| Heterorhabdidae Sars, 1902 ( Arietelloidea ) | |
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| Rem.: | Boxshall & Halsey (2004 (p.68, 126) transfer this genus to the Augaptilidae, however with some doubt as long as the male has not been encountered. |
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| (1) Disseta Giesbrecht, 1889 | |
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| Rem.: | The species of this genus appear to feed mainly on particles by suspension feeding. |
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| Disseta atlantica Wolfenden, 1911 (F,M) |
| Ref.: | Wolfenden, 1911 (p.313, Descr.F,M); Vervoort, 1965 (p.116) |
| Loc: | Atlant.(équat.), Is. Cap Vert E & NE, off W Afr. S |
| Rem.: | Cf. Disseta palumboi |
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| Disseta coelebs Heptner, 1972 (M) |
| Ref.: | Heptner, 1972 (p.1645, Descr.M, figs.M) |
| Loc: | Pacif. W (Fosse de Bougainville) |
| Rem.: | Cf.: Disseta magna |
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| Disseta grandis Esterly, 1906 (F,M) |
| Ref.: | Esterly, 1906 a (p.72, figs.F,M); Sewell, 1947 (p.185, 189: Rem.); Brodsky, 1950 (1967) (p.343, figs.F,M) |
| Loc: | Californie (San Diego) |
| N: | 1 |
| Lg.: | (17) F: 8,3; M: 7,6 |
| Rem.: | Cf. Disseta palumbii. Sewell (1947, p.189) estime que la femelle est synonyme de D. palumboi ce que ne confirme pas Vervoort (1965, p.116). Park (2000, p.13, 17-18) en fait un synonyme junior de D. palumbii. |
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| [1] Disseta magna Bradford, 1971 (F,M) [Figs] | |
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| Disseta maxima Esterly, 1911 (F) |
| Ref.: | Esterly, 1911 (p.330, figs.F); Sewell, 1947 (Rem.: p.189); C.B. Wilson, 1950 (p.198);Tanaka, 1964 a (p.34, 37, Rem.) |
| Loc: | San Diego |
| Lg.: | (11) F: 9,4 |
| Rem.: | Cf. Disseta scopularis |
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| Disseta minuta Grice & Hulsemann, 1965 (F,M) |
| Ref.: | Grice & Hulsemann, 1965 (p.245, figs.F,M); Heptner, 1972 a (p.60, Rem.) |
| Loc: | Atlant. NE |
| Rem.: | Cf. Microdisseta minuta |
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| [2] Disseta palumbii Giesbrecht, 1889 (F,M) [Figs] | |
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| [3] Disseta scopularis (Brady, 1883) (F,M) [Figs] | |
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| Disseta sp. Esterly, 1911 (M) |
| Ref.: | Esterly, 1911 (p.331, Descr.M, figs.M); Sewell, 1947 (p.185, 189, Rem.); Tanaka, 1964 a (p.32) |
| Loc: | San Diego |
| Rem.: | Cf. Disseta palumbii |
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| (2) Hemirhabdus Wolfenden, 1911 | |
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| Rem.: | type: Hemirhabdus grimaldii. 2 spp.: |
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| Hemirhabdus falciformis Wolfenden, 1911 (F) |
| Ref.: | Wolfenden, 1911 (p.310, Descr.F, figs.F); Heptner, 1972 a (p.59) |
| Loc: | Atlant. tropical S |
| Rem.: | Cf. Neorhabdus falciformis |
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| [1] Hemirhabdus amplus Park, 2000 (F,M) [Figs] | |
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| [2] Hemirhabdus grimaldii (Richard, 1893) (F,M) [Figs] | |
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| Hemirhabdus latus Sars, 1905 (F,M) |
| Syn.: | Heterorhabdus latus Sars, 1905 c (p.9, Rem.F); 1920 c (p.11) |
| Ref.: | Sars, 1925 (p.232, figs.F,M); Sewell, 1932 (p.306, 307, Rem.); C.B. Wilson, 1950 (p.238, Rem.); Heptner, 1972 a (p.58) |
| Loc: | Canaries |
| Rem.: | Cf. Neorhabdus latus |
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| Hemirhabdus truncatus A. Scott, 1909 (F) |
| Syn.: | Mesorhabdus truncatus A. Scott, 1909 (p.132, Descr.F, figs.F); ? Heterorhabdus latus Sars, 1905 c (p.9); 1920 c (p.11) |
| Ref.: | Heptner, 1972 a (p.58) |
| Loc: | Mer des Moluques |
| Rem.: | Cf. Neorhabdus latus |
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| (3) Heterorhabdus Giesbrecht, 1898 | |
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| Rem.: | type: Heterorhabdus spinifrons.
