Cyclopoida ( Order )
Corycaeidae Dana, 1852 ( Cyclopoida ) *
Ref.: Claus, 1863 (part., p.83) ; Giesbrecht, 1892 (part., p.83); Esterly, 1905 (part., p.218); van Breemen, 1908 a (part., p.194 ; M. Dahl, 1912 (p.4, Rem.: S.G); Pesta, 1928 c (p.125); Sars, 1918 (p.194); Rose, 1929 (p.67); Oliveira, 1946 (p.474); Gonzalez & Bowman, 1965 (p.265); Owre & Foyo, 1967 (p.119); Chen, Zhang & Zhu, 1974 (p.52); Björnberg & al.,1981 (p.673); Bowman & Abele, 1982 (p.11); Razouls, 1982 (p.684); Zheng Zhong & al., 1984 (1989) (p.266, spp. key); Sazhina, 1985 (p.94, N); Kang & al., 1990 (p.51, sub-genera and species keys); Huys & Boxshall, 1991 (p.284, 461); Razouls, 1993 (p.312); Chihara & Murano, 1997 (p.962); Bradford-Grieve & al., 1999 (p. 887, 973, clé des G. & S-G.); Boxshall & Halsey, 2004 (p.16; 491: Def.; p.493: Genera and species keys; Rem. p.1: These authors are questionable concerning the order of Poecilostomatoida, and the validity of sub-genera); Vives & Shmeleva, 2010 (p.196, Rem., Genera key instead of Sub-genera)
Rem.: 2 G. (maintained provisionally): Corycaeus, Farranula.
* We thank Ruth Böttger-Schnack (Universität Bremen) for having forwarded to us a copy of the families Corycaeidae, Oncaeidae et Sapphirinidae from the Fauna of Japan by Chihara et Murano (1997), as well as the English translation concerning the Oncaeidae family performed by Hiroshi Itoh.
(1) Corycaeus Dana, 1845
Syn.: Corycäus Dana, 1845; Giesbrecht, 1892 (p.85, 659); Agetus Kröyer, 1849 (in Damkaer & Damkaer, 1979, p.43); Giesbrecht, 1892 (p.659)
Ref.: Dahl, 1894 (p.67, clé spp.); Wheeler, 1901 (p.191); Esterly, 1905 (p.224); van Breemen, 1908 a (p.197, clé spp.); A. Scott, 1909 (p.245); Farran, 1911 a (p.282, 287); M. Dahl, 1912 (p.3, Rev., clé spp.); Sars, 1918 (p.194); Pesta, 1920 (p.645); Gurney, 1927 (p.160); Rose, 1929 (p.67); Wilson, 1932 a (p.354, clé spp.); Rose,1933 a (p.324, clé des S-G); Mori; 1937 (1964) (p.130, clé spp.); Dakin & Colefax, 1940 (p.110, clé S.G); Oliveira, 1946 (p.475); Sewell, 1947 (p.272, Rem.); Davis, 1949 (p.75); Carvalho, 1952 a (p.168); Motoda, 1963 (p.209, 213); Gonzalez & Bowman, 1965 (p.266); Owre & Foyo, 1967 (p.119, clé des S-G, clé spp.); Chen, Zhang & Zhu, 1974 (p.52); Gibson & Grice, 1978 (p.66, Developmental stages, figs.); Kabata, 1979 (p.20); Björnberg & al., 1981 (p.675); Razouls, 1982 (p.684); Gardner & Szabo, 1982 (p.97); Zheng Zhong & al., 1984 (1989) (p.266, clé spp., Rem.); Huys & Boxshall, 1991 (p.461); Razouls, 1993 (p.313); Chihara & Murano, 1997 (p.962); Bradford-Grieve & al., 1999 (p.973, clé spp.); Boxshall & Halsey, 2004 (p.491: Def.; p.493: Clé des espèces)
Rem.: 6 S.G.: Agetus, Corycaeus, Ditrichocorycaeus, Monocorycaeus, Onychocorycaeus, Urocorycaeus. Boxshall (1998, p.221 & foll.) and Boxshall & Halsey, 2004 (p.491, 493) consider the sub-genera as genera awaiting an inevitable phylogenetic revision.
Total: 33 espèces + 2 unidentified.
Remarks on dimensions and sex ratio:
The mean female size is 1,297 mm (n= 33; S= 0,514; Cv= 0,396) and the mean male size is 1,143 mm (n= 31; S= 0,402; Cv= 0,352). The size ratio (M/F) is 0,878 ou 87,8 % (n= 31; S= 0,084; Cv= 0,096).
Genus Corycaeus - Plate 1issued from : C.T. Achuthankutty, N. Ramaiah & G. Padmavati in Pelagic biogeograpgy ICoPB II. Proc. 2nd Intern. Conf. Final report of SCOR/IOC working group 93, 9-14 July 1995. Workshop Report No. 142, Unesco, 1998. [p.8, Fig.6].
Salinity ranges for Corycaeus spp. in coastal and estuarine waters of Goa (India).
