Fiche d'espèce de Copépode
Calanoida ( Ordre )
    Clausocalanoidea ( Superfamille )
        Aetideidae ( Famille )
            Aetideus ( Genre )
Aetideus armatus  (Boeck, 1872)   (F,M)
Syn.: Pseudocalanus armatus Boeck, 1872; Brady, 1878 (p.46, figs.F);
Aetidius armatus : Brady, 1883 (part., p.76, figs.F); Sars, 1901 a (1903) (p.25, figs.F,M); Wolfenden, 1911 (p.209, figs.F); With, 1915 (p.75, figs.F); De Decker, 1968 (p.45); Papastephanou & al., 2006 (p.3078, Table 3);
Aethidius armatus: Cleve, 1904 a (p.185); A. tenuirostris Wolfenden, 1904 (p.116, Rem.);
Paraetideus armatus : Paulsen, 1909;
no A. armatus : Giesbrecht, 1892 (p.213); Giesbrecht & Schmeil, 1898 (p.31);
Undinopsis armatus : Shih & Young, 1995 (p.67);
no A. armatus : Esterly, 1905 (p.145, figs.F); Campbell, 1929 (p.311); ? Tanaka, 1957 a (p.31, figs.F,M)
Ref.:
Thompson & Scott, 1903 (p.233, 244); Farran, 1908 b (p.28); A. Scott, 1909 (p.37, figs.F); Lysholm, 1913 (p.5); Pesta, 1920 (p.507); Lysholm & Nordgaard, 1921 (p.11); Sars, 1925 (p.41); Farran, 1926 (p.246); 1929 (p.208, 228); Sewell, 1929 (p.99); Sciacchitano, 1930 (p.12, Rem.); Wilson, 1932 a (p.45, figs.F,M); Rose, 1933 a (p.80, figs.F,M); Jespersen, 1934 (p.52); Farran, 1936 a (p.87); Jespersen, 1940 (p.17); Mori, 1937 (1964) (p.38, figs. juv.F,M); Davis, 1949 (p.25, figs.F); Brodsky, 1950 (1967) (p.144, figs.F,M); Vervoort, 1952 a (n°42, p.3, figs.F,M); 1957 (p.44, Rem.); Vervoort, 1963 b (p.122, Rem.); Gaudy, 1963 (p.22); Matthews, 1964 (p.24, Rem.); Chen & Zhang, 1965 (p.50, figs.F); Owre & Foyo, 1967 (p.43, figs.F,M); Mazza, 1967 (p.108, 138, figs.juv., F,M, Rem.); 1968 (p.531, figs.); Park, 1968 (p.544, Rem.); Ramirez, 1969 (p.57, figs.F,M, Rem.); Vaupel-Klein, 1970 (p.4, 10, Rem.); Bradford, 1971 (p.15, figs.F, Rem); 1972 (p.36, figs.F,M); Razouls, 1972 (p.94, Annexe: p.39, figs.F,M); Park, 1974 (p.215, Rem.); Arcos, 1975 (p.15, figs.F,M); Park, 1978 (p.104); Dawson & Knatz, 1980 (p.6, figs.F,M); Björnberg & al., 1981 (p.604, 630, figs.F,M); Zheng Zhong & al., 1984 (1989) (p.235, figs.F); Razouls, 1994 (p.62, figs.F,M); Markhaseva, 1996 (p.14, figs.F,M, Rem.); Chihara & Murano, 1997 (p.682, Pl.32: F); Bradford-Grieve & al., 1999 (p.879, 920, figs.F,M); G. Harding, 2004 (p.53, 54, figs.F ,M); Vives & Shmeleva, 2007 (p.544, figs.F,M, Rem.); Blanco-Bercial & al., 2011 (p.103, Table 1, Biol. mol, phylogeny)
Espèce Aetideus armatus - Planche 1 de figures morphologiquesissued from : J.M. Bradford in N.Z. Jl Mar. Freshw. Res., 1971, 5 (1). [p.17, Fig.3]. From SW Africa (21°-35°S).
Male : a, habitus (dorsal); b, P5; c, caudal ramus; d, 3rd exopod segment of P3.
Nota: Posterior thoracic points extend up to (but not past), the posterior border of urosomal segment 1. Width: length ratio of caudal rami is 1: 2.58-2.70. Terminal seta on P2-P4 with 24, 22 and 18 teeth respectively.

