Species Card of Copepod
Cyclopoida ( Order )
    Oncaeidae ( Family )
        Triconia ( Genus )
Triconia borealis  (Sars, 1918)   (F,M)
Syn.: Oncaea conifera : Sars, 1900 (p.113, figs.F);
Oncaea borealis Sars, 1918 (p.191, figs.F,M); Früchtl, 1924 b (p.90, Rem.); Campbell, 1929 (p.329, Rem.F,M); Wilson, 1932 a (p.351, figs.F,M); Rose, 1933 a (p.303, figs.F,M); Jespersen, 1934 (p.127); 1940 (p.92); Sewell, 1948 (p.392, 496); Hansen K.V., 1951 (p.231, migration v.s. discontinuity layer); Østvedt, 1955 (p.15: Table 3, p.17, p.79); Wiborg, 1955 (p.47); Gardner & Szabo, 1982 (p.114, figs.F,M); Gagnon & Lacroix, 1982 (p.9, 12); Malt, 1983 b (p.458); 1983 a (p.6, figs.F,M, Rem.); Heron & al., 1984 (p.461, 466, figs.F,M, Rem.); Mikhailovsky, 1986 (p.89) ; Van Hove & al., 2001 (p.303); Heron & Frost, 2000 (p.1034, figs.F, tab.2, 3);
Ref. compl. : Jespersen, 1939 (p.81, Rem., Table 25); Grice, 1962 a (p.101); Mazza, 1966 (p.73); Pertsova, 1967 (p.240); Vinogradov, 1968 (1970) (p.256); Strömgren, 1969 (p.5); Shih & al., 1971 (p.56, 156, 215); Kolosova, 1975 (p.92, fig.3, Table 1); Peterson & Miller, 1975 (p.650); Gibson & Grice, 1977 (p.85, Table 1, copper pollution); Shih & Laubitz, 1978 (p.50); Buchanan & Sekerak, 1982 (p.41, Table 3: vertical distribution); Kovalev & Shmeleva, 1982 (p.86); Groendahl & Hernroth, 1986 (tab.1); Sameoto & al., 1986 (p.53); Kosobokova, 1989 (p.27); Anderson J.T., 1990 (p.127, Rem.: p.131); Vinogradov & al., 1994 (tab.1); Richter, 1994 (tab.4.1a); Abramova, 1996 (tab.1); Kosobokova & al., 1998 (tab.2); Dolganova & al., 1999 (p.13, tab.1); Bragina, 1999 (p.196); Abramova, 1999 (p.161, Table 2); Halvorsen & Tande, 1999 (p.279, figs.3); Kosobokova & Hirche, 2000 (p.2029, tab.2); Lischka & al., 2001 (tab.3, 6); Musaeva & Suntsov, 2001 (p.511); Lischka & al., 2001 (p.186); Fortier M. & al., 2001 (p.1263, fig.6, 7, diel vertical migration); Auel & Hagen, 2002 (p.1013, tab.2);Ringuette & al., 2002 (p.5081, Table 1, Fig.7, population dynamic); Sameoto & al., 2002 (p.13); Kosobokova & al., 2003 (p.697, tab.2); Gislason & Astthorsson, 2004 (p.472, tab.1); Dmoch & Walczowski, 2005 (p.102); Hopcroft & al., 2005 (p.198, table 2); Deibel & Daly; 2007 (p.271, Table 1, 2, Rem.: Arctic polynyas); Humphrey, 2008 (p.84: Appendix A); Darnis & al., 2008 (p.994, Table 1, figs.8, 9); Galbraith, 2009 (pers. comm.); Pomerleau & al., 2011 (p.1779, Table III)
Ref.:
Böttger-Schnack, 1999 (p.43, Rem.); Nishibe & Ikeda, 2004 (p.931, Tab. 2, 4, 5); Boxshall & Halsey, 2004 (p.614: fig.F); Böttger-Schnack & Schnack, 2009 (p.131, Table 5: Rem.); Wi & al., 2010 (p.684, Re-descr.F,M, figs.F,M, Rem.); Vives & Shmeleva, 2010 (p.339, figs.F,M, Rem.); Wi & al., 2011 (p.601: Rem.: variations)
Species Triconia borealis - Plate 1 of morphological figuresissued from : Sars G.O. in An Account of the Crustacea of Norway. Vol. VI. Copepoda Cyclopoida. Published by the Bergen Museum, 1918. [Pl. CVIII]. As Oncaea borealis.
Female & Male (from Oster Fjord, near Bergen).
m = Md + Mx1; mp1 = Mx2; mp2 = Mxp.


