Species Card of Copepod
Cyclopoida ( Order )
    Corycaeidae ( Family )
        Farranula ( Genus )
Farranula gracilis  (Dana, 1849)   (F,M)
Syn.: Corycaeus gracilis Dana, 1849; 1852; Dahl, 1894 c (p.22, Rem.); F. Dahl, 1894 (p.69); Khalil & Abb El-Rahman, 1997 (p.931);
C. pellucidus Dana, 1849; 1852; Brady, 1883 (p.112, figs.F,M); T. Scott, 1894 b (p.110);
C. deplumatus Dana, 1849; 1852;
C. megalops : Brady, 1883 (p.112);
Corycaeus (Corycella) gracilis : M. Dahl, 1912 (p.108, figs.F,M); Sewell, 1948 (p.393); Marques, 1958 a (p.148); Cervigon, 1964 (p.197, figs.F,M); Binet, 1970 (p.74);
Corycella gracilis : Farran, 1929 (p.211, 296); Rose, 1929 (p.72); Klevenhusen, 1933 a (p.84, chart 34); Owre, 1962 (p.489); Björnberg, 1963 (p.85, Rem.); Gaudy, 1963 (p.33, Rem.); 1967 (p.88); Ramirez, 1971 (p.92); Binet & al., 1972 (p.73); Bainbridge, 1972 (p.90, 92, 94); Kovalev & Shmeleva, 1982 (p.86); Bityukov & al., 1984 (tab.1); Selifonova, 2011 (p.77, Table 1, alien species in Black Sea);
C. (Farranula) gracilis : Marques, 1973 (p.250); Böttger-Schnack, 1988 (p.313);
? Corycaeus carinatus : Wheeler, 1901 (p.192, figs.F);
? Corycella carinata : Wilson, 1932 (p.41); 1932 a (p.362, figs.F)
Ref.:
Wilson, 1942 a (p.187, fig.F); Gonzalez & Bowman, 1965 (p.267, figs.F,M, Rem.); Owre & Foyo, 1967 (p.124, figs.F,M); Björnberg, 1972 (p.94, figs., Rem.: Nauplius); Marques, 1975 (p.53); Björnberg, 1981 (p.676, fig.F); Boxshall, 1998 (p.223); Bradford-Grieve & al., 1999 (p.888, 975, figs.F,M); Conway & al., 2003 (p.262, figs.F,M, Rem.); Boxshall & Halsey, 2004 (p.493); Vives & Shmeleva, 2010 (p.225, figs.F,M, Rem.)
Species Farranula gracilis - Plate 1 of morphological figuresissued from : M. Dahl in Ergebnisse der Plankton-Expedition der Humboldt-Stiftung. Bd II, G. f1. I. Die Corycaeinen 1912. [Taf.XIV, Figs.11, 12]. As Corycaeus (Corycella) gracilis..
Female: 11, habitus (dorsal); 12, idem (lateral right side).


Species Farranula gracilis - Plate 2 of morphological figuresissued from : M. Dahl in Ergebnisse der Plankton-Expedition der Humboldt-Stiftung. Bd II, G. f1. I. Die Corycaeinen 1912. [Taf.XIV, Figs.15]. As Corycaeus (Corycella) gracilis..
Female: 15, A2.


Species Farranula gracilis - Plate 3 of morphological figuresissued from : M. Dahl in Ergebnisse der Plankton-Expedition der Humboldt-Stiftung. Bd II, G. f1. I. Die Corycaeinen 1912. [Taf.XIV, Figs.18, 19]. As Corycaeus (Corycella) gracilis..
Female: 18, P1; 19, P2.


Species Farranula gracilis - Plate 4 of morphological figuresissued from : M. Dahl in Ergebnisse der Plankton-Expedition der Humboldt-Stiftung. Bd II, G. f1. I. Die Corycaeinen 1912. [Taf.XIV, Figs.20, 21]. As Corycaeus (Corycella) gracilis..
Female: 20, P3: 21, P4.