Female: e, habitus (dorsal); f, idem (lateral right side); g, rostrum; h, genital segment and posterior thoracic points (lateral left side); i, idem (lateral right side); j, rostrum.
Nota: Posterior thoracic points just passed the posterior border of the genital segment, seminal receptacles with a thick, short tube joining the dorsal and ventral sacs. Rostrum: width/depth about 0.86-1.3 (falling within the range of the 2 specimens from Norway). Anterior cephalothorax vaulted, heavily chitinized and when viewed laterally indented above the base of A1 (characteristic not noted in the Norwegian specimens).


Espèce Aetideus armatus - Planche 2 de figures morphologiquesissued from : Sars G.O. in An Account of the Crustacea of Norway. Vol. IV. Copepoda Calanoida. Published by the Bergen Museum, 1903. [Pl. XIII]. As Aetidius armatus.
Female & Male.
C = forehead (lateral); R = rostrum (frontal view);


Espèce Aetideus armatus - Planche 3 de figures morphologiquesissued from : Sars G.O. in An Account of the Crustacea of Norway. Vol. IV. Copepoda Calanoida. Published by the Bergen Museum, 1903. [Pl. XIV]. As Aetidius armatus.
Female & Male.
M = Md; m = Mx1; mp1 = Mx2; mp2 = Mxp.


Espèce Aetideus armatus - Planche 4 de figures morphologiquesissued from : J.M. Bradford in N.Z. Jl Mar. Freshw. Res., 1971, 5 (1). [p.15, Fig.2].
Female (drawn from mounted specimens from Sars' collection): a, A2; b, Md (cutting edge); c, Md (mandibular palp); d, Mx1; e, Mx2; f, Mxp; g, P1; h, P2; i, P4.

Nota: Anterior integument covered with very small chitinous thickenings (visible only under phase contrast). Terminal spines of P2-P4 with 16, 17, 15 teeth respectively (although the numbers vary slightly between specimens). Seminal receptacle similar to A. australis


Espèce Aetideus armatus - Planche 5 de figures morphologiquesissued from : F.C. Ramirez in Contr. Inst. Biol. mar., Buenos Aires, 1969, 98. [p.56, Lam. IX, figs.57, 58, 61].
Female (from off Mar del Plata): 57, habitus (lateral left side); 58, idem (dorsal).

Male: 61, P5.
Scale bars in mm: 0.6 (57, 58); 0.1 (61).


Espèce Aetideus armatus - Planche 6 de figures morphologiquesissued from : A. Scott in Siboga-Expedition, 1909, XIX a. [Plate IV, Figs.14-25].
Female (from Indonesia-Malaysia): 14, habitus (dorsal); 15, forehead (lateral); 16, last thoracic and genital segments (left side); 17, rostrum; 18, A1; 19, A2; 20, Md; 21, Mx2; 22, Mxp; 23, P1; 24, P2; 25, P3.


Espèce Aetideus armatus - Planche 7 de figures morphologiquesissued from : R.N. Wolfenden in Die Marinen Copepoden der Deutschen Südpolar-Expedition 1901-1903, 1911. [p.208, Fig.4]. As Aetidius armatus.
Female: a-b, habitus (dorsal and lateral, respectively); c, rostrum


Espèce Aetideus armatus - Planche 8 de figures morphologiquesissued from : J.M. Bradford in Mem. N. Z. Oceonogr. Inst., 1972, 54. [p.37, Fig.6 (1-3)].
Female (from Kaikoura, New Zealand): 1, habitus (lateral).

Male: 2, habitus (dorsal); 3, P5.

Scale bars: 1 mm (1, 2); 0.1 mm (3).


Espèce Aetideus armatus - Planche 9 de figures morphologiquesissued from : C. Razouls in Th. Doc. Etat Fac. Sc. Paris VI, 1972, Annexe. [Fig.41, A, C].
Female: A, habitus (lateral).