Species Triconia borealis - Plate 2 of morphological figuresissued from : G.A. Heron & B.W. Frost in Proc. Biol. Soc. Washington, 2000, 113 (4). [p.1021, Fig.1, E]. As Oncaea borealis.
Female: P2-P4 endopod terminal ''spine set'' (combination of the lengths, shapes, and position of the three or two terminal spines on the endopods of swimming legs 2-4 , from left to right in figure, for different Oncaea' species).

Nota: endopod of P4 with subterminal and lateral spine lengths both about half that of terminal spine.


Species Triconia borealis - Plate 3 of morphological figuresissued from : G.A. Heron, T.S. English & D.M. Damkaer in J. Crustacean Biol., 1984, 4 (3). [p.464, Fig.9, D-G]. As Oncaea borealis.
Female (from Arctic): D, habitus (lateral; scale bar: y); E-F, posterior of last prosomal segment and urosome (lateral and dorsal, respectively; scale bar: w); G, right A2 (scale bar: w).


Species Triconia borealis - Plate 4 of morphological figuresissued from : G.A. Heron, T.S. English & D.M. Damkaer in J. Crustacean Biol., 1984, 4 (3). [p.467]. As Oncaea borealis.
Female & Male: Armature of swimming legs P1-P4. Roman numerals: spines; arabic numerals: setae. Si = ineer setae; st = terminal setae; se = outer setae.


Species Triconia borealis - Plate 5 of morphological figuresissued from : G.A. Heron, T.S. English & D.M. Damkaer in J. Crustacean Biol., 1984, 4 (3). [p.468, Fig.10]. As Oncaea borealis.
Female: A, labrum (ventral; scale bar: x); B, right Md (scale bar: x); C, left Mx1 (scale bar: x); D, right Mx2 (scale bar: x); E, right Mxp (scale bar: z); F-I, P1 to P4 (scale bar: w); J, P5 (scale bar: x).


Species Triconia borealis - Plate 6 of morphological figuresissued from : G.A. Heron, T.S. English & D.M. Damkaer in J. Crustacean Biol., 1984, 4 (3). [p.469, Fig.11, A-D]. as Oncaea borealis.
Male: A, habitus (lateral; scale bar: w); B, posterior of last peosomal segment and urosome ( dorsal; scale bar: w); C, right Mxp (scale bar: x); D, P5 (scale bar: x).


Species Triconia borealis - Plate 7 of morphological figuresissued from : J.H. Wi, R. Böttger-Schnack & H.Y. Soh in J. Crustacean Biol., 2010, 30 (4). [p.685, Fig.8].
Female (from 36°68.33'N, 129°52.42'E): A-B, habitus (dorsal and lateral, respectively); C, A1; D, A2 (ornamentation on coxobasis obscured, not discerned); E, Md; F, Mx1; G, Mxp; H, P5; I, anal somite and caudal ramus (dorsal view), setae IV and V.
Scale bars in micrometers.


Species Triconia borealis - Plate 8 of morphological figuresissued from : J.H. Wi, R. Böttger-Schnack & H.Y. Soh in J. Crustacean Biol., 2010, 30 (4). [p.687, Fig.10 A].
Female (SEM photograph): A, A2, distal endopodal segment (arrow indicatig ornamentation).
Scale bar in micrometers.


Species Triconia borealis - Plate 9 of morphological figuresissued from : J.H. Wi, R. Böttger-Schnack & H.Y. Soh in J. Crustacean Biol., 2010, 30 (4). [p.687, Fig.10 C].
Female (SEM photograph): C, P1 (part of basis and proximal segments of endopod (anterior view), showing inner basal seta.
Scale bar in micrometers.


