès Sciences, Univ. P & M. Curie, Paris , Tome annexe, 1972, Fig.37 (unpublished).
Female (from Banyuls): A, habitus (lateral right side); B, urosome (ventral); C, P5; D, basipod 2 of P2; E, basipod 2 of P3; F, idem; G, exopod 3 of P2; H, exopod 3 of P3; I, A1; J, basipod 2 of P2; K, basipod 2 of P3.


Espèce Clausocalanus pergens - Planche 4 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.78, Table 4a].
Clausocalanus pergens Females: Measurements and ratios .
TL = total body length ; SL = spermatophore length ; P :U = ratio of prosome length to urosome length ; U :U1 = ratio of total urosome length to length of 1st urosomal segment (genital segment) ; S :U1 = ratio of spermatophore sac length to U1 length of female on which spermatophore is attached ; r = sample range; m = sample mean; n = number of specimens measured; s = sample standard deviation.


Espèce Clausocalanus pergens - Planche 5 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.79, Table 4b].
Clausocalanus pergens Males: Measurements and ratios .
TL = total body length; P :U = ratio of prosome length to urosome length ; P:U2 = ratio of prosome length to length od 2nd urosomal segment; U2:2P5 = ratio of U2 length to length of 2nd segment of longer ramus of P5; U2:3P5 = ratio of U2 length of 3rd segment of longer ramus of P5; 3P5:2P5 = ratio of length of 3P5 of longer ramus to length of 2P5 of longer ramus; r = sample range; m = sample mean; n = number of specimens measured; s = sample standard deviation.


Espèce Clausocalanus pergens - Planche 6 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.21, Fig.4].
Exopodal segment 3:terminal seta length ratio of P4 (ordinate) plotted against length of exopodal segment 3 of P4 (abscissa) for females of Clausocalanus pergens.


Espèce Clausocalanus pergens - Planche 7 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.20, Fig.3].
Exopodal segment 3:terminal seta length ratio of P3 (ordinate) plotted against length of exopodal segment 3 of P3 (abscissa) for females of Clausocalanus pergens.


Espèce Clausocalanus pergens - Planche 8 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.205, Pl.52, i, j].
Female: i, exopodal segment 3 and terminal seta of P3; j, exopodal segment 3 and terminal seta of P4.


Espèce Clausocalanus pergens - Planche 9 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.205, Pl.52, c-d, e-f].
Female: c-d, basipodite 2 of P2; e-f, basipodite 2 of P3.


Espèce Clausocalanus pergens - Planche 10 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.207, Pl.53, c-d].
Female: c-d, P5 (from different specimens).


Espèce Clausocalanus pergens - Planche 11 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.211, Pl.55, b, c, d].
Male: b, posterior part of last thoracic segment and urosome (left lateral); c, P5 (right posterolateral); d, right P5.


Espèce Clausocalanus pergens - Planche 12 de figures morphologiquesissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.209, Pl.54, d, e, f].
Male: d, basipodite 2 of P2; e-f, basipodite 2 of P3; From different specimens.


Espèce Clausocalanus pergens - Planche 13 de figures morphologiquesissued from : J.M. Bradford in Mem. N. Z. Oceonogr. Inst., 1972, 54. [p.35, Fig.5 (25-26)].
Female (from Kaikoura, New Zealand): 25, habitus (lateral); 26, P5.
Scale bars: 0.1 mm (25, 26).


Espèce Clausocalanus pergens - Planche 14 de figures morphologiquesissued from : C. Razouls in Th. Doc. Etat Fac. Sc. Paris VI, 1972, Annexe. [Fig.37].
Female (from Banyuls, G. of Lion): A, habitus (lateral); B, urosome (dorsal); C, P5; D, basipod 2 of P2; E, basipod 2 of P3; F, basipod 2 of P3; G, exopod 3 of P2; H, exopod 3 of P3; I, A1; J, basipod 2 of P2; K, basipod 2 of P3.


Espèce Clausocalanus pergens - Planche 15 de figures morphologiquesissued from : A. Ianora, B. Scotto di Carlo, M.G. Mazzocchi & P. Mascellaro in J. Plankton Res., 1990, 12 (2). [p.255, Fig.6].
Longitudinal section (A) of Clausocalanus pergens parasitized by the phycomycetes (Coelomomycetaceae) which has destroyed most of the body tissue and led to total sexual castration of the host.
Close-up of parasite with (B) light microscope and (C) TEM (x 4300).
mg = midgut; fc = phycomycetal cells.
Scale bar = 0.100 mm.


Espèce Clausocalanus pergens - Planche 16 de figures morphologiquesIssued from F. Alvarez-Marques in Rev. Fac. Cienc. Univ. Oviedo (Ser. Biologia), (1979-80), 20-21, 1981. [p.165, Pl. II, Figs.4-7].
Female (from Gijon, NW Spain): 4, habitus (lateral); 5, genital segment and spermatheca (lateral); 6 & 7, P5.

