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List species and varieties by family |
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| Eucalanidae Giesbrecht, 1892 ( Eucalanoidea ) | |
| | | (1) Eucalanus Dana, 1852 ( Eucalaninae ) | |
| | Rem.: | Type: Calanus elongatus (Dana,1849).
Not all authors follow Geletin (1976) in his defenition of the genera Pareucalanus and Subeucalanus (Brodsky & al., 1983, p.203).
Fleminger (1973) splits up 17 spp. into 4 groups: 'subtenuis', 'pileatus', 'elongatus', 'attenuatus'.
17 spp. + 1 debated (E. pseudattenuatus ) + 2 to specify (E. muticus , E. peruanus ) + 3 very doubtful.
Bradford-Grieve (1994) rehabilitates the two genera Pareucalanus and Subeucalanus.
The genus Eucalanus comprises 6 spp. + 1 doubtful.
Diagnosis from Bradford-Grieve (1994, p.74) :
- As in the family definition.
- Urosome female 4-segmented, and posterior borders of pedigers 2-4 and urosome segment 1 without spines.
- Integumental organs located at Mx1 and Mx2 sites vary from 8-18 in number are not arranged in a rectangular pattern, posterior sets more lateral than the anterior sets. Mxp sites are in 4 or more pairs. Pediger segments 2 and 3 with 1 or 2 lateral sites ; pediger segment 4 lacking paired sites lying outside the dorsal quadrangle of hair sensilla ; abdominal segment 1-2 with at least 2 dorsal perforations.
- A2 endopod elongate, exopodal segments 1 and 2 separate.
- Md palp basis and exopod elongate ; endopod small, inserted just beyond midlength on basis which also bears 1 or 3 setae.
- Mx1 with 3 inner lobes bearing setae.
- P1 with a 2-segmented endopod and 3-segmented exopod.
P5 absent in female.
- P5 uniramous on both sides in male. |
| | Eucalanus acus Farran, 1929 (F) | | Ref.: | Farran, 1929 (p.218, figs.F); Sewell, 1948 (p.513, Rem.) | | Rem.: | Cf. Subeucalanus longiceps |
| | Eucalanus atlanticus Wolfenden, 1904 (M) | | Ref.: | Wolfenden, 1904 (p.113, figs.M); With, 1915 (p.48, Rem.); Candeias, 1926 (1929) (p.27, figs.M); Rose, 1933 a (p.69, figs.M, Rem.) | | Rem.: | Cf. Eucalanus elongatus (juv.) |
| | Eucalanus attenuatus Dana, 1849 (F,M) | | Syn.: | Calanus attenuatus Dana,1849 | | Ref.: | Bradford-Grieve, 1994 (p.80, Rem.) | | Rem.: | Cf. Pareucalanus attenuatus |
| | [1] Eucalanus bungii Giesbrecht, 1892 (F,M) [Figs] | |
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| | Eucalanus bungii bungii Johnson, 1938 (F,M) | | Ref.: | M.W Johnson, 1938 (p.167, figs.F,M); Vervoort, 1946 (p.91, Rem.); Sewell, 1948 (p.561, Rem.); Brodsky, 1950 (1967) (p.101, 102, figs.F, M); Mazza, 1967 (p.71, Rem.); Vidal, 1971 a (p.16, figs.F,M); Shih & al., 1971 (p.146, 204); Tanaka, 1956 (p.267, Rem.) | | Ref. compl.: | Harding, 1966 (p.17, 69); Dunbar & Harding, 1968 (p.319, 322); Omori, 1969 (p.5); Morris, 1970 (p.2300); Shih & Laubitz, 1978 (p.53); Van der Spoel & Heyman, 1983 (p.94, fig.58); Gordon & al., 1985 (p.91, 92); Sasaki & al., 1988 (tab.1); Coyle & al., 1990 (p.763); Hirakawa & al., 1990 (tab.3); Hattori, 1991 (tab.1, Appendix); Shih & Young, 1995 (p.69); Kotani & al., 1996 (tab.2); Park & Choi, 1997 (Appendix); Kosobokova & Hirche, 2000 (p.2029, tab.2) | | Loc: | Arct. (Fletcher's Ice Is.), Alaska (Auke Bay), off I. Reine Charlotte, Colombie Britannique, Arct. (bassin polaire) (rare), Japon, off Hokkaido, Corée, Mer de Chine E, Pacif. central N (subarct.) | | N: | 19 | | Lg.: | (22) F: 8-6,5; M: 5,4-4,8 | | Rem.: | Delalo (1968, p.137) signale cette forme en Mer Rouge et G. d'Aden ce qui nécessite confirmation. Cf. Eucalanus bungii. |
