List species and varieties by family
Oncaeidae Giesbrecht, 1893 ["1892"] ( Cyclopoida )
(1) Archioncaea Böttger-Schnack & Huys, 1997
Rem.: Total: 1 sp.
For Böttger-Schnack & Schnack (2013, p.18) the genus has a unique leg armature and combines primitive and advanced characters, which separates this taxon from all other known oncaeids. Most remarkable is the trisetose P5 exopod. On all four swimming legs, the middle and distal segments of the endopods are completely fused and the leg armature (Table 5, type 4) is characterized by the absence of an outer subdistal spine on the fused endopodal segments of P3 and P4 and showing only 2 outer spines on exopodite 3 of P2.

[1] Archioncaea arabica  Böttger-Schnack & Huys, 1997   (F)    [Figs]
(2) Conaea Giesbrecht, 1891
Rem.: type: Conäa rapax Giesbrecht,1891. 4 spp.
For Böttger-Schnack & Schnack (2013, p.19) within Conaea s.L. two subgroups can be distinguished (Table 3): the type species of the genus, C. rapax, (defined as subgroup Conaea s.str.) differs from all other Conaea species (difined as expressa-subgroup) by a different endopodal spine count on P2 (Table 5, type 3), and is also separated by details of the lateral elements on enddopodite 2 of A2, especially the lack of the typical Conaea-like spinulose ornamentation of antennary elements and a very small seta IV. Further differences are found in the proportional lengths of the urosomites.

[1] Conaea expressa  (Gordeeva, 1973)   (F,M)    [Figs]

Conaea gracilis    Dana, 1849   (F,M)
Syn.: Antaria gracilis Dana, 1849; 1852; ? non Conäa rapax Giesbrecht, 1891; 1892 (p.605); ? non Oncaea gracilis T. Scott, 1894 b (p.116, figs.F); Malt, 1982 (p.193); 1983 a (p.7); non Conaea gracilis : A. Scott, 1909 (p.244); Wilson, 1942 a (p.178); 1950 (p.191, figs.F,M); Owre & Foyo, 1964 a (p.369); 1967 (p.112); Boxshall, 1977 a (p.145); 1977 b (p.549)
Ref.: De Decker & Mombeck, 1964 (p.12, 18); Owre & Foyo, 1967 (p.112, figs.F,M); De Decker, 1968 (p.45); Binet & Dessier, 1968 (p.7); Furuhashi, 1966 (p.313); Deevey, 1971 (p.224); Heron, 1977 (p.86, 90, Rem.); Deevey & Brooks, 1977 (tab.2); Frontier, 1977 a (p.17); Vives, 1982 (p.296); Rudyakov, 1982 (p.209)
Loc: Afr. S (E & W), off Mauritanie, off Bermuda, Madagascar (Nosy Bé), Indien SW, Japon, Kuroshio zone
Lg.: (139) F: 1,05-0,88; M: 0,86-0,76; {F: 0,88-1,05; M: 0,76-0,86}
Rem.: espèce mal décrite. Voir remarques à Conaea rapax. ? Cf. Conaea rapax ou Oncaea gracilis

[2] Conaea hispida  Heron, 1977   (F,M)    [Figs]

[3] Conaea rapax  Giesbrecht, 1891   (F,M)    [Figs]

[4] Conaea succurva  Heron, 1977   (F)    [Figs]
(3) Epicalymma Heron, 1977
Rem.: Malt (1982, p.185) regards this genus as a synonym of Oncaea, contrary to Heron & al. (1984) and Heron & Bradford-Grieve (1995).
Type: Epicalymma schmitti. 7 spp.:
For Böttger-Schnack & Schnack, (2013, p.19) within the genus, one species, E. umbonata, may deserve a separate group status, based on morphological grounds (labral structure, antennary and maxillipedal elements) as discussed by Böttger-Schnack (2009), but not shown in detail in the present study. hence, the species has been subsumed under Epicalymma S.l. (in Table 3, p.12) to denote the potential existence of two lineages within this genus.

[1] Epicalymma ancora  (Gordeeva, 1973)   (F)    [Figs]

[2] Epicalymma brittoni  Heron, English & Damkaer, 1984   (F)    [Figs]

[3] Epicalymma bulbosa  Böttger-Schnack, 2009   (F,M)    [Figs]

[4] Epicalymma exigua  (Farran, 1908)   (F,M)    [Figs]

[5] Epicalymma schmitti  Heron, 1977   (F,M)    [Figs]

[6] Epicalymma umbonata  Heron, 1977   (F,M)    [Figs]

[7] Epicalymma vervoorti  Heron, English & Damkaer, 1984   (F,M)    [Figs]
(4) Monothula Böttger-Schnack & Huys, 2001
Rem.: type: Oncaea subtilis Giesbrecht,1892. Total: 1 sp.