Species Farranula gracilis - Plate 5 of morphological figuresissued from : M. Dahl in Ergebnisse der Plankton-Expedition der Humboldt-Stiftung. Bd II, G. f1. I. Die Corycaeinen 1912. [Taf.XV, Figs.15, 16]. As Corycaeus (Corycella) gracilis..
Female: 15, posterior part of metasome and urosome (dorsal); 16, idem (lateral left side; with spermatophores).


Species Farranula gracilis - Plate 6 of morphological figuresissued from : M. Dahl in Ergebnisse der Plankton-Expedition der Humboldt-Stiftung. Bd II, G. f1. I. Die Corycaeinen 1912. [Taf.XIV, Figs.13, 14]. As Corycaeus (Corycella) gracilis..
Male: 13, habitus (dorsal); 14, idem (lateral left side).


Species Farranula gracilis - Plate 7 of morphological figuresissued from : M. Dahl in Ergebnisse der Plankton-Expedition der Humboldt-Stiftung. Bd II, G. f1. I. Die Corycaeinen 1912. [Taf.XIV, Figs.16, 17]. As Corycaeus (Corycella) gracilis..
Male: 16, A2; 17, Mxp.


Species Farranula gracilis - Plate 8 of morphological figuresissued from : M. Dahl in Ergebnisse der Plankton-Expedition der Humboldt-Stiftung. Bd II, G. f1. I. Die Corycaeinen 1912. [Taf.XV, Figs.29, 30]. As Corycaeus (Corycella) gracilis..
Male: 29, posterior part of metasome and urosome (lateral right side); 30, idem (dorsal).


Species Farranula gracilis - Plate 9 of morphological figuresissued from : F. Cervigon iin Mem. Soc. Cienc. nat. La Salle, 1964, 24 (68). [p.198, Lam.24]. As Corycella gracilis.
Female (Venezuela): 1, prosome (dorsal); 2, habitus (lateral); 3, portion anteroventral of genital segment; 4, A2; 5, P4.

Nota: Lengths ratio abdomen and caudal rami 68:32 and 71:29.


Species Farranula gracilis - Plate 10 of morphological figuresissued from : F. Cervigon iin Mem. Soc. Cienc. nat. La Salle, 1964, 24 (68). [p.19, Lam.25]. As Corycella gracilis.
Male: 1-2, habitus (dorsal and lateral, respectively); 3, A2; 4, P4.

Nota: Lengths abdomen and caudal rami 67.5:32.5; 67:33 and 69:31.


Species Farranula gracilis - Plate 11 of morphological figuresFemale: Urosome about 2 times as long as the caudal rami and, in lateral view, appears more or less humpbacked. In lateral view of the urosome, part of the dorsal margin, before it tapers , lies almost parallel to the ventral margin; the length of this portion of the urosome is about equal to the height of the urosome.

Male: Urosome less than 3 times as long as the caudal rami; rami much more than twice as long as broad. Posterior margin of genital enlargements separated from the posterior end of the urosome by more than 1/3 the length of the urosome.