Species Triconia borealis - Plate 10 of morphological figuresissued from : J.H. Wi, R. Böttger-Schnack & H.Y. Soh in J. Crustacean Biol., 2010, 30 (4). [p.687, Fig.10 D].
Female: D, endopodal segments 3 of P3 and P4 (arrows indicating apical pores on conical processes)
Scale bar in micrometres.


Species Triconia borealis - Plate 11 of morphological figuresissued from : J.H. Wi, R. Böttger-Schnack & H.Y. Soh in J. Crustacean Biol., 2010, 30 (4). [p.687, Fig.10 B].
Female (SEM photograph): B, genital apertures (arrows indicating spinous processes close to seta of P6).
Scale bar in micrometers.


Species Triconia borealis - Plate 12 of morphological figuresissued from : J.H. Wi, R. Böttger-Schnack & H.Y. Soh in J. Crustacean Biol., 2010, 30 (4). [p.687, Fig.10 E].
Female (SEM photograph): E, caudal rami (ventral view).
Scale bar in micrometers.

Compl. Ref.:
Blachowiak-Samolyk & al., 2006 (p.101, tab.1); Hop & al., 2006 (p.182, Table 4, 5: inter-annual variability, fig.15); Blachowiak-Samolyk & al., 2007 (p.2716, Table 2); Nishibe & Ikeda, 2008 (p.398, tab.1); Hopcroft & al., 2009 (p.9, Table 3); Kosobokova & Hopcroft, 2010 (p.96, Table 1, fig.7); Hopcroft & al., 2010 (p.27, Table 2, fig.9); Kosobokova & al., 2011 (p.29, Table 2, figs.4, 7, Rem.: Arctic Basins); DiBacco & al., 2012 (p.483, Table S1, ballast water transport)
NZ: 8

Distribution map of Triconia borealis by geographical zones
Species Triconia borealis - Distribution map 3issued from : P. Jespersen, 1939 in Meddr Grønland, 1939, 119 (9). [p.82, Fig. 34].
The occurrence of Oncaea borealis (= Triconia borealis at the ''Dana'' stations in 1932 and 1933.
black circles: positive occurrence (common occurrence); white circle (in few numbers); x: negative occurrence.
Loc:
Médit. W (Chenal Liguro-provençal: in Mazza, 1966), Woods Hole, off Nova Scotia E, Estuaire du St. Laurent, off Newfoundland, Cap Flemish, , Groenland (W, Kejser Franz Joseph and Scoresby Sund Fjord), Baffin (Lancaster Sound, W baffin Bay); Mer du Groenland, Fram Strait, Kongsfjorden ( W Spitsberg), Islande, Mer de Norvège, Oslo fjord, Lysefjorden, Mer du Nord*, Arct. (Mer Blanche, Mer de Barents, Pechora Sea, Mer de Laptev, Chupa Inlet, Lomonosov Ridge, Mer de Kara, Fram Strait, Ellesmere Is., Canadian abyssal plain, Nansen Basin, Amunsen Basin, Barrow Strait, Makarov Basin, Sibérie E, Mer de Chukchi, off NW Alaska, SE Beaufort Sea, Canada Basin, Mer de Beaufort), Mer du Japon, Oyashio region, Kouriles S, Station "P", Strait of Géorgia, Washington inland, Oregon (Yaquina)
N: 71
Lg.:
(45) F: 0,7-0,6; M: 0,45-0,35; (133) F: 0,7; M: 0,4; (134) F: 0,75; (141) F: 0,7; M: 0,4; (675) F: 0,76-0,64; M: 0,56-0,44; (796) F: 0,8-0,69; M: 0,54-0,46; (1064) F: 0,564-0,687; M: 0,38-0,398; {F: 0,56-0,76; M: 0,35-0,56}
Rem.: epi-mesopelagic in the polar basin.
Collected by Heron & al. (1984, p.467) in samples mainly from above 500 m.
Heron & Frost (2000, p.1017) do not agree that the generic definition of Triconia by Böttger-Schnack (1999) is sufficient to justify inclusion of all of the divergent species she assigned to the new genus.
For Wi & al. (2010, p.688) the species from Korean waters shows the typical morphological characteristics of T. borealis (cf. Sars, 1918); Heron & al. (1984) including some important details lacking in Sars' description.
Last update : 10/05/2013
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