[1] Monothula subtilis  (Giesbrecht, 1892)   (F,M)    [Figs]
Myspictosum Kazatchenko & Andreev, 1977
Rem.: Cf. Oncaea
(5) Oncaea Philippi, 1843
Rem.: Malt (1982, p.191, 192) regards the 2 genera Conaea and Epicalymma as synonyms of Oncaea, whereas Heron & al. (1984) and Heron & Bradford-Grieve (1995) believe that they should be maintained.
Type: Oncaea venusta Philippi,1843. Total: 71 spp. + 1 unidentified.:

After Ohtsuka & Nishida (2017, p.582), Oncaea abd related genera are associated with discarded appendicularian houses, feeding on plankters retained on the appendicularian feeding nets (Ohtsuka & Kubo, 1991; Ohtsuka & &l., 1993; 1996a).

[1] Oncaea africana  Shmeleva, 1979   (F)    [Figs]

[2] Oncaea alboranica  Shmeleva, 1979   (F)    [Figs]

Oncaea ancora    Gordeeva, 1973   (F)
Rem.: Cf. Epicalymma ancora

[3] Oncaea anglica  Brady, 1905   (F)
Rem.: species doubtful.

Oncaea antarctica    Heron, 1977   
Ref.: Heron, 1977 (p.42, figs.F,M)
Rem.: Cf. Triconia antarctica

[4] Oncaea atlantica  Shmeleva, 1967   (F,M)    [Figs]

[5] Oncaea bathyalis  Shmeleva, 1968   (F)    [Figs]

[6] Oncaea bispinosa  Böttger-Schnack, 2002   (F,M)    [Figs]

Oncaea borealis    Sars, 1918   (F,M)
Syn.: Oncaea conifera : Sars, 1900 (p.113, figs.F)
Ref.: Sars, 1918 (p.191, figs.F,M)
Rem.: Cf. Triconia borealis

[7] Oncaea bowmani  Heron, 1977   (F)    [Figs]

[8] Oncaea brocha  Heron, 1977   (F,M)    [Figs]

[9] Oncaea brodskii  Shmeleva, 1968   (F,M)    [Figs]

Oncaea canadensis    Heron & Frost, 2000   (F,M)
Ref.: Heron & Frost, 2000 (p.1019, figs.F,M, tab.2, 3)
Ref. compl.: Nishibe & Ikeda, 2004 (Rem. p.933)
Rem.: Cf. Triconia canadensis

[10] Oncaea clevei  Früchtl, 1923   (F,M)    [Figs]

[11] Oncaea compacta  Heron, 1977   (F,M)    [Figs]

Oncaea conifera    Giesbrecht, 1891   (F,M)
Syn.: Oncäa conifera Giesbrecht, 1891; 1892 (p.591, 603, 774, figs.F,M); non O. conifera : Sars, 1900 (p.113); non Oncaea conifera var. III Giesbrecht, 1902 (p.41, Pl.13, figs.7-11); non O. conifera : Tanaka, 1960 (p.66, figs.F,M); B.D. Lee, 1966; non Oncaea conifera : Farran, 1936 a (p.127, figs.F: formes b & c); non O. conifera : Moulton, 1973 (p.147 & suiv.:'minus');
Rem.: Cf. Triconia conifera

Oncaea conifera furcula    Farran, 1936   (F,M)
Ref.: Farran, 1936 a (p.127, figs.F); Sewell, 1947 (p.260, Rem.); Gardner & Szabo, 1982 (p.116, figs.F,M); Malt, 1983 (p.457)
Rem.: Cf. Triconia furcula

[12] Oncaea convexa  Heron, 1977   (F)    [Figs]

[13] Oncaea cristata  Böttger-Schnack, 2005   (F,M)    [Figs]

[14] Oncaea crypta  Böttger-Schnack, 2005   (F,M)    [Figs]

[15] Oncaea curta  Sars, 1916   (F,M)    [Figs]

[16] Oncaea curvata  Giesbrecht, 1902   (F,M)    [Figs]

[17] Oncaea damkaeri  Heron, 1977   (F,M)    [Figs]

[18] Oncaea delicata  Heron, English & Damkaer, 1984   (F,M)    [Figs]

Oncaea dentipes    Giesbrecht, 1891   (F,M)
Syn.: Oncäa dentipes Giesbrecht, 1891; 1892 (p.591, 603, 774, Descr.F, figs.F)
Rem.: Cf. Triconia dentipes

Oncaea derivata    Heron & Bradford-Grieve, 1995   (F,M)
Syn.: Oncaea conifera : Moulton, 1973 (p.142 & suiv.: 'bumped')
Ref.: Heron & Bradford-Grieve, 1995 (p.25, Descr.F, figs.F); Heron & Frost, 2000 (p.1028, Descr.M, figs.F,M, tab.2, 3)
Rem.: Cf. Triconia derivata