Male: C, P5.


Espèce Aetideus armatus - Planche 10 de figures morphologiquesissued from : D.F.R. Arcos in Gayana, Zool., 1975, 32. [Lam.IV, Figs.29-34].
Female (from Bahia de Concepcion, Chile): 29-30, habitus (dorsal and lateral, respectively); 31, last thoracic segment and urosome (dorsal); 32, genital segment (lateral, left side); 34, exopodite segment 3 of P4.

male: 33, P5.


Espèce Aetideus armatus - Planche 11 de figures morphologiquesissued from : D.F.R. Arcos in Gayana, Zool., 1975, 32. [Lam.III, Fig.28].
Male: 28, last thoracic segment and urosome (dorsal).


Espèce Aetideus armatus - Planche 12 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf. 36 , Figs.7, 8, 9]. As Aëtidius armatus.
Female: 7, forehead (lateral); 8, urosome (ventral); 9, last thoracic segment and urosome (lateral, left side).


Espèce Aetideus armatus - Planche 13 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf. 36 , Fig.6]. As Aëtidius armatus.
Male: 6, habitus (dorsal).


Espèce Aetideus armatus - Planche 14 de figures morphologiquesissued from : C. With in The Danish Ingolf-Expedition, 1915, III (4). [p.75, Text-Fig.16]. As Aetidius armatus.
Female (from 61°-67°N, 9°-55°W):distal part of prosome and urosome (lateral, left side).


Espèce Aetideus armatus - Planche 15 de figures morphologiquesissued from : C. With in The Danish Ingolf-Expedition, 1915, III (4). [Pl. II, Figs.1a, 1b]. As Aetidius armatus.
Female: 1a, genital segment (left side); 1b, same (ventral).


Espèce Aetideus armatus - Planche 16 de figures morphologiquesissued from : C. With in The Danish Ingolf-Expedition, 1915, III (4). [Pl. II, Figs.1c, 1d]. As Aetidius armatus.
Female: 1c, basipod of left P4 (posterior); 1d, lamina labialis.
Nota: The basipod 2 of P4 has, in addition to marginal row of setae, posteriorly and just proximally to the inner seta a transverse row of 4 short spines (1c), which corresponds to the two spines mentioned by Giesbrecht but not by Sars and Scott. Glandular pores were found at the base of outer seta of exopod 2 and outer seta of exopod 3 in the three posterior pair of legs, and in addition to these a pore was found at the base of outer seta of exopod 1 in P3-P4.
The lamina labialis and the setae in front of it have a structure as shown in fig. 1d.


Espèce Aetideus armatus - Planche 17 de figures morphologiquesissued from : G. Harding in Key to the adullt pelagic calanoid copepods found over the continental shelf of the Canadian Atlantic coast. Bedford Inst. Oceanogr., Dartmouth, Nova Scotia, 2004. [p.53].
Female.

P1: Exopodite 1 without spine on the outer margin (arrowed). P2: Endopodal segments 1 and 2 incompletely separated (arrowed).


Espèce Aetideus armatus - Planche 18 de figures morphologiquesissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.26, Fig.122].
Female: 122, forehead (lateral).