For Vervoort, 1951 (p.140) thie genus Alloiorhabdus is based exclusively on the number of setae on the various segments of the swimming legs. It appears, however, that wolfenden’s observations are inaccurate as no differences in this respect exists between Heterorhabdus stricto sensu and the proposed genus Alloiorhabdus. The structure of the oral appendages in these genera too does not justify such a distinction.
Brodsky , 1950 (1967, p.345) establishes the 2 subgenera : Heterorhabdus (type: H. robustus) et Euheterorhabdus (types : H. papilliger & H. norvegicus.
Park establishes 4 groups ("spinifrons", "papilliger", "fistulosus", "abyssalis".
31 spp. (+ 4 doubtful).
The species of this genus appear typical carnivores preying upon copepodids, polychaetes, and othe rmacrozooplanktoners (see litt. in Ohtsuka & al., 1997). |
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| Ref.: | Park, 2000 (p.90, Déf., clé des spp.) |
| Rem.: | 7 spp.: |
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| [1] Heterorhabdus ankylocolus Park, 2000 (F,M) [Figs] | |
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| [2] Heterorhabdus caribbeanensis Park, 1970 (F,M) [Figs] | |
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| [3] Heterorhabdus heterolobus Park, 2000 (F,M) [Figs] | |
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| [4] Heterorhabdus insukae Park, 2000 (F,M) [Figs] | |
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| [5] Heterorhabdus quadrilobus Park, 2000 (F,M) [Figs] | |
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| [6] Heterorhabdus spinifrons (Claus, 1863) (F,M) [Figs] | |
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| [7] Heterorhabdus subspinifrons Tanaka, 1964 (F,M) [Figs] | |
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| Ref.: | Park, 2000 (p.102, Déf., clé spp.) |
| Rem.: | 5 spp.: |
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| [1] Heterorhabdus guineanensis Park, 2000 (F,M) [Figs] | |
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| [2] Heterorhabdus lobatus Bradford, 1971 (F,M) [Figs] | |
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| [3] Heterorhabdus papilliger Claus, 1863 (F,M) [Figs] | |
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| [4] Heterorhabdus prolatus Park, 2000 (F,M) [Figs] | |
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| [5] Heterorhabdus spinifer Park, 1970 (F,M) [Figs] | |
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| Ref.: | Park, 2000 (p.110, Déf., clé spp.) |
| Rem.: | 2 spp.: |
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| [1] Heterorhabdus egregius Heptner, 1972 (F,M) [Figs] | |
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| [2] Heterorhabdus fistulosus Tanaka, 1964 (F,M) [Figs] | |
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| Ref.: | Park, 2000 (p.114, Déf., clé spp.) |
| Rem.: | 17 spp.: |
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| [1] Heterorhabdus abyssalis (Giesbrecht, 1889) (F,M) [Figs] | |
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| [2] Heterorhabdus americanus Park, 2000 (F,M) [Figs] | |
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| [3] Heterorhabdus austrinus Giesbrecht, 1902 (F,M) [Figs] | |
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| Heterorhabdus austrinus var. Farran, 1929 (F,M) |
| Ref.: | Farran, 1929 (p.265: petites formes); Vervoort, 1957 (p.132, Rem.) |
| Loc: | Antarct., sub-Antarct., off Nouvelle-Zélande E |
| Lg.: | (35) F: 3,18-2,97; M: 2,88-2,84 |
| Rem.: | Pour Vervoort (1957) on n’observe pas de différence structurelle entre cette variété et l’espèce précédente. |
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| [4] Heterorhabdus clausi (Giesbrecht, 1889) (F,M) [Figs] | |
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| [5] Heterorhabdus cohibilis Park, 2000 (F,M) [Figs] | |
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| [6] Heterorhabdus confusibilis Park, 2000 (F,M) [Figs] | |
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| [7] Heterorhabdus habrosomus Park, 2000 (F,M) [Figs] | |