[19] Oncaea englishi  Heron, 1977   (F,M)    [Figs]

Oncaea exigua    Farran, 1908   (F,M)
Ref.: Farran, 1908 b (p.93); Malt, 1983 a (p.7, figs.F,M, Rem.); Greze & al., 1985 (p.8)
Rem.: Cf. Epicalymma exigua

Oncaea expressa    Gordeeva, 1973   (F,M)
Ref.: Gordeeva, 1973 (p.1573, figs.F,M); Malt, 1982 (p.192, 193)
Loc: off I. Ste-Hélène W
Rem.: Cf. Conaea expressa

[20] Oncaea frosti  Heron, 2002   (F,M)    [Figs]

Oncaea furcula    Farran, 1936   (F,M)
Syn.: Oncaea conifera var. furcula Farran, 1936 a (p.127, figs.F: forme b); Bradford, 1972 (part., p.50); Oncaea conifera : Moulton, 1973 (p.142 & suiv.: 'Long' form); Oncaea furcula : Heron & Bradford-Grieve, 1995 (p.24, Redescr.F,M, figs.F,M)
Rem.: Cf. Triconia furcula

[21] Oncaea furnestini  Shmeleva, 1979   (F)    [Figs]

[22] Oncaea glabra  Heron & Frost, 2000   (F,M)    [Figs]

Oncaea gracilis    Dana, 1849   (F)
Syn.: Antaria gracilis Dana,1849; Giesbrecht, 1892 (p.591: sp. douteuse, 592)
Ref.: Malt, 1983 a (p.7, figs.F,M, Rem.); Böttger-Schnack, 1994 (p.277); Chihara & Murano, 1997 (p.980, Pl.221: F,M)
Rem.: La synonymie entre Conaea rapax Giesbrecht,1891 et cette forme insuffisamment décrite n'est pas admise par tous les auteurs. Giesbrecht (1892, p.592) estime qu'il s'agit d'une variété de Oncaea venusta . Voir remarques à Conaea gracilis et C. rapax

[23] Oncaea grossa  Heron & Frost, 2000   (F,M)    [Figs]

Oncaea hawii    Böttger-Schnack & Boxshall, 1990   (F,M)
Syn.: Oncaea sp. Böttger-Schnack, 1988; Böttger-Schnack & Boxshall, 1990 (p.865, Descr.F, figs.F, tab.III)
Rem.: Cf. Triconia hawii

[24] Oncaea heronae  Malt, 1982   (F)    [Figs]

Oncaea hispida    Malt, 1982   (F,M)
Ref.: Malt, 1982 (p.187, figs.F,M)
Loc: off NE Is. du Cap Vert
Lg.: (680) F: 0,56; M: 0,55
Rem.: Heron & Bradford-Grieve (1995, p.46) ne considèrent pas la forme décrite par Malt (1982) comme synonyme de Conaea hispida Heron, 1977

[25] Oncaea illgi  Heron, 1977   (F,M)    [Figs]

[26] Oncaea infantula  Gordejeva, 1972   (F,M)    [Figs]

Oncaea inflexa    Heron, 1977   (F,M)
Ref.: Heron, 1977 (p.45, figs.F,M); Malt,1983 (p.458, Rem.,); Heron & al., 1984 (p.465); Heron & Bradford-Grieve, 1995 (p.15, figs.F,M)
Rem.: Cf. Triconia inflexa

[27] Oncaea insolita  Heron & Frost, 2000   (F,M)    [Figs]

Oncaea ivlevi    Shmeleva, 1966   (F,M)
Ref.: Shmeleva, 1966 (p.932, figs.F,M); Böttger Schnack, 2003 (p.192, Rem.)
Rem.: Cf. Spinoncaea ivlevi.

Oncaea ivlevi 1       
Ref.: Böttger-Schnack, Hagen & Schnack-Shiel,2001 (p.1029, tab.1,2)
Loc: G. d'Aqaba, Mer Rouge (N-S)

Oncaea ivlevi 2       
Ref.: Böttger-Schnack, Hagen & Schnack-Shiel,2001 (p.1029, tab.1,2)
Loc: Mer Rouge (N-S)

[28] Oncaea lacinia  Heron, English & Damkaer, 1984   (F,M)    [Figs]

[29] Oncaea latimana  Gordejeva, 1975   (F,M)    [Figs]

[30] Oncaea longipes  Shmeleva, 1968   (F,M)    [Figs]

[31] Oncaea longiseta  Shmeleva, 1968   (F,M)    [Figs]

[32] Oncaea macilenta  Heron, 1977   (F,M)    [Figs]

