Species Card of Copepod
Calanoida ( Order )
    Arietelloidea ( Superfamily )
        Augaptilidae ( Family )
            Haloptilus ( Genus )
Haloptilus ornatus  (Giesbrecht, 1892)   (F,M)
Syn.: Hemicalanus ornatus Giesbrecht, 1892 (p.384, 399, 773, figs.F,M); Fish, 1962 (p.21)
Ref.:
Giesbrecht & Schmeil, 1898 (p.120, Rem.F,M, fig.F); A. Scott, 1909 (p.141, Rem.); Wolfenden, 1911 (p.323); Sars, 1925 (p.247, figs.F); Sewell, 1932 (p.328); Rose, 1933 a (p.213, figs.F); Mori, 1937 (1964) (p.75, figs.F); Lysholm & al., 1945 (p.37); Sewell, 1947 (p.190, 194, fig.F); Chiba, 1956 (p.52, figs.M); Chiba & al., 1957 a (p.11); Grice, 1962 (p.223, figs.F); Tanaka, 1964 b (p.43, Rem.F); Vervoort, 1965 (p.126, Rem.); Owre & Foyo, 1967 (p.81, figs.F,M); Tanaka & Omori, 1971 (p.253, Rem.F); Matthews, 1972 (p.52); Björnberg & al., 1981 (p.653, 654, figs.F,M); Park, 1988 (p.14, Descr.F, figs.F, Rem.); Chihara & Murano, 1997 (p.728, Pl.58, 61: F,M); Bradford-Grieve & al., 1999 (p.883, 941, figs.F,M); Bradford-Grieve,1999 b (p.63, figs.F,M, Rem., figs.172, 190); Vives & Shmeleva, 2007 (p.244, figs.F,M, Rem.)
Species Haloptilus ornatus - Plate 1 of morphological figuresissued from : J.M. Bradford-Grieve in The Marine Fauna of New Zealand: Pelagic Calanoid Copepoda. National Institute of Water and Atmospheric Research (NIWA). NIWA Biodiversity Memoir, 111, 1999. [p.63, Fig.34].
Female: A, habitus (dorsal); B, forehead (lateral right side); C, Mx1; D, Mx2; E, P5.


Species Haloptilus ornatus - Plate 2 of morphological figuresissued from : T. Park in Antarct. Res. Ser. Washington, 1988, 47. [p.17, Fig.9].
Female: a, habitus (dorsal); b, forehead (lateral); c, idem (dorsal); d, urosome (dorsal); e, idem (lateral right side); f, genital segment (ventral); g, A2; h, Md; i, Md (cutting edge).


Species Haloptilus ornatus - Plate 3 of morphological figuresissued from : T. Park in Antarct. Res. Ser. Washington, 1988, 47. [p.18, Fig.10].
Female: a, Mx1; b, Mx2; c, Mxp; d, P1 (anterior).

Nota: A1 extending beyond distal end of caudal ramus by last 4 or 5 segments. Spermatheca more or less round when viewed laterally.


Species Haloptilus ornatus - Plate 4 of morphological figuresIssued from : G.O. Sars in Résult. Camp. Scient. Prince Albert I, 69, pls.1-127 (1924). [Pl.LXXIII, figs.1-5].
Female: 1, habitus (dorsal); 2, forehead (lateral); 3, Md; 4, Mx1; 5, Mx2.


Species Haloptilus ornatus - Plate 5 of morphological figuresissued from : R.B.S. Sewell in The John Murray Expedition, 1933-34, Scientific Reports, VIII (1), 1947. [p.195, Fig.50, H].
Female (from Arabian Sea): H, Mx2.


Species Haloptilus ornatus - Plate 6 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.27, Fig.38]. As Hemicalanus ornatus.
Male: 38, P5 (posterior view). Ps = left leg; Pd = right leg; Re = exopod; Ri = endopod; Se = outer spine; Si = inner spine; St = terminal spine.


Species Haloptilus ornatus - Plate 7 of morphological figuresissued from : T. Mori in The pelagic Copepoda from the neighbouring waters of Japan, 1937 (2nd edit., 1964). [Pl.39, Figs.5-8].
Female: 5, habitus (dorsal); 6, Mx2; 7, Md; 8, forehead (lateral).


Species Haloptilus ornatus - Plate 8 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.27, Figs.6, 7]. As Hemicalanus ornatus.
Female: 6, Mx1 (posterior view); 7, Mx2 (postrior view).


Species Haloptilus ornatus - Plate 9 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.27, Figs.14, 15, 21, 24]. As Hemicalanus ornatus.
Female: 14, Md (mandibular palp); 15, Md (mandibular blade); 21, exopod of P5; 24, endopod of P3.


Species Haloptilus ornatus - Plate 10 of morphological figuresissued from : G.D. Grice in Fish. Bull. Fish and Wildl. Ser., 1962, 61. [p.224, Pl.25, Figs.12-16].
Female (from 01°10'S, 134°57'W): 12, A2; 13, Md (biting edge); 14, Mx1; 15, Mx2; 16, P5.


Species Haloptilus ornatus - Plate 11 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.42, Fig.1]. As Hemicalanus ornatus.
Female: 1, habitus (dorsal).


Species Haloptilus ornatus - Plate 12 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.42, Figs.17, 19, 22]. As Hemicalanus ornatus.
Female: 17, exopodal segment 3 of P2; 19, urosome ( ventral); 22, endopod of A2.


Species Haloptilus ornatus - Plate 13 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.42, Fig.9]. As Hemicalanus ornatus.
Male: 9, habitus (dorsal).


Species Haloptilus ornatus - Plate 14 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.42, Fig.24]. As Hemicalanus ornatus.
Male: 24, endopod of A2.