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| [8] Heterorhabdus longisegmentus Park, 2000 (F,M) [Figs] | |
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| [9] Heterorhabdus norvegicus (Boeck, 1872) (F,M) [Figs] | |
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| [10] Heterorhabdus oikoumenikis Park, 2000 (F,M) [Figs] | |
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| [11] Heterorhabdus pacificus Brodsky, 1950 (F,M) [Figs] | |
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| [12] Heterorhabdus paraspinosus Park, 2000 (F,M) [Figs] | |
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| [13] Heterorhabdus prolixus Park, 2000 (F,M) [Figs] | |
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| [14] Heterorhabdus pustulifer Farran, 1929 (F,M) [Figs] | |
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| [15] Heterorhabdus spinosus Bradford, 1971 (F,M) [Figs] | |
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| [16] Heterorhabdus tanneri (Giesbrecht, 1895) (F,M) [Figs] | |
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| [17] Heterorhabdus tuberculus Park, 2000 (F,M) [Figs] | |
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| Espèces citées comme Heterorhabdus en synonymie ou douteuses: |
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| [1] Heterorhabdus atlanticus Wolfenden, 1905 (M) | |
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| Heterorhabdus brevicaudatus Wolfenden, 1905 (F) |
| Ref.: | Wolfenden, 1905 a (p.12, Descr.F, fig.F) |
| Rem.: | Cf. Mesorhabdus brevicaudatus |
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| [2] Heterorhabdus brevicornis (F. Dahl, 1894) (F) | |
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| Heterorhabdus compactoides Heptner, 1971 (F,M) |
| Ref.: | Heptner, 1971 (p.152, figs.F,M); 1972 a (p.59) |
| Rem.: | Cf. Paraheterorhabdus (Paraheterorhabdus) compactoides |
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| Heterorhabdus compactus Sars, 1900 (F,M) |
| Ref.: | Heterochaeta compacta Sars, 1900 (p.83, Descr.F, figs.F) |
| Rem.: | Cf. Paraheterorhabdus (Antirhabdus) compactus |
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| Heterorhabdus farrani Brady, 1918 (F,M) |
| Ref.: | Brady, 1918 (p.27, figs.F,M) |
| Rem.: | Cf. Paraheterorhabdus (Paraheterorhabdus) farrani |
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| Heterorhabdus fistulosus tuberculata Heptner, 1971 (F) |
| Ref.: | Heptner, 1971 (p.149, figs.F) |
| Rem.: | Cf. Heterorhabdus fistulosus |
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| Heterorhabdus fistulosus typica Heptner, 1971 (F,M) |
| Ref.: | Heptner, 1971 (p.149, figs.F,M) |
| Rem.: | Cf. Heterorhabdus fistulosus |
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| Heterorhabdus grandis Wolfenden, 1904 (F,M) |
| Ref.: | Wolfenden, 1904 (p.120, fig.F); 1905 a (p.8, Descr.F,M, fig.M) |
| Loc: | Atlant. NE |
| Rem.: | Cf. Disseta palumbii |
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| Heterorhabdus grimaldii Richard, 1893 (F,M) |
| Syn.: | Heterochaeta grimaldii Richard, 1893 a (p.151, Descr.F) |
| Ref.: | Giesbrecht & Schmeil, 1898 (p.117, Rem.); Wolfenden,1905 a (p.10, Rem.F,M); Sewell, 1913 (p.354) |
| Loc: | Canaries, Açores N, Irlande, Indien (G. du Bengale) |
| Rem.: | Cf. Hemirhabdus grimaldii |
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| Heterorhabdus latus Sars, 1905 (F) |
| Ref.: | Sars, 1905 c (p.9, Rem.F); Heptner, 1972 a (p.58) |
| Loc: | Canaries |
| Rem.: | Cf. Neorhabdus latus |
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| Heterorhabdus longicornis Giesbrecht, 1889 (F,M) |
| Syn.: | Heterochäta longicornis Giesbrecht,1889; 1892 (p.373, figs.F) |
| Ref.: | Giesbrecht & Schmeil, 1898 (p.116) |
| Rem.: | Cf. Heterostylites longicornis |
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| Heterorhabdus longispinus Davis, 1949 (M) |
| Ref.: | Davis, 1949 (p.58, Descr.M, figs.M) |
| Loc: | Pacif. NE (off C. Flattery) |