[33] Oncaea media  Giesbrecht, 1891   (F,M)    [Figs]

Oncaea media major    Sewell, 1947   (F,M)
Ref.: Sewell, 1947 (p.261, Rem.F); 1951 (p.375, Rem.); Malt, 1983 a (p.5, 7); Böttger-Schnack & al., 1989 (p.1089); 1995 (p.92) 1997 (p.409); Ohtsuka & al., 1996 a (p.91); Suarez-Morales & Gasca, 1998 a (p.112)
Ref. compl.: Lakkis & al., 2005 (p.152)
Lg.: (11) F: 0,817-0,733; (46) F: 0,82-0,55; (180) F: 0,76-0,7; (788) F: 0,77-0,65; {F: 0,55-0,82}

Oncaea media minor    Sewell, 1947   (F,M)
Syn.: ? Oncaea sp.1 Ferrari, 1975 (p.228, figs.)
Ref.: Sewell, 1947 (p.261, Rem.F); 1951 (p.375, Rem.); Malt,1983 a (p.5, 7); Böttger-Schnack & al., 1989 (p.1089); Huys & Boxshall, 1991 (p.292, 304, 311, figs.F,M); Böttger-Schnack, 1992 (p.304); Ohtsuka & al., 1996 a (p.91); Böttger-Schnack, 1997 (p.409); Boxshall & Halsey, 2004 (p.614: fig.M)
Lg.: (11) F: 0,65-0,58; M: 0,683-0,533; (180) F: 0,65-0,61 (non ! 1,65- 1,61); (788) F: 0,52-0,48; M: 0,44-0,41; {F: 0,48-0,65; M: 0,41-0,68}
Rem.: souvent confondu avec Oncaea scottodicarloi.

[34] Oncaea mediterranea  (Claus, 1863)   (F,M)    [Figs]

[35] Oncaea memorata  Gordejeva, 1973   (F,M)    [Figs]

[36] Oncaea minima  Shmeleva, 1968   (F,M)    [Figs]

[37] Oncaea minor  Shmeleva, 1979   (M)    [Figs]

Oncaea minuta    Giesbrecht, 1892   
Syn.: Oncäa minuta Giesbrecht, 1892 (p.591, 603, 774, Descr.F, figs.F);
Rem.: Cf. Triconia minuta

[38] Oncaea mollicula  Gordejeva, 1975   (F,M)    [Figs]

[39] Oncaea neobscura  Razouls, 1969   (F)    [Figs]
Rem.: Malt, 1982 a (p. 140) considers that it concerns a juvenile stage 5 of a male.

[40] Oncaea notopus  Giesbrecht, 1891   (F, ?M)    [Figs]

Oncaea obtusa    Dana, 1852   (F,M)
Ref.: Brady, 1883 (p.120, figs.F,M); Grindley & Lane, 1979 (p.113); Kovalev & Shmeleva, 1982 (p.86)
Lg.: (135) F: 1,3
Rem.: Cf. Oncaea venusta

[41] Oncaea obscura  Farran, 1908   (F,M)    [Figs]
Rem.: Malt (1982 a) believes that it concerns male juvenile forms (stage 5).
After Shiganova & al. (2012, p.99) this species new in the Black Sea (as other species) arise from a rise of the water surface layer temperature and/or increasing of shipping activity during the last decades.

[42] Oncaea oceanica  Gordeeva, 1972   (F,M)    [Figs]

[43] Oncaea olsoni  Heron, 1977   (F,M)    [Figs]

[44] Oncaea ornata  Giesbrecht, 1891   (F,M)    [Figs]

[45] Oncaea ovalis  Shmeleva, 1966   (F,M)    [Figs]

[46] Oncaea parabathyalis  Böttger-Schnack, 2005   (F,M)    [Figs]

[47] Oncaea paraclevei  Böttger-Schnack, 2001   (F)    [Figs]

Oncaea paraobscura    Shmeleva, 1979   
Ref. compl.: Razouls & Durand, 1991 (p.46)
Loc: Médit.
N: 1
Rem.: espèce douteuse, non retenue

[48] Oncaea parila  Heron, 1977   (F,M)    [Figs]

Oncaea parobscura    Shmeleva, 1979   (F)
Ref.: Shmeleva, 1979 (p.494, Descr.F, figs.F)
Ref. compl.: Kovalev & Shmeleva, 1982 (p.86); Greze & al., 1985 (p.8); Uysal & al., 2002 (p.18, tab.1)
Loc: Médit. (Mer d'Alboran, Bassin Levantin)
N: 3
Lg.: (683) F: 0,36
Rem.: Malt, 1982 a (p.140) estime que cette espèce est une forme juvénile mâle.