| | Eucalanus bungii californicus Johnson, 1938 (F,M) | | Syn.: | Eucalanus bungii : Davis, 1949 (p.16, figs.F,M) | | Ref.: | M. W. Johnson, 1938 (p.169, figs.F); Vervoort, 1946 (p.91, Rem.); Sewell, 1948 (p.561, Rem.); Mazza, 1967 (p.71, Rem.); Morris, 1970 (p.2300); Shih & al., 1971 (p.147); Hernandez-Trujillo, 1991 (1993) (tab.I); Van der Spoel & Heyman, 1983 (p.94, fig.58) | | Loc: | Détr. Juan de Fuca, I. Malcolm | | Rem.: | Cf. Eucalanus californicus |
| | [2] Eucalanus californicus Johnson, 1938 (F,M) [Figs] | |
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| | Eucalanus crassus Giesbrecht, 1888 (F,M) | | Syn.: | Eucalanus oculanus Marukawa, 1921 (p.11); Brodsky, 1950 (1967) (p.105) | | Ref.: | Bradford-Grieve, 1994 (p.87) | | Rem.: | Cf. Subeucalanus crassus |
| | Eucalanus danai Lubbock, 1860 | | Syn.: | Calanus danai Lubbock,1860 | | Ref.: | Giesbrecht, 1892 (p.132); Giesbrecht & Schmeil, 1898 (p.22, Rem.) | | Loc: | G. de Guinée | | Rem.: | sp. douteuse |
| | Eucalanus dentatus A. Scott, 1909 (F) | | Ref.: | A. Scott, 1909 (p.22, figs.F); Geletin, 1976 (p.84); Fleminger, 1973 (p.978, 999, figs.F) | | Loc: | Indonésie-Malaisie | | Rem.: | Cf. Subeucalanus dentatus |
| | [3] Eucalanus elongatus (Dana, 1849) (F,M) [Figs] | |
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| | Eucalanus elongatus bungii Giesbrecht, 1892 | | Ref.: | Giesbrecht, 1892 (p.149); Vervoort, 1946 (p.91, Rem.); Davis, 1949 (p.16) | | Rem.: | Cf. Eucalanus bungii ou californicus |
| | Eucalanus elongatus hyalinus Claus, 1866 (F,M) | | Syn.: | Calanella hyalina Claus,1866; Eucalanus spinifer T. Scott,1894 b ; Eucalanus atlanticus Wolfenden,1904 | | Ref.: | Giesbrecht, 1892 (p.131, 149, figs.F,M, Rem.); M.W. Johnson, 1938 (p.167, 169, fig.F); Vervoort, 1946 (p.91, 92, Rem.); 1957 (p.32, Rem.); 1963 b (Rem., p.88); Sewell, 1948 (p.561, Rem.); Davis, 1949 (p.16); Marques, 1968 (p.395); Corral Estrada, 1970 (p.72, Rem.); Mazzocchi & al., 1995 (p.116, figs.F, Rem.) | | Rem.: | Cf. Eucalanus hyalinus |
| | Eucalanus elongatus inermis Giesbrecht, 1892 | | Ref.: | Giesbrecht, 1892 (p.149); Vervoort, 1946 (p.91, 93, Rem.); 1963 b (p.89, Rem.); Sewell, 1948 (p.561, Rem.); Davis, 1949 (p.16) | | Rem.: | Cf. Eucalanus inermis |
| | Eucalanus giesbrechti Mori, 1937 (F,M juv.) | | Syn.: | Eucalanus elongatus bungii Giesbrecht,1892 (part.); Eucalanus elongatus (F): Esterly,1905 (p.131,figs.F,M) | | Ref.: | Mori, 1937 (1964) (p.22, figs.F, juv.M); Davis, 1949 (p.17, Rem.); Vervoort, 1957 (p.32) | | Rem.: | Cf. Eucalanus californicus |
| | [4] Eucalanus hyalinus (Claus, 1866) (F,M) [Figs] | |