| Rem.: | Cf. Paraheterorhabdus (Paraheterorhabdus) longispinus |
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| Heterorhabdus major F. Dahl, 1894 (M) |
| Syn.: | Heterochaeta major F. Dahl,1894 |
| Ref.: | Giesbrecht & Schmeil, 1898 (p.117) |
| Rem.: | Cf. Heterostylites major |
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| Heterorhabdus medianus Park, 1970 (F,M) |
| Ref.: | Park, 1970 (p.477, 525, figs.F, non M) |
| Rem.: | Cf. Paraheterorhabdus (Paraheterorhabdus) medianus |
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| Heterorhabdus nigrotinctus Brady, 1918 (M) |
| Ref.: | Brady, 1918 (p.27, Rem.M, figs.M) |
| Rem.: | Cf. Heterostylites nigrotinctus. |
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| Heterorhabdus pilosus Tanaka, 1953 |
| Ref.: | Tanaka, 1953 (p.134, nom.nud.); 1964 a (p.11) |
| Rem.: | Cf. Heterorhabdus tanneri |
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| [3] Heterorhabdus profundus (F. Dahl, 1894) (F,M) [Figs] | |
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| Heterorhabdus proximus Davis, 1949 (F,M) |
| Ref.: | Davis, 1949 (p.57, figs.F,M); non Bradford, 1971 a (p.126, figs.F,M) |
| Rem.: | Cf. Heterorhabdus tanneri |
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| Heterorhabdus robustoides Brodsky, 1950 (F,M) |
| Ref.: | Brodsky, 1950 (1967) (p.346, figs.F,M); Tanaka, 1964 a (p.20, Rem.); Vervoort, 1965 (p.118, Rem.); Heptner, 1971 (p.156, 158, Rem.); Gardner & Szabo, 1982 (p.368, figs.F,M) |
| Loc: | Pacif. NW, Mer de Béring |
| Rem.: | Cf. Heterorhabdus robustus |
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| Heterorhabdus robustus Farran, 1908 (F,M) |
| Ref.: | Farran, 1908 b (p.65, figs.F,M, Rem.) |
| Rem.: | Cf.: Paraheterorhabdus (Paraheterorhabdus) robustus. |
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| Heterorhabdus tenuis Tanaka, 1964 (M) |
| Ref.: | Tanaka, 1964 a (p.20, Descr.M, figs.M) |
| Rem.: | Cf. Paraheterorhabdus (Paraheterorhabdus) vipera |
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| [4] Heterorhabdus tropicus (F. Dahl, 1894) (F,M) | |
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| Heterorhabdus vipera Giesbrecht, 1889 (F,M) |
| Syn.: | Heterochäta vipera Giesbrecht,1889; 1892 (p.373, 383, figs.F, non M); |
| Ref.: | Giesbrecht & Schmeil, 1898 (p.116) |
| Rem.: | Cf. Paraheterorhabdus (Paraheterorhabdus) vipera |
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| Heterorhabdus zetesios Wolfenden, 1902 (M) |
| Syn.: | Heterochaeta zetesios Wolfenden, 1902 (p.367, Descr.M); Fowler, 1903 (p.124) |
| Ref.: | Vervoort, 1957 (p.135) |
| Loc: | Féroé |
| Rem.: | Cf. Heterostylites major |
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| (4) Heterostylites Sars, 1920 | |
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| Rem.: | The species of this genus appear to feed mainly on particles by suspension feeding, additionally feeding on zooplankters, such as copepods (see in Harding, 1974 and Ohtsuka & al., 1997). |
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| [1] Heterostylites echinatus Park, 2000 (F,M) [Figs] | |
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| [2] Heterostylites longicornis (Giesbrecht, 1889) (F,M) [Figs] | |
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| [3] Heterostylites longioperculis Park, 2000 (F,M) [Figs] | |
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| [4] Heterostylites major (F. Dahl, 1894) (F,M) [Figs] | |
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| [5] Heterostylites nigrotinctus (Brady, 1918) (F,M) [Figs] | |
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| [6] Heterostylites submajor Park, 2000 (F,M) [Figs] | |
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| (5) Mesorhabdus Sars, 1905 | |
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| Rem.: | type: Mesorhabdus annectens (= Heterorhabdus brevicaudatus). 5 spp.: |
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| [1] Mesorhabdus angustus Sars, 1907 (F,M) [Figs] | |
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| Mesorhabdus annectens Sars, 1905 (F) |