[49] Oncaea petila  Heron, 1977   (F)    [Figs]

[50] Oncaea philippinensis  (Kazatchenko & Andreev, 1977)   (M)

[51] Oncaea platysetosa  Boxshall & Böttger, 1987   (F)    [Figs]

Oncaea praeclara    Humes, 1988   (F,M)
Ref.: Humes, 1988 (p.475, figs.F,M)
Ref. compl.: Suarez-Morales & Gasca, 1998 a (p.112)
Loc: Galapagos Rift, Pacif. Rise E, G. de Californie (Guaymas Basin)
N: 1
Lg.: (699) F: 1,3-1,01; M: 0,98-0,91; {F: 1,01-1,30; M: 0,91-0,98}
Rem.: Cette espèce a été prélevée à très grande profondeur( 2000 à 2635 m) au moyen des submersibles 'Alvin' et 'Cyana' sur des sites d'hydro-thermalisme du Pacifique Est. Leurs dimensions sont parmi les plus grandes chez les Oncaea . Pour Böttger-Schnack, 2001(p.30, 52) cette espèce est synonyme de Oncaea venusta.

[52] Oncaea prendeli  Shmeleva, 1966   (F,M)    [Figs]

[53] Oncaea prolata  Heron, 1977   (F,M)    [Figs]

[54] Oncaea pumilis  Heron, 1977   (F,M)    [Figs]

Oncaea quadrata    Heron & Bradford-Grieve, 1995   (F,M)
Ref.: Heron & Bradford-Grieve, 1995 (p.21, figs.F,M)
Rem.: Cf. Triconia quadrata

Oncaea redacta    Heron & Bradford-Grieve, 1995   (F,M)
Syn.: Oncaea conifera : Farran, 1936 a (p.127, figs.F: forme c); Bradford, 1972 (part., p.50); O. conifera : Moulton, 1973 (p.147 & suiv.:'minus');
Ref.: Heron & Bradford-Grieve, 1995 (p.29, figs.F, non M); Heron & Frost, 2000 (p.1032, Redescr.M, figs.M, tab.2, 3)
Rem.: Cf. Triconia redacta

[55] Oncaea rimula  Heron & Frost, 2000   (F,M)    [Figs]

[56] Oncaea rotata  Heron & Frost, 2000   (F,M)    [Figs]

[57] Oncaea rotunda  Heron, 1977   (F,M)    [Figs]

[58] Oncaea rotundata  Boxshall, 1977   (F,M)    [Figs]

Oncaea rufa    Boxshall & Böttger, 1987   (F)
Ref.: Boxshall & Böttger, 1987 (p.553, Descr.F, figs.F)
Rem.: Cf. Triconia rufa

Oncaea schmitti    Heron, 1977   (F,M)
Syn.: Epicalymma schmitti Heron, 1977 (p.82)
Ref.: Malt, 1982 (p.190, figs.F, Rem.)
Rem.: Cf. Epicalymma schmitti

[59] Oncaea scottodicarloi  Heron & Bradford-Grieve, 1995   (F,M)    [Figs]

[60] Oncaea serrulata  Böttger-Schnack, 2011   (F,M)    [Figs]

[61] Oncaea setosa  Heron, 1977   (F,M)    [Figs]

[62] Oncaea shmelevi  Gordejeva, 1972   (F,M)    [Figs]

Oncaea similis    Sars, 1918   (F,M)
Ref.: Sars, 1918 (p.193, Descr.F, figs.F); Rose, 1933 a (p.303, figs.F)
Rem.: Cf. Triconia similis

Oncaea subtilis    Giesbrecht, 1892   (F,M)
Syn.: Oncäa subtilis Giesbrecht, 1892 (p.591, 603, Descr.F, figs.F)
Rem.: Cf. Monothula subtilis

Oncaea succurva    Heron, 1977   (F)
Syn.: Conaea succurva Heron, 1977 (p.90)
Ref.: Malt, 1982 (p.192, 193, Rem.)
Rem.: Cf. Conaea succurva

[63] Oncaea tenella  Sars, 1916   (F)    [Figs]

Oncaea tenella var. pacifica    Olson, 1949   (F,M)
Ref.: Böttger-Schnack, 2002 (p.1120, Rem.: p.1125)
Rem.: Cf. Oncaea zernovi

[64] Oncaea tenuimana  Giesbrecht, 1891   (F,M)    [Figs]

Oncaea thoresoni    Heron & Frost, 2000   (F,M)
Ref.: Heron & Frost, 2000 (p.1026, figs.F,M, tab.2, 3)
Rem.: Cf. Triconia thoresoni

[65] Oncaea tregoubovi  Shmeleva, 1968   (F,M)    [Figs]