Ref. compl.:
Yamazi, 1958 (p.148, Rem.); Grice & Hart, 1962 (p.287, table 3, p.293: Rem.); Duran, 1963 (p.14, Rem.); De Decker & Mombeck, 1964 (p.12); Anraku & Azeta, 1965 (p.13, Table 2, fish predator); Grice & Hulsemann, 1965 (p.223); Mazza, 1966 (p.70); Furuhashi, 1966 a (p.295, vertical distribution in Kuroshio region, Table 10); Fleminger, 1967 a (tabl.1); ? Delalo, 1968 (p.137); Champalbert, 1969 a (? part., p.633); Apostolopoulou, 1972 (p.327, 343); Björnberg, 1973 (p.311, 385); Corral Estrada & Pereiro Muñoz, 1974 (tab.I); Peterson & Miller, 1975 (p.642, 650, Table 3, interannual abundance); 1976 (p.14, Table 1, 2, 3, abundance vs interannual variations); 1977 (p.717, Table 1, seasonal occurrence); Carter, 1977 (1978) (p.35); Kovalev & Schmeleva, 1982 (p.83); Vives, 1982 (p.290); Scotto di Carlo & al., 1984 (p.1041); Tremblay & Anderson, 1984 (p.6: Rem.); Regner, 1985 (p.11, Rem.: p.28); Jansa, 1985 (p.108, Tabl.V); Brenning, 1985 a (p.28, Table 2); Greze & al., 1985 (p.7); Lozano Soldevilla & al., 1988 (p.58); Cervantes-Duarte & Hernandez-Trujillo, 1989 (tab.3); Pancucci-Papadopoulou & al., 1990 (p.199); Hirakawa & al., 1990 (tab.3); Ianora & al., 1990 (p.249, fig.; parsitism effects); Scotto di Carlo & al., 1991 (p.271); Yoo, 1991 (tab.1); Seguin & al., 1993 (p.23); Landry & al., 1994 (p.55, abundance, grazing); Landry & al., 1994 a (p.73, grazing, gut evacuation); Shih & Young, 1995 (p.72); Kotani & al., 1996 (tab.2); Go & al., 1997 (tab.1); Park & Choi, 1997 (Appendix); Noda & al., 1998 (p.55, Table 3, occurrence); Hure & Krsinic, 1998 (p.100); Siokou-Frangou & al., 1999 (p.205, Table 5); Seridji & Hafferssas, 2000 (tab.1); Siokou-Frangou, 1999 (p.476); Razouls & al., 2000 (p.343, Appendix); Moraitou-Apostolopoulou & al., 2000 (tab.I); Holmes, 2001 (p.42); Fragopoulu & al., 2001 (p.49, tab.1); Uysal & al., 2002 (p.17, tab.1); Peterson & al.2002 n (p.381, Table 2, interannual abundance); Sameoto & al., 2002 (p.12); Mackas & Galbraith, 2002 a (p.423, Table 2); Keister & Peterson, 2003 (p.341, Table 1, 2, abundance, cluster species vs hydrological events); Vukanic, 2003 (139, tab.1); Hsiao & al., 2004 (p.325, tab.1); Rezai & al., 2004 (p.489, tab.2, Rem.); Peralba & Mazzocchi, 2004 (p.645, figs.3,6); Fernandez de Puelles & al., 2004 (p.654, fig.7); Lo & al., 2004 (p.89, tab.1); Wang & Zuo, 2004 (p.1, Table 2, dominance, origin); Vukanic & Vukanic, 2004 (p.9, tab. 2, 3); Fernandez & al., 2004 (p.501, tab.5); Mackas & al., 2005 (p.1011, 1021, tab.2); Manning & Bucklin, 2005 (p.233, Table 1)); Zuo & al., 2006 (p.163: tab.1); Isari & al., 2006 (p.241, tab.II); Mackas & al., 2006 (L22S07, Table 2); Zervoudaki & al., 2006 (p.149, Table I); Hooff & Peterson, 2006 (p.2610); Fernandez de Puelles & al., 2007 (p.338, fig.7); Valdés & al., 2007 (p.103: tab.1); Khelifi-Touhami & al., 2007 (p.327, Table 1); Dur & al., 2007 (p.197, Table IV); McKinnon & al., 2008 (p.844: Tab.1); Cabal & al., 2008 (289, Table 1); Morales-Ramirez & Suarez-Morales, 2008 (p.519); Fernandes, 2008 (p.465, Tabl.2); Rossi, 2008 (p.90: Tableau XII); C.-Y. Lee & al., 2009 (p.151, Tab.2); Peterson, 2009 (p.73, Rem.: p 78); Galbraith, 2009 (pers. comm.); Chiba & al., 2009 (p.1846, Table 1, occurrence vs temperature change); Zhang W & al., 2009 (p.261); Brugnano & al., 2010 (p.312, Table 3, fig.8); Lidvanov & al., 2010 (p.356, Table 3); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical/tropical, Fig.4, 6, 7, Tabe 4); Mazzocchi & Di Capua, 2010 (p.425); Hafferssas & al., 2010 (p.1281, Table III, abundance vs spatial distribution); Hsiao S.H. & al., 2011 (p.475, Appendix I); Selifonova, 2011 a (p.77, Table 1, alien species in Black Sea); Keister & al., 2011 (p.2498, interannual variation); Mazzocchi & al., 2011 (p.1163, Table II, fig.6, long-term time-series 1984-2006); Isari & al., 2011 (p.51, Table 2, abundance vs distribution); Mazzocchi & al., 2012 (p.135, annual abundance 1984-2006); Shiganova & al., 2012 (p.61, Table 4); Jang M.-C & al., 2012 (p.37, abundance and seasonal distribution); Uysal & Shmeleva, 2012 (p.909, Table I); Salah S. & al., 2012 (p.155, Tableau 1); Miloslavic & al., 2012 (p.165, Table 2, transect distribution); Brugnano & al., 2012 (p.207, Table 2); Jean & al., 2012 (p.12, Table 3, protein vs environmental metal stress); Hidalgo & al., 2012 (p.134, Table 2); Takahashi M. & al., 2012 (p.393, Table 2, water type index); in CalCOFI regional list (MDO, Nov. 2013; M. Ohman, comm. pers.); Tachibana & al., 2013 (p.545, Table 1, seasonal change 2006-2008); Lidvanov & al., 2013 (p.290, Table 2, % composition); Pansera & al., 2014 (p.221, Table 2, abundance); Mazzocchi & al., 2014 (p.64, Table 3, 4, 5, spatial & seasonal composition %); Fernandez de Puelles & al., 2014 (p. in press, Table 3)
NZ: 18