| Ref.: | Sars, 1905 c (p.9, Rem.F) |
| Loc: | Açores |
| Rem.: | Cf. Mesorhabdus brevicaudatus |
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| [2] Mesorhabdus brevicaudatus (Wolfenden, 1905) (F,M) [Figs] | |
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| [3] Mesorhabdus gracilis Sars, 1907 (F,M) [Figs] | |
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| [4] Mesorhabdus paragracilis Park, 2000 (F,M) [Figs] | |
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| [5] Mesorhabdus poriphorus Park, 2000 (F,M) [Figs] | |
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| Mesorhabdus truncatus A. Scott, 1909 (F) |
| Ref.: | A. Scott, 1909 (p.132, Descr.F, figs.F); Heptner, 1972 a (p.58) |
| Loc: | Mer des Moluques |
| Rem.: | Cf. Neorhabdus latus |
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| (6) Microdisseta Heptner, 1972 | |
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| Rem.: | Boxshall & Halsey (2004, p.126) do not follow the opinion of Park on the uncertainty of the place of this genus within this family. 1 sp.
The feeding habits of species are still enigmatic, but probably mainly employs suspension feeding on the basis of their similarities in the mouthpart structures to those of Disseta and Heterostylites (see the fine structures of Md and labrums in Ohtsuka & al., 1997). |
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| [1] Microdisseta minuta (Grice & Hulsemann, 1965) (F,M) [Figs] | |
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| (7) Neorhabdus Heptner, 1972 | |
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| Rem.: | type: Neorhabdus latus. 5 spp.: |
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| [1] Neorhabdus brevicornis Park, 2000 (F,M) [Figs] | |
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| [2] Neorhabdus capitaneus Park, 2000 (F,M) [Figs] | |
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| [3] Neorhabdus falciformis (Wolfenden, 1911) (F,M) [Figs] | |
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| [4] Neorhabdus latus (Sars, 1905) (F,M) [Figs] | |
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| [5] Neorhabdus subcapitaneus Park, 2000 (F) [Figs] | |
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| Neorhabdus truncatus A. Scott, 1909 (F) |
| Syn.: | Mesorhabdus truncatus A. Scott, 1909 (p.132, Descr.F, figs.F); ? Hemirhabdus latus Sars, 1925 (p.232, figs.F,M); C.B. Wilson, 1950 (p.238); non Hemirhabdus truncatus : Sewell, 1932 (p.306, figs.F, Rem.); 1947 (p.181); 1948 (p.330, 530, 533, 547) |
| Ref.: | Heptner,1972 a (p.58); Park, 1999 (p.58: Rem., 59) |
| Loc: | Mer Arabe, Is. Laquedives, Inde S, Mer des Moluques |
| N: | 3 |
| Lg.: | (5) F: 7; (11) F: 9,667; (44) F: 7,44 |
| Rem.: | Cf. Neorhabdus latus |
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| (8) Paraheterorhabdus Brodsky, 1950 | |
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| [1] Paraheterorhabdus (Antirhabdus) compactus (Sars, 1900) (F,M) [Figs] | |
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| [2] Paraheterorhabdus (Paraheterorhabdus) compactoides (Heptner, 1971) (F,M) [Figs] | |
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| [3] Paraheterorhabdus (Paraheterorhabdus) farrani (Brady, 1918) (F,M) [Figs] | |
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| [4] Paraheterorhabdus (Paraheterorhabdus) illgi Park, 2000 (F,M) [Figs] | |
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| [5] Paraheterorhabdus (Paraheterorhabdus) longispinus (Davis, 1949) (F,M) [Figs] | |
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| [6] Paraheterorhabdus (Paraheterorhabdus) medianus (Park, 1970) (F,M) [Figs] | |
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| [7] Paraheterorhabdus (Paraheterorhabdus) robustus (Farran, 1908) (F,M) [Figs] | |
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| [8] Paraheterorhabdus (Paraheterorhabdus) vipera (Giesbrecht, 1889) (F,M) [Figs] | |
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