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| | [5] Eucalanus inermis Giesbrecht, 1892 (F,M) [Figs] | |
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| | Eucalanus langae Fleminger, 1973 (F,M) | | Ref.: | Fleminger, 1973 (p.978, 990, 996, figs.) | | Rem.: | Cf. Pareucalanus langae |
| | Eucalanus longiceps Matthews, 1925 (F,M) | | Syn.: | Eucalanus acus Farran,1929 (p.218, figs.F); ? E. attenuatus : With, 1915 (p.52); Subeucalanus longiceps : Geletin, 1976 (p.91) | | Ref.: | Bradford-Grieve, 1994 (p.87) | | Rem.: | Cf. Subeucalanus longiceps |
| | Eucalanus monachus Giesbrecht, 1888 (F,M) | | Syn.: | Eucalanus vadicola F. Dahl,1894; non E. monachus : Dakin & Colefax, 1940 | | Ref.: | Bradford-Grieve, 1994 (p.88) | | Rem.: | Cf. Subeucalanus monachus |
| | Eucalanus mucronatus Giesbrecht, 1888 (F,M) | | Syn.: | non E. mucronatus : Mori, 1937 (1964) (p.25, figs.F,M) | | Ref.: | Bradford-Grieve, 1994 (p.88) | | Rem.: | Cf. Subeucalanus mucronatus |
| | [6] Eucalanus muticus C.B. Wilson, 1950 (F,M) [Figs] | |
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| | Eucalanus oculanus Marukawa, 1921 (F,M) | | Ref.: | Brodsky, 1950 (1967) (p.105, figs.F,M) | | Rem.: | Mori, 1937 (1964) (p.24) considère l'espèce comme synonyme de Eucalanus crassus (juv.). Cf. Subeucalanus crassus |
| | Eucalanus parki Fleminger, 1973 (F,M) | | Syn.: | Eucalanus attenuatus Park, 1968 (p.533, grandes formes); E. quadrisetosus Geletin, 1973 (p.1786: M) | | Ref.: | Bradford-Grieve, 1994 (p.80, Rem.) | | Rem.: | Cf. Pareucalanus parki |
| | Eucalanus peruanus Volkov, 1971 (F,M) | | Ref.: | Volkov, 1971 (p.141, figs.F,M) | | Rem.: | Cf. Pareucalanus peruanus |
| | Eucalanus pileatus Giesbrecht, 1888 (F,M) | | Syn.: | Eucalanus monachus : Dakin & Colefax,1940; E. saravatae Oliveira, 1945; E. subcrassus-pileatus : Deevey, 1960 (p.16); Björnberg, 1963 (p.20) | | Ref.: | Bradford-Grieve, 1994 (p.88) | | Rem.: | Cf. Subeucalanus pileatus |
| | Eucalanus pseudattenuatus Sewell, 1947 (F,M) | | Syn.: | Eucalanus pseudoattenuatus : Brodsky, 1962 c (p.112, figs.F,M); Chen & Zhang, 1965 (p.34, figs.F); Deevey, 1971 (p.224); Sazhina, 1985 a (p.491, tab.1) | | Ref.: | Sewell, 1947 (p.47, figs.F); Rose, 1956 (p.459); Kasturirangan, 1963 (p.17, 19, fig.F); Vervoort, 1963 b (p.90); Geletin, 1973 (p.1783, 1786, figs.); 1976 (p.83); Fleminger, 1973 (p.967, 990, 992); Chen Q-c, 1980 (p.794); Brodsky & al., 1983 (p.208); Zheng Zhong & al., 1984 (1989) (p.229, figs.F,M); Shih & Young, 1995 (p.69); Wong & al, 1998 (tab.2); Mauchline, 1998 (tab.30) | | Lg.: | (290) F: 3,75-3,8; (864) F: 4-4,2 | | Rem.: | Cf. Pareucalanus attenuatus et sewelli |
| | Eucalanus quadrisetosus Geletin, 1973 (M) | | Ref.: | Geletin, 1973 (p.1786, figs.M, non F); Brodsky & al., 1983 (p.212, Rem.) | | Rem.: | Cf. Paereucalanus parki |
| | Eucalanus saravatae Oliveira, 1945 (F) | | Ref.: | Oliveira, 1945 a (p.456, fig.F); Björnberg & al., 1981 (p.604, Rem.) | | Rem.: | Cf. Eucalanus pileatus |
| | Eucalanus setiger Brady, 1883 (F,M) | | Ref.: | Brady, 1883 (p.39,figs.F,M); Thompson, 1888 d (p.139); Giesbrecht, 1892 (p.132); T. Scott, 1894 b (p.30); Giesbrecht & Schmeil, 1898 (p.22) | | Rem.: | Cf. ? Eucalanus subtenuis ou Subeucalanus crassus |