Carte de distribution de Clausocalanus pergens par zones géographiques
Espèce Clausocalanus pergens - Carte de distribution 4issued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [p.68, Chart 10, a].
Occurrence of C. pergens in samples examined; closed circles represent samples examined; open circles represent samples in which adults were found; bars through open circles represent samples from which specimens were removed for measurements.
Espèce Clausocalanus pergens - Carte de distribution 5issued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [p.69, Chart 10, b].
Occurrence of C. pergens in samples examined; closed circles represent samples examined; open circles represent samples in which adults were found; bars through open circles represent samples from which specimens were removed for measurements.
Loc:
sub-Antarct. (Indian), South Africa (E), off S Cape Verde Is., off Morocco-Mauritania, Cap Ghir (Morocco), Canary Is., off Madeira, Azores, Bay of Biscay, S Brazil, G. of Mexico, New-York, G. of Maine, off Nova Scotia E, off S Newfoundland, off Ireland (S & W), Gijon (NW Spain), Iberian Shelf, Morocco-Mauritania, Ibero-moroccan Bay, Medit. (Alboran Sea, Gulf of Annaba, Baleares, Banyuls, Marseille, Toulon Harbour, Ligurian Sea, Napoli, Tyrrhenian Sea, Lake Faro, Strait of Messina, Adriatic Sea, Aegean Sea, Thracian Sea, Lebanon Basin, Black Sea), South Africa (Natal), S Indian, Bay of Bengal, Australia (North West Cape), Straits of Malacca, China Seas (Yellow Sea, East China Sea), Taiwan (SW, E, N: Mienhua Canyon, NW), Okinawa, S Korea, Korea Strait, Nagazaki, Kuchinoerabu Is., Japan, Tokyo Bay, Bikini Is., Pacif. (central N), Pacif. (E equatorial), Vancouver Is., off Washington coast, Oregon (Yaquina, off Newport), California, Santa Monica Basin, W Baja California, W Costa Rica, S Tasmania, New Zealand (N, Kaikoura, North Island SE), off SE Australia, Ecuador, Chile, off Santiago
N: 136
Lg.:
(30) F: 1,1-0,7; M: 0,67-0,52; (35) F: 1-0,9; (38) F: 1,05-0,9; (55) F: 0,99; (72) F: 0,94-0,85; (116) F: 0,94; (374) F: 1,025-0,75; (1138) F: 0,98-0,81; M: 0,58-0,55; {F: 0,70-1,10; M: 0,52-0,67}

The mean female size is 0.924 mm (n = 14; SD = 0.1147), and the mean size male is 0.580 mm (n = 4; SD = 0.0648). The size ratio (male : female) is about 0.65.
Rem.: épipélagique-bathypélagique.
Sampling depth (sub-Antarct.) : 100-250 m.
Certaines localisations sont à confirmer.
Voir aussi les remarques en anglais
Dernière mise à jour : 31/10/2017

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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2017. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Disponible sur http://copepodes.obs-banyuls.fr 
[Accédé le 18 décembre 2017]

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