Oncaea umbonata    Heron, 1977   (F)
Syn.: Epicalymma umbonata Heron, 1977 (p.84)
Ref.: Malt, 1982 (p.191, 193)
Rem.: Cf. Epicalymma umbonata

Oncaea umerus    Böttger-Schnack & Boxshall, 1990   (F)
Syn.: Oncaea sp. Böttger-Schnack, 1988, 1990
Ref.: Böttger-Schnack & Boxshall, 1990 (p.861, Descr.F, figs.F, tab.III);
Rem.: Cf. Triconia umerus

[66] Oncaea venella  Farran, 1929   (F,M)    [Figs]
Rem.: Farran (1929) characterises this variety by its dimensions. Tanaka (1960, p.72) observed in his material a small form (0/90-1.00 mm) corresponding to the venella form. The proportional lengths of the anterior and posterior regions of the body female are as 60 to 40. The anterior region about 1.6 times as long as broad (61 : 39), this is more slender than in forma typica (= O. venusta. The posterior region has the segments in the proportional lengths 10 : 45 : 7 : 7 : 9 : 22 = 100. The caudal rami 4 times as long as broad. No structural differences were observed between the two forms venusta and venella.
Heron (2002) considers this variety as a true species (after essentially its dimensions and some minor morphological details), and distinguishes it as a new species O. frosti (see remark for this species).
After Heron (2002, p.150) this is the first time that Oncaea venusta var. venella is elevated to the species level. The difference in size specifications has caused some confusion for investigators. The smaller size reported in Farran (1936) is considered the most authentic, since it was based on additional material. Because specimens have been found conforming to Farran’s size range from three different areas, the smaller size for O. venella appears to be confirmed.
Its presence in the eastern Pacific is confirmed; however, Böttger-Schnack & Huys (2004) contest the geographic localisations of this form which is not always validly identified, and even the existence of this species that corresponds only to a simple form.
See remarks in Oncaea venusta.
After Böttger-Schnack & Schnack ( 2013, p. 7, Table 2) the species from Heron (2002) is insufficiently described .
Observed in the ballast waters of ships in San Francisco.

[67] Oncaea venusta  Philippi, 1843   (F,M)    [Figs]
Rem.: Epi- to mesopelagic, also demersal and semi-parasitic.
Sampling depth (Antarct., sub-Antarct.) : 400 m. Overall depth Range in Sargasso Sea: 0-500 m (Deevey & Brooks, 1977, Station "S"). 0-1236 m at Station T-1 (E Tori Is, E middle Japan) from Furuhashi (1966 a).
This species presents 2 forms (sub-species?) that are characterised by their dimensions. Sewell (1947) suggests also that their spawning season is different. For this author the forma typica is characterized by: The proportional lengths of the anterior and posterior regions of the body as 59 : 41, and the length and breadth of the anterior region are as 59 to 29.. The proportional lengths of the segments of the posterior region (5th thoracic segment to caudal rami) as 13 : 46: 7: 7: 9 : 18. In the forma venellaThe proportional lengths of the anterior and posterior regions of the body as 60 to 40, and the length and breadth of the anterior region as 60 to 24. The proportional lengths of the segments of the posterior region (5th thoracic segment to caudal rami) as 13 : 43 : 7 : 8 : 9: 20 = 100. The proportions of the caudal rami are as 23 to 6.
For Tanaka (1960, p.71) the two forrms : large and small, are present in his material. Female: The proportional lengths of the anterior and posterior regions of the body are as 57 to 43. The anterior region is 1.7 times as long as wide (63 : 37). The proportional lengths of the segments of the posterior region (caudal rami included) as 13 : 48 : 5 : 5 : 9 : 20 = 100. Caudal rami 4 times as long as broad.
Heron (2002) considers the two varieties O. venusta typica and O. venusta venella as two species (see remarks in Oncaea venella and frosti), position objected to by Böttger-Schnack & Huys (2004).
After KazKazmi (2004, p.232), although free living and pelagic, the species has been found on fish gills (Kazatchenko & Adeev, 1977) or in a sponge (Ho, 1984) and now from the sand (Kazmi & Naushaba, 2000).
After Benedetti & al. (2018, p.1, Fig.2) this species belonging to the functional group 5 corresponding to small sac-spawning detritivorous.