| | Eucalanus sewelli Fleminger, 1973 (F,M) | | Syn.: | E. attenuatus : Vervoort, 1946 (part., p.95); Sewell, 1947 (part., p.39, fig. 7B); Brodsky, 1962 (p.113, fig.14); Ramirez, 1969 (p.44); Björnberg, 1973 (p.226) | | Ref.: | Bradford-Grieve, 1994 (p.80, 82, Rem.) | | Rem.: | Cf. Pareucalanus sewelli |
| | [7] Eucalanus spinifer T. Scott, 1894 (F,M) [Figs] | |
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| | Eucalanus subcrassus Giesbrecht, 1888 (F,M) | | Ref.: | Bradford-Grieve, 1994 (p.88) | | Rem.: | Cf. Subeucalanus subcrassus |
| | Eucalanus subtenuis Giesbrecht, 1888 (F,M) | | Ref.: | Giesbrecht, 1892 (p.132, 150, figs.F,M); Sars, 1925 (p.21); Farran, 1929 (p.218); Vervoort, 1946 (p.106); 1963 b (p.91); Carvalho, 1952 a (p.141, figs.M); Fukase, 1957 (p.17, figs.F,M, Rem.); Marques, 1968 (p.396); 1974 (p.12); 1982 (p.753); Fleminger, 1973 (p.969, 978, 979, 982, 999, figs.F); 1975 (p.395); Geletin, 1976 (p.83, 91, Rem.); Greenwood, 1976 (p.16); Van der Spoel & Heyman, 1983 (p.40, fig.52) | | Rem.: | Cf. Subeucalanus subtenuis |
| | Eucalanus subtenuis japonica Fukase, 1957 (F,M) | | Syn.: | Eucalanus subtenuis : Tanaka,1935; Eucalanus mucronatus : Mori, 1937 (1964) (p.25, figs.F,M) | | Ref.: | Fukase, 1957 (p.19, figs.F,M, Rem.); Grice, 1962 (p.181, figs.F,M, Rem.) | | Loc: | Japon | | Rem.: | Cf. Subeucalanus subtenuis japonica |
| | Eucalanus vadicola Dahl, 1894 (F,M) | | Ref.: | Dahl, 1894 c (p.20, Rem. F,M); Giesbrecht & Schmeil, 1898 (p.21); Björnberg, 1963 (p.21, Rem.) | | Loc: | Estuaire de l'Amazone | | Rem.: | Cf. Subeucalanus monachus |
| | Eucalanus sp. Candeias, 1926 (? M juv) | | Ref.: | Candeias, 1926 (1929) (p.28, figs.juv) | | Loc: | Portugal |
| | Eucalanus sp. Tanaka, 1935 (F,M) | | Ref.: | Tanaka, 1935 (p.146, figs.F,M) | | Loc: | Japon (Sagami Bay) | | Lg.: | (389) F: 6,19-5,72; M: 4,67-4,5; {F: 5,72-6,19; M: 4,5-5,72} | | Rem.: | Cette forme nécessite confirmation. | | | (2) Pareucalanus Geletin, 1976 ( Eucalaninae ) | |
| | Rem.: | type: Paracalanus attenuatus Dana,1849. Total: 6 spp. + 1 undetermined.
Brodsky & al., 1983 (p.203) do not acknowledge the creation of this genus. The genus is confirmed by Bradford-Grieve (1994)
Diagnosis from Bradford-Grieve (1994, p.80) :
- As in the family definition.
- Urosome female 3-segmented, and posterior borders of pedigers 2-4 and urosome segment 1 without spines.
- Integumental organs located at M11 and Mx2 with a total of 10 sites forming a rectangle and arranged anterior to posterior in 2 bilateral sets of 1 pair, 1 single and 1 pair of sites ; Mxp site has 6 pairs ; pedigers segments 2 and 3 in lateral view with 3 sites ; pediger segment 4 lacking paired sites lying outside of the dorsal quadrangle of hair sensilla ; urosome segment 4-5 with at least 3 dorsal sites.
- A2 endopod elongate, exopod segments 1 and 2 separate.
- Md palp basis and exopod elongate ; endopod small, inserted just at midlength on basis which also bears 2 or 4 setae.
- Mx1 with 3 inner lobes bearing setae.