Shaded area indicates the range of higher abundance.
Agetus Kröyer, 1849
Ref.: Kröyer, 1849 (in Damkaer & Damkaer, 1979, p.43); M. Dahl, 1912 (p.11, 30); Rose, 1933 a (p.325); Chihara & Murano, 1997 (p.964, Pl.211); Bradford-Grieve & al., 1999 (p.974: clé spp.); Boxshall & Halsey, 2004 (p.494)
Rem.: type: Corycaeus typicus
Corycaeus Dana, 1845
Ref.: M. Dahl, 1912 (p.12); Rose, 1933 a (p.325); Dakin & Colefax, 1940 (p.111, clé spp.); Sewell, 1947 (p.272, Rem.); Kang & al., 1990 (p.51, clé spp.); Chihara & Murano, 1997 (p.966, Pl.211); Bradford-Grieve & al., 1999 (p.974: clé spp.); Boxshall & Halsey, 2004 (p.494)
Rem.: type: Corycaeus speciosus Dana,1849
Ditrichocorycaeus M. Dahl, 1912
Ref.: M. Dahl, 1912 (p.11, 51); Rose, 1933 a (p.325); Dakin & Colefax, 1940 (p.113, clé spp.); Sewell, 1947 (p.278, Rem.); Tanaka, 1957 (p.85, Rem.); Vilela, 1968 (p.37, Rem.); Gonzalez & Bowman, 1965 (p.266); Bradford, 1978 (p.138); Kang & al., 1990 (p.52, clé spp.); Chihara & Murano, 1997 (p.963, Pl.210); Bradford-Grieve & al., 1999 (p.974: clé spp.); Boxshall & Halsey, 2004 (p.493)
Rem.: type: Corycaeus. (D-) angicus
Monocorycaeus M. Dahl, 1912
Ref.: M. Dahl, 1912 (p.11, 27); Rose, 1933 a (p.325); Chihara & Murano, 1997 (p.962, Pl.211); Bradford-Grieve & al., 1999 (p.974); Boxshall & Halsey, 2004 (p.494)
Rem.: type: ?
Onychocorycaeus M. Dahl, 1912
Ref.: M. Dahl, 1912 (p.11, 82); Rose, 1933 a (p.325); Dakin & Colefax, 1940 (p.113, clé spp.); Sewell, 1947 (p.283); Kang & al., 1990 (p.53, clé spp.); Chihara & Murano, 1997 (p.965, Pl.211); Bradford-Grieve & al., 1999 (p.974, 975: clé spp.); Boxshall & Halsey, 2004 (p.493)
Rem.: type: Corycaeus latus
Urocorycaeus M. Dahl, 1912
Ref.: M. Dahl, 1912 (p.10, 41); Rose, 1933 a (p.325); Dakin & Colefax, 1940 (p.111, clé spp.); Chihara & Murano, 1997 (p.962, Pl.210); Bradford-Grieve & al., 1999 (p.973, 975: clé spp.); Boxshall & Halsey, 2004 (p.493)
Rem.: type: Corycaeus lautus
Corycella Farran, 1911
Ref.: Farran, 1911 a (p.283); C. B. Wilson, 1950 (p.227, Rem.)
Rem.: Cf. Farranula
(2) Farranula Wilson, 1932
Syn.: Corycella Farran, 1911 a (p.283); Wilson, 1932 (p.41, Rem.); 1932 a (p.361); Dakin & Colefax, 1940 (p.110, clé spp.); Sewell, 1951 (p.385, Rem.); Kasturirangan, 1963 (p.79); Corycaeus ( Corycella ) M. Dahl, 1912 (p.9, 106, clé spp.)
Ref.: Wilson, 1942 a (p.186: Rem.); 1950 (p.227); Motoda, 1963 (p.252); Gonzalez & Bowman, 1965 (p.267); Owre & Foyo, 1967 (p.119, clé des spp., Rem.); Vilela, 1968 (p.41); Corral Estrada, 1970; Razouls, 1982 (p.703); Kang & al., 1990 (p.54); Huys & Boxshall, 1991 (p.287); Razouls, 1993 (p.313); Chihara & Murano, 1997 (p.967, Pl.210); Bradford-Grieve & al., 1999 (p.888, 973, 975: clé spp.); Boxshall & Halsey, 2004 (p.493); Wi & Soh, 2013 (p.21)
Rem.: Böttger-Schnack (1988, p.313, 315: tab.1, 2) consider this genus as a sub-genus of Corycaeus, but no (p.314, tab.3). In 1990 a and 1990 b the author maintains the sub-genus following the initial position of M. Dahl (1912).
Total: 7 species.
Remarks on dimensions and sex ratio:
The mean female size is 0,872 mm (n= 6; S= 0,083; Cv= 0,095) and the mean male size is 0.785 mm (n= 6; S= 0,071; Cv= 0,09). The size ratio (M/F) is 0,901 ou 90,1 % (n= 6; S= 0,031; Cv= 0,034). The sex-ratio is 1.

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