Species Card of Copepod
Calanoida ( Order )
    Diaptomoidea ( Superfamily )
        Candaciidae ( Family )
            Candacia ( Genus )
Candacia norvegica  (Boeck, 1864)   (F,M)
Syn.: Candace norvegica Boeck, 1864; Giesbrecht, 1892 (p.425)
Ref.:
Giesbrecht & Schmeil, 1898 (p.131); Sars, 1902 (1903) (p.134, figs.F,M); Farran, 1908 b (p.85); Lysholm, 1913 (p.6); Lysholm & Nordgaard, 1921 (p.27); Sewell, 1932 (p.336, Rem.); Wilson, 1932 a (p.138, figs.F,M); Rose, 1933 a (p.251, figs.F,M); Jespersen, 1934 (p.119); 1940 (p.66); Lysholm & al., 1945 (p.41); Farran, 1948 b (n°13, p.3, figs.F,M); Marques, 1953 (p.120, figs.F,M); Grice & Jones, 1960 (p.288, figs.F,M, Rem.); Grice, 1963 (p.178, figs.F,M); Tanaka, 1964 c (p.246, figs.F,M, Rem.: variety); Silas, 1972 (p.649); Lawson, 1977 (p.71, tab.2,3,4, fig.4, 5); Huys & Boxshall, 1991 (p.66, fig.); Mazzocchi & al., 1995 (p.76, figs.F,M, Rem.); Chihara & Murano, 1997 (p.754, Pl.75,77: F,M); Bradford-Grieve & al., 1999 (p.885, 956, figs.F,M); Bradford-Grieve, 1999 b (p.172, figs.F,M, Rem., figs.183,193); Boxshall & Halsey, 2004 (p.85: fig.F); Vives & Shmeleva, 2007 (p.453, figs.F,M, Rem.)
Species Candacia norvegica - Plate 1 of morphological figuresissued from : O. Tanaka in Publs Seto Mar. Biol. Lab., 1964, XII (3). [p.247, Fig.228].
Female: a, habitus (dorsal); b, last thoracic segment and urosome (left lateral side); c, proximal portion of A1; d, P5. g, h, juvenile (urosome and P5, respectively).

Male: e, last thoracic segment and urosome (dorsal); ; f, P5.


Species Candacia norvegica - Plate 2 of morphological figuresissued from : Bradford-Grieve J.M. in The Marine Fauna of New Zealand: Pelagic Calanoid Copepoda. National Institute of Water and Atmospheric Research (NIWA). NIWA Biodiversity Memoir, 111, 1999. [p.173, Fig.123].
Female: A, last thoracic segments and urosome (dorsal); B, urosome (left side); C, exopod segment 3 of P2; D, P5.
Nota: The female genital somite of the Southwest Pacific specimens appears to differ slightly from the description by Sars (1903) in that the swollen anterior part is bordered by small hairs.

Male: E, last thoracic segments and urosome (dorsal); F, dorso lateral view of anterior urosome; G, right A1; H, P5.


Species Candacia norvegica - Plate 3 of morphological figuresissued from : G.O. Sars in An Account of the Crustacea of Norway. Vol. IV. Copepoda Calanoida. Published by the Bergen Museum, 1903. [Pl.LXXXIX].
Female.
Nota: R = rostrum; M = Md; m = Mx1.


Species Candacia norvegica - Plate 4 of morphological figuresissued from : G.O. Sars in An Account of the Crustacea of Norway. Vol. IV. Copepoda Calanoida. Published by the Bergen Museum, 1903. [Pl.XC].
Female & Male.
Nota: mp1 = Mx2; mp2 = Mxp


Species Candacia norvegica - Plate 5 of morphological figuresIssued from: M.G. Mazzocchi, G. Zagami, A. Ianora, L. Guglielmo & J. Hure in Atlas of Marine Zooplankton Straits of Magellan. Copepods. L. Guglielmo & A. Ianora (Eds.), 1995. [p.77, Fig.3.9.1].
Female: A, habitus (dorsal); B, urosome (dorsal).
Nota: Proportional lengths of urosomites and furca 48:21:15:16 = 100. In dorsal view, genital somite bears short bifid processes at midpoint of lateral margins (right process shorter than left; above bifid processes, genital somite bears tufts of hairs on either side.

Male: C, P5.
Nota: Proportional lengths of urosomites and furca 28:19:19:11:8:15 = 100.