See in DVP Conway & al., 2003 (version 1)

Oncaea venusta typica    Farran, 1929   (F,M)
Syn.: Oncaea venusta : Farran, 1929 (p.284)
Ref.: Farran, 1929 (p.284); Sewell, 1947 (p.263, Rem.F); 1951 (p.371, Rem.: parasites); Corral Estrada, 1970 (p.217); Boxshall, 1977 a (p.127, Rem.); 1977 b (p.548); Malt, 1983 (p.454, figs.F); 1983 a (figs.F,M, Rem., p.5, 7); Ho, 1984 (p.42, Rem.); Böttger-Schnack & al., 1989 (p.1089); Böttger-Schnack, 1994 (p.277); 1995 (p.92); Ohtsuka & al., 1996 a (p.91); Chihara & Murano, 1997 (p.981, Pl.223: F,M); Nakata & al., 2001 (p.335, tab.4, 5, 7); Böttger-Schnack, 2001 (p.31, 46, figs.F,M, Rem.); Rezai & al., 2004 (p.490, tab.2); Böttger-Schnack & Huys, 2004 (p.1, Rem.); Böttger-Schnack & al., 2004 (p.1130, tab.1, Rem.)
Ref. compl.: Böttger-Schnack & al., 2001 (p.1029, tab.1); Razai & al., 2004 (p.490, tab.2)
Lg.: (11) F: 1,25-1,185; (139) F: ± 1,13; (788) F: 1,23-1; M: 0,95-0,88; (819) F: 1-1,2; M: 0,88-0,95; (833) F: 1,23-1,09; M: 0,98-0,89; {F: 1,00-1,25; M: 0,88-0,95}
Rem.: Cette variété est caractérisée d’après Farran (1929) par ses dimensions: F = 1,08 -1,16 mm; M = 0,78-0,81 mm

Oncaea venusta “robust form”       
Ref.: Boxshall, 1977 a (p.127, fig.F, Rem.); 1977 b (p.546)
Lg.: (139) F: 1,04-0,88; {F: 0,88-1,04}

Oncaea venusta venella    Farran, 1929   
Ref.: Farran, 1929 (p.210, 284, fig.F); Boxshall, 1977 a (p.127: Rem.); Böttger-Schnack & al., 2004 (p.1130, tab.1, Rem.)
Loc: Atlant. (tropical), Pacif. SW, Grande Barrière)
Lg.: (35) F: 1,07-0,92
Rem.: Cf. Oncaea venella. Voir remarques in Böttger-Schnack & Huys, 2004 (p. 2 & suiv.)

[68] Oncaea vodjanitskii  Shmeleva & Delalo, 1965   (F,M)    [Figs]

[69] Oncaea waldemari  Bersano & Boxshall, 1994    (F,M)    [Figs]

[70] Oncaea walleni  Heron, 1977   (F)    [Figs]

[71] Oncaea zernovi  Shmeleva, 1966   (F,M)    [Figs]

[72] Oncaea sp.1  [Heron & Frost, 2000]   (F,M)    [Figs]

Oncaea sp.1    Vilela, 1968   
Ref.: Vilela, 1968 (p.33, figs. Juv.F)
Loc: Portugal
Lg.: (327) juv.F: 0,99-0,81

Oncaea sp.2    Vilela, 1968   (M)
Ref.: Vilela, 1968 (p.33, figs.M)
Loc: Portugal
Lg.: (327) M: 0,87

Oncaea sp. A    Razouls, 1974   (F)
Ref.: Razouls, 1972 (p.95, Annexe: p.118, figs.F); 1974 b (p.241, figs.F)
Loc: Médit. (G. du Lion: Banyuls)
Rem.: Cf. Oncaea minuta

Oncaea sp.1    Ferrari, 1975   (F)
Ref.: Ferrari, 1975 (p.228, figs.F); Boxshall, 1977 a (p.143); Böttger-Schnack, 2001 (p.29, Rem.)
Loc: G. du Mexique
Lg.: (687) F: 0,71-0,68
Rem.: Cf. ? Oncaea media ou curta

Oncaea sp.2    Ferrari, 1975   (F)
Ref.: Ferrari, 1975 (p.228, figs.F); Boxshall, 1977 a (p.143); Böttger-Schnack, 2001 (p.29, Rem.)
Loc: G. du Mexique
Lg.: (687) F: 0,58-0,54
Rem.: Cf. ? Oncaea curta ou venusta venella

Oncaea sp. K    Böttger-Schnack, Hagen & Schnack-Schiel, 2001   
Ref.: Böttger-Schnack & al., 2001 (p.1029, tab.1, 2)
Loc: G. d'Aqaba, Mer Rouge (N-S)

Oncaea 'zernovi'    Böttger-Schnack, Hagen & Schnack-Schiel, 2001   
Ref.: Böttger-Schnack & al., 2001 (p.1029, tab.1, 2)
Loc: G. d'Aqaba, Mer Rouge (N-S)
(6) Spinoncaea Böttger-Schnack, 2003
Rem.: type: Oncaea ivlevi. Total: 3 spp.
For Böttger-Schnack & Schnack (2013, p.19) the genus displays intrageneric variation in the spine count on the exopod segment of P2 (Table 5, type 6). Similar variation has also been found in the longipes-group and the zernovi -group (Table 5: type 5 and type 7a, respectively). In allthese cases, however, all the other morphological characters of the species, including details of ornamentation, were similar (unlike the situation within Conaea s.l.), indicating that they are closelt related sibling/sister species within the same group. Hence, differences in exopodal spine counts on the distal segment of P2 alone are not regaeded as an indication for defining separate subgroups within any group defined in the present study.