Species Haloptilus ornatus - Plate 15 of morphological figuresissued from : J.M. Bradford-Grieve, E.L. Markhaseva & C.E.F. Rocha & B. Abiahy in South Atlantic Zooplankton, Edit. D. Boltovskoy. 1999. Vol.2. Copepoda. [p.1043, Fig. 7.276].
Female: Mx1.

Nota: Endopodite of Mx1 with 2 setae; exopodite with 10 setae; outer lobe 1 with 9 setae.

Compl. Ref.:
Wilson, 1942 a (p.189); Massuti Alzamora, 1942 (p.111); Sewell, 1948 (p.329, 504, 509, 522, 530, 532, 548, 558, 567); C.B. Wilson, 1950 (p.236); V.N. Greze, 1963 a (tabl.2); De Decker & Mombeck, 1964 (p.12); Grice & Hulsemann, 1965 (p.224); Mazza, 1966 (p.72); Mullin, 1966 (p.546, Table I, III, diet); Fleminger, 1967 a (tabl.1); Delalo, 1968 (p.138, Rem.: ?); Morris, 1970 (p.2301); Park, 1970 (p.478); Deevey, 1971 (p.224); Roe, 1972 (p.277, tabl.1, tabl.2); 1972 c (p.1032); Apostolopoulou, 1972 (p.328, 362); Deevey & Brooks, 1977 (p.156, tab.2, Station "S"); Carter, 1977 (1978) (p.36); Vives, 1982 (p.293); Kovalev & Shmeleva, 1982 (p.84); Cummings, 1984 (p.163, Table 2); Guangshan & Honglin, 1984 (p.118, tab.); Scotto di Carlo & al., 1984 (1044); Longhurst, 1985 (tab.2); Brenning, 1985 a (p.28, Table 2); 1986 (p.14, Rem.); Madhupratap & Haridas, 1986 (p.105, tab.1); Lozano Soldevilla & al., 1988 (p.60); Cervantes-Duarte & Hernandez-Trujillo, 1989 (tab.3); Heinrich, 1990 (p.18); Suarez & al., 1990 (tab.2); Scotto di Carlo & al., 1991 (p.270) ; Suarez & Gasca, 1991 (tab.2); Hernandez-Trujillo, 1991 (1993) (tab.I); Suarez, 1992 (App.1); Shih & Young,1995 (p.68); Hure & Krsinic, 1998 (p.102); Alvarez-Cadena & al., 1998 (t.4); Padmavati & al., 1998 (p.347); Suarez-Morales & Gasca, 1998 a (p108); Lopez-Salgado & al., 2000 (tab.1); Fernandez-Alamo & al., 2000 (p.1139, Appendix); Holmes, 2001 (p.13); Vukanic, 2003 (p.139, tab.1); Hsiao & al., 2004 (p.325, tab.1) Lo & al., 2004 (p.89, tab.1); Zuo & al., 2006 (p.162: tab.1); Lavaniegos & Jiménez-Pérez, 2006 (Tab.2, 4, Rem.); Dur & al., 2007 (p.197, Table IV); Morales-Ramirez & Suarez-Morales, 2008 (p.518); Williamson & McGowan, 2010 (p.273, Table 3, Pacific central gyres: N and S); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical/tropical); Mazzocchi & Di Capua, 2010 (p.424); Hsiao S.H. & al., 2011 (p.475, Appendix I); Medellin-Mora & Navas S., 2010 (p.265, Tab. 2)
NZ: 16

Distribution map of Haloptilus ornatus by geographical zones
Loc:
Afr. S (E), SE Atlantic, G. de Guinée, off I. Ascension SW, off I. Trindade, off I. St . Paul SE & E, Brésil S, off Maroc-Mauritanie, Canaries, Madère, off Portugal, Açores, off Amazone, I. Barbade, Caribbean Sea, Caribbean Colombia, Yucatan, Jamaïque, G. du Mexique, Floride, Mer des Sargasses, off Bermuda (Station "S"), off Irlande W, Médit. (Mer d'Alboran, NW, Ligurian Sea, Naples, Strait of Messina, Adriatique, Ionienne, Mer Egée, bassin Levantin), G. d'Akaba, Mer Rouge, Mer Arabe, Is. Laquedives, off I. La Réunion, Natal, Indien (equatorial central & South), Indonésie-Malaisie, Philippines, mers de Chine (East China Sea, South China Sea), off Taiwan, Taiwan (SW, E, N: Mienhua Canyon), Japon (Izu), Bikini, Californie, Basse Californie W, G. de Californie, G. de Tehuantepec, off Guatemala W, W Costa Rica, Pacif. (tropical), off Hawaii N, Costa Rica W, Galapagos SW, off Pérou, Pacif. SE (tropic.), Pacific central gyres: N and S, off Chile (33°S), Pacif. W (équatorial), off SW Australia, Nouvelle-Zélande (North Island SW, SE, Mer de Tasman N)
N: 85
Lg.:
(1) F: 5; (11) F: 4,5-4,4; (16) F: 4,8-4,3; (46) F: 4,8-4,6; M: 3,05-2,75; (69) F: 4,54; (90) F: 4,94-2,88; (91) F: 5-3; (101) F: 4,56; (199) F: 5,09-4,1; M: 3,04-2,96; (785) F: 4; (866) F: 3-5,33; (909) F: 4,75; 4,95; (1108) F: 4,94; 5,17; {F: 3,00-5,33; M: 2,75-3,05}
Rem.: epi-mesopelagic. Known to occur from the surface to below 150 m (Matthews, 1972). Sargasso Sea: 0-1000 m (Deevey & Brooks, 1977, Station "S");
Last update : 26/04/2013
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