Species Candacia norvegica - Plate 6 of morphological figuresIssued from: M.G. Mazzocchi, G. Zagami, A. Ianora, L. Guglielmo & J. Hure in Atlas of Marine Zooplankton Straits of Magellan. Copepods. L. Guglielmo & A. Ianora (Eds.), 1995. [p.78, Fig.3.9.2].
Female (SEM preparation): A, habitus (dorsal); B, genital somite (dorsal); C, urosome (lateral right side); D, detail of bifid process on genital somite (arrows in B and C); E, P5.
bars: A 0.200 mm; C 0.100 mm; B, E 0.050 mm; D 0.010 mm.


Species Candacia norvegica - Plate 7 of morphological figuresissued from : G.D. Grice & E.C. Jones in Pacific Science, 1960, XIV (3). [p.289, Figs.46-65].
Female: 46, habitus (lateral left side); 47, urosome (dorsal); 48, left genital spinous process (lateral view); 49, right genital spinous process (lateral view); 50, 51, edge biting of mandibular blades from one individual, showing unlike basal teeth; 52, Mx1; 53, Mx2; 54, P5 (with 2 external spinous processes on each foot); 55, P5 (showing 3 external spinous processes on one foot and 2 on the other); 63, urosome (dorsal); 64, 65, genital spinous process (left and right, respectively).

Male: 56, habitus (dorsal); 57, idem (lateral right side); 58, 4th and 5th thoracic segments and genital segment (dorsal); 59, genital segment (ventral); 60, 4th and 5th thoracic segments and genital segment (right lateral oblique view); 61, right A1 (segments 17 through 19); 62, P5.

Figures 46-62: Pacific specimens; 63-65: Atlantic specimen.


Species Candacia norvegica - Plate 8 of morphological figuresIssued from : R.B.S. Sewell in Mem. Indian Mus., 1932, X (continued). [p.337, Fig.110]. As Candacia norvegica var. tropica
Female (from Laccadive Sea): a, urosome (dorsal); b, proximal segments of A1; c, Mx2; d, P5.


Species Candacia norvegica - Plate 9 of morphological figuresissued from : R. Huys & G.A. Boxshall in Copepod Evolution. The Ray Society, 1991, 159. [p.66, Fig.2.2.13, A].
From Norway: Christiana Fjord, Sars collection: A, A2.

Compl. Ref.:
Wilson, 1942 a (part., p.175); Sewell, 1948 (p.350, 408, 496, 506); Mazza, 1966 (p.72); Roe, 1972 (p.277, tabl.1, tabl.2); Falconetti & Seguin, 1977 (p.188); Vives, 1982 (p.294); Madhupratap & Haridas, 1986 (p.105, tab.1); Go & al., 1997 (tab.1, as Canadacia: lapsus calami); Holmes, 2001 (p.30); Beaugrand & al., 2002 (p.179, figs.5, 6); Gaard & al., 2008 (p.59, Table 1, N Mid-Atlantic Ridge)
NZ: 16

Distribution map of Candacia norvegica by geographical zones
Loc:
Angola, G. de Guinée, off Maroc-Mauritanie, Canaries, Caraïbes, Mer des Sargasses, Woods Hole, off Terre-Neuve E, Mer du Groenland, Détr. de Davis S, Islande S, off Irlande S & W, off Ecosse W, Norvège, Is. Lofoten, Mer du Nord, baie de Lübeck, ? Médit. W, Indien (E Laccadives Is.), Pacif. N (central), Japon, ? [Philippines, mers de Chine, Japon, Aléoutiennes, Hawaii, Amér. centrale, Equateur W, Galapagos S, Chili (in Wilson, 1942 a , 1950, p.181) et (in Fagetti, 1962, p.33)], Détr. de Magellan, Australie SE, Nouvelle-Zélande (North Island SW, South Island E)
N: 35 (Sub-Antarct.: 1; Atlant. N: 23; Atlant. S: 2; Indien: 3; Pacif.: 6)
Lg.:
(36) F: 3,14-2,9; M: 2,96-2,82; (45) F: 3,35-3; M: 3,2-2,9; (65) F: 3,2; M: ±3,2; (120) F: 3,63; M: 3,25; (149) F: 3,24-2,28; M: 2,68; 3,23; (199) F: 2,74; (909) F: 2,75-3,1; M: 2,6; {F: 2,28-3,63; M: 2,60-3,25}
Rem.: epi-mesopelagic.
According to Tanaka (1964 c, p.248), the specimen might be regarded as a variety: C. norvegica intermedia.
Grice & Jones (1960, p. 290) point to differences in the details of the lateral spines of the female genital segment between Pacific and Atlantic forms. The Pacific females specimens agree with tropica Sewell (1932).
According to Mazzocchi & al. (1995) the females from the Straits of Magellan (Pacific area) belong to the variety tropica from Sewell (1932).
Last update : 24/07/2011
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