Ref. compl.:
Rose, 1925 (p.152); Wilson, 1942 a (p.170); Lysholm & al., 1945 (p.11); Sewell, 1948 (p.347, 499, 508, 512, 516, 525, 545, 549, 555, 559, 566); C.B. Wilson, 1950 (p.159); Østvedt, 1955 Table 3, p.59); Yamazi, 1958 (p.148, Rem.); Fagetti, 1962 (p.17); Gaudy, 1962 (p.93, 99, Rem.: p.105); Duran, 1963 (p.15); V.N. Greze, 1963 a (tabl.2); Shmeleva, 1963 (p.141); Grice, 1963 a (p.495); Gaudy, 1963 (p.22, Rem.); Unterüberbacher, 1964 (p.21, Rem.M); De Decker & Mombeck, 1964 (p.11); Shmeleva, 1965 b (p.1350, lengths-volume -weight relation); Grice & Hulsemann, 1965 (p.223); Mazza, 1966 (p.70); Pavlova, 1966 (p.43); Matthews, 1967 (p.159, Table 1, Rem.); Fleminger, 1967 a (tabl.1); Grice & Hulsemann, 1967 (p.14); Berdugo & Kimor, 1968 (p.448); Dowidar & El-Maghraby, 1970 (p.267); Park, 1970 (p.475); Shih & al., 1971 (p.31, 142); Deevey, 1971 (p.224); Gamulin, 1971 (p.382, tab.3); Carli, 1971 (p.373, tab.2); Apostolopoulou, 1972 (p.327, 345); Boucher & Thiriot, 1972 (p.47, Tableau 4); Roe, 1972 (p.277, tabl.1, tabl.2); 1972 a (p.332); Björnberg, 1973 (p.326, 384); Vives & al., 1975 (p.41, tab.II, III, IV); Vives, 1976 (p.104); Deevey & Brooks, 1977 (p.256, tab.2, Station "S"); Carter, 1977 (1978) (p.35); Arashkevich, 1978 (p.118, Table: diets); Dessier, 1979 (p.204); Vaissière & Séguin, 1980 (p.23, tab.1); Vives, 1982 (p.290); Kovalev & Schmeleva, 1982 (p.83); Huntley & al., 1983 (p.143, Table 2); Brenning, 1983 (p.1, spatial distribution, T-S diagram, Rem.); De Decker, 1984 (p.316); Guangshan & Honglin, 1984 (p.118, tab.); Roe, 1984 (p.356); Scotto di Carlo & al., 1984 (p.1044); Tremblay & Anderson, 1984 (p.3); Sameoto, 1984 (p.767, vertical migration); Brenning, 1985 a (p.28, Table 2); Regner, 1985 (p.11, Rem.: p.30, as Aetidus ); Madhupratap & Haridas, 1986 (p.105, tab.1); Lozano Soldevilla & al., 1988 (p.58); Jimenez-Perez & Lara-Lara, 1988; Hernandez-Trujillo, 1989 a (tab.1); Cervantes-Duarte & Hernandez-Trujillo, 1989 (tab.3); Fransz & al., 1991 (p.9); Yoo, 1991 (tab.1); Hernandez-Trujillo, 1991 (1993) (tab.I); Hattori, 1991 (tab.1, Appendix); Hays & al., 1994 (tab.1); Kouwenberg, 1994 (tab.1); Hajderi, 1995 (p.542); Shih & Young, 1995 (p.66); Errhif & al., 1997 (p.422); Park & Choi, 1997 (Appendix); Suarez-Morales & Gasca, 1998 a (p107); Hure & Krsinic, 1998 (p.100); Padmavati & al., 1998 (p.349); Siokou-Frangou, 1999 (p.476); Halvorsen & Tande, 1999 (p.279, tab.2, 3, Rem.: p.282); Fernandez-Alamo & al., 2000 (p.1139, Appendix); Lapernat, 2000 (tabl. 3, 4); Razouls & al., 2000 (p.343, Appendix); Seridji & Hafferssas, 2000 (tab.1); Lopez-Salgado & al., 2000 (tab.1); Moraitou-Apostolopoulou & al., 2000 (tab.I); d'Elbée, 2001(tabl. 1); Holmes, 2001 (p.45); Hunt & al., 2001 (p.374, tab.1); Beaugrand & al., 2002 (p.1692); Beaugrand & al., 2002 (p.179, figs.5, 6); Sameoto & al., 2002 (p.12); Vukanic, 2003 (139, tab.1); CPR, 2004 (p.49, fig.134); Berasategui & al., 2005 (p.313, fig.2); Hop & al., 2006 (p.182, Table 4); Isari & al., 2006 (p.241, tab.II); Kosobokova & al., 2007 (p.929: Tab.7); Valdés & al., 2007 (p.103: tab.1); Castro & al., 2007 (p.486, fig.3, 5, Table 1, 2, 3); Cabal & al., 2008 (289, Table 1); Morales-Ramirez & Suarez-Morales, 2008 (p.517); Fernandes, 2008 (p.465, Tabl.2); Wishner & al., 2008 (p.163, Table 2, fig.8, oxycline); Gaard & al., 2008 (p.59, Table 1, N Mid-Atlantic Ridge); Raybaud & al., 2008 (p.1765, Table A1); Galbraith, 2009 (pers. comm.); Licandro & Icardi, 2009 (p.17, Table 4); C.E. Morales & al., 2010 (p.158, Table 1); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical/tropical); Hidalgo & al., 2010 (p.2089, Table 2); Mazzocchi & Di Capua, 2010 (p.423); Medellin-Mora & Navas S., 2010 (p.265, Tab. 2); Selifonova, 2011 (p.77, Table 1, alien species in Black Sea); Salah S. & al., 2011 (Tableau 1); Shiganova & al., 2012 (p.61, Table 4): Zizah & al., 2012 (p.79, Tableau I)
NZ: 24