Compl. Ref.:
C.B. Wilson, 1950 (p.228); Fagetti, 1962 (p.50); Grice, 1963 a (p.496); Neto & Paiva, 1966 (p.34, Table III); Deevey, 1971 (p.224); Björnberg, 1973 (p.369, 386); Boxshall, 1977 b (p.556); Deevey & Brooks, 1977 (p.156, tab.2, Station "S"); Binet, 1979 (p.400); Dessier, 1979 (p.131, 202, 208); Vives, 1982 (p.296); Turner & Dagg, 1983 (p.17, 22); Greze & al., 1985 (p.8); Lindo, 1991 (tab.3); Böttger-Schnack, 1995 (p.92); Webber & Roff, 1995 (tab.1); Webber & al., 1996 (tab.1); Suarez-Morales & Gasca, 1997 (p.1525); Alvarez-Cadena & al., 1998 (t.1,2,3,4); Hopcroft & al., 1998 (tab.2); Lopes & al., 1998 (p.195, tab.2); Suarez-Morales & Gasca, 1998 a (p.112); Lopes & al., 1999 (p.215, tab.1); Lavaniegos & Gonzalez-Navarro, 1999 (p.239, Appx.1); Neumann-Leitao & al., 1999 (p.153, tab.2); Razouls & al., 2000 (p.343, tab. 5, Appendix); Suarez-Morales & Gasca, 2000 (1247, tab.1); Lopez-Salgado & al., 2000 (tab.1); Dunbar & Webber, 2003 (tab.1); Camisotti & al., 2005 (p.99); Alvarez-Silva & al., 2005 (p.39); Dias & Araujo, 2006 (p.85, Rem., chart); Neumann-Leitao & al., 2008 (p.799: Tab.II, fig.6); Morales-Ramirez & Suarez-Morales, 2008 (p.516, 525); Miyashita & al., 2009 (p.815, Tabl.II); Lidvanov & al., 2010 (p.356, Table 3); Dias & al., 2010 (p.230, Table 1); Medellin-Mora & Navas S., 2010 (p.265, Tab. 2); Shiganova & al., 2012 (p.61, Table 4); Uysal & Shmeleva, 2012 (p.909, Table I)
NZ: 19

Distribution map of Farranula gracilis by geographical zones
Species Farranula gracilis - Distribution map 4issued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. 1967. [p.124, Table 66].
Vertical distribution of Farranula gracilis at the ''40-Mile station'' in the Florida Current (E Miami: ± 25°35'N, 79°27'W).
SL 53: 18 V 1958; A: during midday; B: during midnight.
Loc:
Antarct. (Mer de Ross), sub-Antarct. (Pacif. SW), Angola, Baia Farta, Congo, Is. San Tomé et Principe, G. de Guinée, Ivorian shelf, Patagonie, Brazil (S, Vitoria-Cabo de Sao Tomé, off Macaé, off Natal, Amazone), Is. du Cap Vert, off Maroc-Mauritanie, off Madère, I. Barbade, Mer des Antilles, Caribbean Colombia, Yucatan, Venezuela, Jamaïque (Kingston Harbour), Porto Rico, E Costa Rica, G. du Mexique, Cuba, Floride, Long Island, Mer des Sargasses, off Bermuda (Station "S"), off Açores W, Médit. (Mer d'Alboran, Ionienne-Chypre, N Levantin Basin, Black Sea), Mer Rouge (G. d'Aqaba), Natal, Maldives, Madagascar, Indien S, Philippines, mers de Chine, off Nouvelle-Zélande S, Is. Fidji, off Hawaii, Galapagos, off Washington, Californie, G. de Californie, Zihuatanejo Bay, W Costa Rica, Equateur, Chili N, Mer de Béring
N: 76
Lg.:
(35) (Atlant. trop.) F: 0,96-0,9; M: 0,96-0,8; (Atlant. N) F: 0,79; M: 0,75-0,66; (Antarct.) F: 0,99; (179) F: 0,94; (237) F: 0,93-1,1; M: 0,8-1,0; (336) F: 1,1-0,9; M: 0,9-0,7; (618) F: 1,03-0,93; M: 0,86-0,83; (651) F: 0,99-0,92; M: 0,89-0,77; (666) F: 1; M: 0,8; (668) F: 0,98-0,96; M: 0,92-0,86; (991) F: 0,96-0,98; M: 0,86-0,92; {F: 0,79-1,10; M: 0,66-1,00}
Rem.: épi-méso-bathypélagic.
Sampling depth (Antarct.) : 0-450 m. Overall Depth Range in Sargasso Sea: 0-500 m (Deevey & Brooks, 1977, Station "S");
See remark on Farranula carinata.
After Björnberg (1963, p.86) this species is the most frequent Corycaeid, besides C. giesbrechti in the Brazilian waters. When in large numbers, it is a good indicator of very saline (above 35.5 p.1000) and warm waters (above 21°C).

See in DVP Conway & al., 2003 (version 1)
Last update : 02/05/2013
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[Accessed May 23, 2013]

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