[1] Spinoncaea humesi  Böttger-Schnack, 2003   (F,M)    [Figs]

[2] Spinoncaea ivlevi  (Shmeleva, 1966)   (F,M)    [Figs]

Spinoncaea ivlevi "robust form"       
Ref.: Böttger-Schnack, 2003 (p.204, figs.F)

Spinoncaea ivlevi "elongate form"       
Ref.: Böttger-Schnack, 2003 (p.204, figs.F)

[3] Spinoncaea tenuis  Böttger-Schnack, 2003   (F,M)    [Figs]

Spinoncaea tenuis "typical form"       
Ref.: Böttger-Schnack, 2003 (p.222, Rem.)

Spinoncaea tenuis "elongate form"       
Ref.: Böttger-Schnack, 2003 (p.222, Rem.)
(7) Triconia Böttger-Schnack, 1999
Rem.: type: Oncaea conifera Giesbrecht,1891. Total: 26 spp. + 1 indet.

Generic name derived from the Latin tres, tria, meaning three and conus, meaning cone, and refers to the presence of a conical projection on the distal endopodal segments of the last three pairs of swimming legs.
Böttger-Schnack (1999) established the genus Triconia which is composed of all oncaeids having a large conical process on the distal margin of the endopod of swimming legs 2-4.
This genus was subdivided into 3 subgroups: similis, conifera, dendipes.
After reexamination, the similis sub-group includes 2 groups, which can be distinguished by the proportional spine lengths on the endopods of swimming P2: 1- containingT. minuta, T. umerus, T. gonopleura, T. parasimilis, present a relatively short distal endopodal spine: 2- including T. similis, T. hawii, T. recta, T. denticulata, with a relatively long distal endopodal spine.

[1] Triconia antarctica  (Heron, 1977)   (F,M)    [Figs]

[2] Triconia borealis  (Sars, 1918)   (F,M)    [Figs]

[3] Triconia canadensis  (Heron & Frost, 2000)   (F,M)    [Figs]

[4] Triconia conifera  (Giesbrecht, 1891)   (F,M)    [Figs]

[5] Triconia constricta  Wi, Böttger-Schnack & Soh, 2012   (F,M)    [Figs]

[6] Triconia denticula  Wi, Shih & Soh, 2011   (F)    [Figs]

[7] Triconia dentipes  (Giesbrecht, 1891)   (F,M)    [Figs]

[8] Triconia derivata  (Heron & Bradford-Grieve, 1995)   (F,M)    [Figs]

[9] Triconia elongata  Böttger-Schnack, 1999   (F,M)    [Figs]

[10] Triconia furcula  (Farran, 1936)   (F,M)    [Figs]

[11] Triconia giesbrechti  Böttger-Schnack, 1999   (F, M)    [Figs]

[12] Triconia gonopleura  Böttger-Schnack, 1999   (F)    [Figs]

[13] Triconia hawii  (Böttger-Schnack & Boxshall, 1990)   (F,M)    [Figs]

[14] Triconia hirsuta  Wi, Böttger-Schnack & Soh, 2010   (F)    [Figs]

[15] Triconia inflexa  (Heron, 1977)   (F,M)    [Figs]

[16] Triconia minuta  Giesbrecht, 1892   (F, M)    [Figs]

[17] Triconia pacifica  Cho, Kim, Böttger-Schnack, Lee, 2013   (F,M)    [Figs]

[18] Triconia pararedacta  Wi, Böttger-Schnack & Soh, 2012   (F)    [Figs]

[19] Triconia parasimilis  Böttger-Schnack, 1999   (F)    [Figs]

[20] Triconia quadrata  (Heron & Bradford-Grieve, 1995)   (F,M)    [Figs]

[21] Triconia recta  Böttger-Schnack, 1999   (F,M)    [Figs]

[22] Triconia redacta  (Heron & Bradford-Grieve, 1995)   (F,M)    [Figs]

[23] Triconia rufa  (Boxshall & Böttger, 1987)   (F,M)    [Figs]

[24] Triconia similis  (Sars, 1918)   (F,M)    [Figs]

[25] Triconia thoresoni  (Heron & Frost, 2000)   (F,M)    [Figs]

[26] Triconia umerus  (Böttger-Schnack & Boxshall, 1990)   (F,M)    [Figs]

[27] Triconia sp.  Böttger-Schnack, 1999       [Figs]

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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2024. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed April 27, 2024]

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