Carte de distribution de Aetideus armatus par zones géographiques
Espèce Aetideus armatus - Carte de distribution 4issued from : A.A. Shmeleva in Bull. Inst. Oceanogr., Monaco, 1965, 65 (n°1351). [Table 6: 17 ]. Aetideus armatus (from South Adriatic).
Dimensions, volume and Weight wet. Means for 50-60 specimens. Volume and weight calculated by geometrical method. Assumed that the specific gravity of the Copepod body is equal to 1, then the volume will correspond to the weight.
Espèce Aetideus armatus - Carte de distribution 5issued from : U. Brenning in Wiss. Z. Wilhelm-Pieck-Univ. Rostock - 32. Jahrgang 1983. Mat.-nat. wiss. Reihe, 5. [p.1, Figs.1, 2].
Spatial distribution and T-S Diagram for Aetideus armatus, Aetideus giesbrechti and Aetideopsis carinata from 8° S - 26° N; 16°- 20° W.
Loc:
Cosmopolite: tropical & tempéré (except Arct.), Also Antarct. (Atlant. SW, Indien), sub-Antarct. (Atlant. SW, off Prince Edward W, Indien, Pacif. SW, SE), G. du St. Laurent, off SE Nova Scotia, Détr. de Davis S, Kongsfjorden (E Spitzbergen), Islande, off Féroé, Mer de Norvège, Raunefjorden, Mer du Nord, Manche, Medit., Marmara Sea, Black Sea, Arabian Sea, mers de Chine (Yellow Sea, East China Sea, South China Sea), Japan, Chili (N-S), Caribbean Colombia (San Andrés y Providencia), Station "S" (32°10'N, 64°30'W),
N: 195
Lg.:
(7) F: 1,93; (14) F: 1,7-1,65; (22) F: 1,95-1,80; M: 1,53-1,4; (34) F: 1,44-1,33; (35) F: 1,84-1,68; (37) F: 2-1,6; M: 1,53-1,3; (38) F: 1,95-1,8; M: 1,53; ? (39) F: 1,82; M: 1,8; (45) F: 1,8; M: 1,45-1,25; (59) F: 2-1,8; M: 1,5-1,4; (65) F: 1,8; M: 1,45; (72) F: 2,02-1,8; (116) F: 1,78; M: 1,56; (199) F: 1,9-1,6; (202) F: 1,78-1,95; M: 1,4-1,56; (204) F: 1,9-1,6; M: 1,5-1,3; (205) F: 2,25-2,15; M: 2,1-2; (244) F: 1,9; (290) F: 1,6-1,75; (432) F: 1,98-1,83; (449) F: 1,8; M: 1,45; (866) F: 1,65-1,95; M: 1,4-1,8; (920) F: 1,51; (1109) F: 1,58-1,87; {F: 1,33-2,25; M: 1,25-2,10}
Rem.: épi-bathypélagique.
Sampling depth (Antarct., sub-Antarct.) : 0-600; 100-1000 m.
Voir aussi les remarques en anglais
Dernière mise à jour : 22/05/2013

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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2012. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Disponible sur http://copepodes.obs-banyuls.fr 
[Accédé le 23 mai 2013]

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