- P1 with a 2-segmented endopod and 3-segmented exopod.
- P5 female absent.
- P5 male uniramous on both sides with right leg not extending beyond the distal part of left leg segment 2. |
| | [1] Pareucalanus attenuatus (Dana, 1849) (F,M) [Figs] | |
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| | [2] Pareucalanus langae (Fleminger, 1973) (F,M) [Figs] | |
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| | [3] Pareucalanus parki (Fleminger, 1973) (F,M) [Figs] | |
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| | [4] Pareucalanus peruanus (Volkov, 1971) (F,M) | |
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| | [5] Pareucalanus sewelli (Fleminger, 1973) (F,M) [Figs] | |
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| | [6] Pareucalanus smithae Prusova, 2007 (F,M) [Figs] | |
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| | [7] Pareucalanus sp. Goetze & Ohman, 2010 [Figs] | |
| | | | Rem.: | Cf. Rhincalanidae. | | | | (3) Subeucalanus Geletin, 1976 | |
| | Rem.: | type: Eucalanus subtenuis Giesbrecht,1888. Total: 9 spp. + 3 undet.
Brodsky & al. (1983, p.203) do not acknowledge the creation of this genus. Bradford-Grieve (1994) affirms the validity of the genus.
Diagnosis from Bradford-Grieve (1994, p.69) :
- As in the family definition.
- Female urosome 3-segmented, and posterior borders of pedigers 2-4 and urosome segment 1 without spines.
- Integumental organs located at Mxp site in 2 pairs ; pediger segments 3 and 4 with a single mid-dorsal site lying between the anterior 2 hair sensilla ; pediger segment 4 with 1 pair of sites lateral to the quadrangle formed by 4 hair sensilla.
- A2 endopod elongate, exopod segments 1 and 2 fused.
- Md palp basis and exopod elongate ; endopod small, inserted on the terminal one third or quarter of basis which also bears 2 or 3 setae.
- Mx1 with 2 inner lobes bearing setae, 1 lobe absent.
- P1 with a 2-segmented endopod and 3-segmented exopod.
- P5 female absent.
- P5 male uniramous with right leg absent. |
| | [1] Subeucalanus crassus (Giesbrecht, 1888) (F,M) [Figs] | |
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| | [2] Subeucalanus dentatus (A. Scott, 1909) (F) [Figs] | |
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| | [3] Subeucalanus flemingeri Prusova, Al-Yamani & Al-Mutairi, 2001 (F,M) [Figs] | |
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| | [4] Subeucalanus longiceps (Matthews, 1925) (F,M) [Figs] | |
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| | [5] Subeucalanus monachus (Giesbrecht, 1888) (F,M) [Figs] | |
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| | [6] Subeucalanus mucronatus (Giesbrecht, 1888) (F,M) [Figs] | |
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| | [7] Subeucalanus pileatus (Giesbrecht, 1888) (F,M) [Figs] | |
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| | [8] Subeucalanus subcrassus (Giesbrecht, 1888) (F,M) [Figs] | |
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| | [9] Subeucalanus subtenuis (Giesbrecht, 1888) (F,M) [Figs] | |
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| | Subeucalanus subtenuis japonica Fukase, 1957 (F,M) | | Syn.: | Eucalanus subtenuis japonica Fukase, 1957 (p.19, figs.F,M, Rem.); Tanaka, 1960 (p.21, Rem.); Grice, 1962 (p.181, figs.F,M, Rem.); Vervoort, 1963 b (p.92, Rem.); Eucalanus subtenuis : Tanaka, 1935 a ; Eucalanus mucronatus : Mori, 1937 (1964) (p.25, figs.F,M); Chihara & Murano, 1997 (p.789, Pl.99 part.,101: F,M); | | Loc: | Mer de Chine E, Japon, Pacif. équatorial | | Lg.: | (91) F: ± 3; M: 2,9-2,7; (101) F: 3,12-2,4; M: 2,52-2,45; {F: 2,40-3,12; M: 2,45-2,90} |
| | [10] Subeucalanus sp. Goetze & Ohman, 2010 [Figs] | |
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| | [11] Subeucalanus sp.2 Goetze & Ohman, 2010 [Figs] | |
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| | [12] Subeucalanus sp.3 Goetze & Ohman, 2010 [Figs] | |
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 Any use of this site for a publication will be mentioned with the following reference :
Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2024. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed April 26, 2024] © copyright 2005-2024 Sorbonne University, CNRS
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