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Calanoida ( Order ) |
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Eucalanoidea ( Superfamily ) |
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Rhincalanidae ( Family ) |
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Rhincalanus ( Genus ) |
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Rhincalanus cornutus (Dana, 1849) (F,M) | |
| | | | | | | Syn.: | no Calanus rostrifrons (F) Dana,1849;
No Rhincalanus cornutus : Brady, 1883, p.41, figs.F); Giesbrecht, 1892 (part., p.153, 160, 774, figs.F,M); Vives & Shmeleva, 2007 (p.886, figs.F,M, Rem.);
Rhincalanus cornutus atlantica Schmaus, 1917 (p.306); 1927 (p.358, 360);
R. gigas : Sewell, 1914 a (p.203);
no R. cornutus : Tanaka, 1956 (p.271);
Rhincalanus cornutus f. typica : Mazza, 1967 (p.321);
Rhincalanus cornutus : Owre & Foyo, 1967 (part., p.37, no the drawings: figs.196-198) | | | | Ref.: | | | Giesbrecht & Schmeil, 1898 (part. p.23, Rem. F;M, figs.F); [Thompson & Scott, 1903 (p.233, 242)]; [Cleve, 1904 a (p.196, as Rhinocalanus cornutus)]; [A. Scott, 1909 (p.23)]; Wolfenden, 1911 (p.194); [Sewell, 1912 (p.353, 358)]; Willey, 1918 (p.186, figs. juv.F,M); Sars, 1925 (p.22); [Sewell, 1929 (p.58)]; Farran, 1929 (part., p.208, 220); ? Sciacchitano, 1930 (p.10, figs.F,M); Wilson, 1932 a (p.35, figs.F,M); [Dakin & Colefax, 1933 (p.204)]; Rose, 1933 a (p.71, figs.F,M); Gurney, 1934 a (p.209, N, juv.1); [Farran, 1936 a (p.79)]; [Mori, 1937 (1964) (p.27, figs.F,M)]; [Dakin & Colefax, 1940 (p.95, fig. juv.)]; Wilson, 1942 a ( part., p.205); Lysholm & al., 1945 (p.9); [Vervoort, 1946 (p.116, Rem.: anomalies, figs.F)]; [C.B. Wilson, 1950 (p.318)]; Farran & Vervoort, 1951 b (n°34, p.3, fig.F); [Krishnaswamy, 1953 (p.112)]; [King & Hida, 1955 (p.11)]; [Chiba, 1956 (p.51, figs.F,M); Yamazi, 1958 (p.147, Rem.)]; [Grice, 1962 (p.181, figs.M, juv.M)]; [Ganapati & Shanthakumari, 1962 (p.7, 16)]; [Kasturirangan, 1963 (p.14, figs.F,M)]; Vervoort, 1963 b (p.100, Rem.); Unterüberbacher, 1964 (p.15, figs.F, Rem.); [De Decker, 1964 (p.16, 24, 28)]; [De Decker & Mombeck, 1964 (p.13)]; [Chen & Zhang, 1965 (p.49, 129: Rem., figs.F,M)]; Dawson, 1966 (p.176); [Grice & Hulsemann, 1967 (p.14)]; [Mullin, 1966 (p.546)]; [Saraswathy, 1966 (1967) (p.76)]; Mazza, 1967 (p.93, figs.F,M, juv., Rem.); [Vinogradov, 1968 (1970) (p.79, 81, 91, 94, 277)]; Marques, 1968 a (p.4); [Itoh, 1970 (tab.1)]; [Morris, 1970 (p.2300, 2302)]; Corral Estrada, 1970 (p.81, figs.F, Rem.: "forma atlantica"); Rader, 1970 (p.75); Shih & al., 1971 (p.39); Bowman, 1971 b (p.18, Rem.); Björnberg, 1972 (p.14, figs., Rem.N); [Heinrich, 1973 (p.95)]; Marques, 1974 (p.12); Harding, 1974 (p.141, tab.2); Vives & al., 1975 (p.35, tab.II, XII); [Marques, 1976 (p.988)]; [Tranter, 1977 (p.596)]; [Carter, 1977 (1978) (p.35)]; [Goswami & Goswami, 1979 (p.103, figs.)]; Björnberg & al., 1981 (p.621, figs.F,M); Sazhina, 1982 (p.35, figs.N); [Marques, 1982 (p.754)]; Brodsky & al., 1983 (p.202, fig.F, Rem.F,M); Roe, 1984 (p.356); Baessa de Aguiar, 1986 (1989) (p.59, Rem.); 1988 (1990) (p.183, figs.F,M, Rem. F,M, juv.); [Zheng Zhong & al., 1984 (1989) (p.229, figs.F,M)]; [Jimenez-Perez & Lara-Lara, 1988]; [Yoo, 1991 (tab.1)]; [Park & Choi, 1997 (Appendix)]; [Chihara & Murano, 1997 (p.787, Pl.96: F,M)]; [Wong & al, 1998 (tab.2)]; Bradford-Grieve & al., 1999 (p.878, 912, fig.F); Goetze, 2003 (p.2322 & suiv.); G. Harding, 2004 (p.39, figs.F,M); Boxshall & Halsey, 2004 (p.176, p.177: fig.M); Mulyadi, 2004 (p.120, figs.F,M, Rem.); Phukham, 2008 (p.127, figs.F, M); Blanco-Bercial & al., 2011 (p.103, Table 1, Biol. mol, phylogeny) |  issued from : J. Corral Estrada in Tesis Doct., Univ. Madrid, A-129, Sec. Biologicas, 1970. [Lam.12, figs.5-6]. As Rhincalanus cornutus forma \"atlantica\". Female (from Canarias Is.): 5, posterior part cephalothorax and urosome (dorsal); 6, left P5. Nota: lateral spines of thoracic segments with anomalies.
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 issued from : T.E. Bowman in Smithson. Contr. Zool., 1971, 96. [p.24, Fig.19, c-d]. Female (from Florida): c, P5; d, distal segments of A1 (dorsal).
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 issued from : N. Phukham in Species diversity of calanoid copepods in Thai waters, Andaman Sea (Master of Science, Univ. Bangkok). 2008. [p.206, Fig.80]. Female (from W Malay Peninsula): a, habitus (dorsal); b, posterior part of thoracx and urosome (dorsal); c, forehead (dorsal); d, P5. Male: e-f, habitus (dorsal and lateral); g, last thoracic segments and urosome (dorsal); h, P5. Body length after drawings: F = 3.00 mm; M = 2.286 mm.
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 issued from : Mulyadi in Published by Res. Center Biol., Indonesia Inst. Sci. Bogor, 2004. [p.121, Fig.69]. Female (from Indonesian Seas): a, habitus (dorsal); b, forehead (ventral); c, P5. Male: d, habitus (dorsal); e, P5.
| | | | | Compl. Ref.: | | | Oliveira, 1945 (p.191); C.B. Wilson, 1950 (part., p.318); Gaudy, 1962 (p.93, Rem.: p.102); 1963 (p.20, Rem.); Björnberg, 1963 (p.23, Rem.); Grice & Hulsemann, 1965 (p.223); ? Mullin, 1966 (p.546, Table I, III, diet); Neto & Paiva, 1966 (p.20, Table III); Mazza, 1966 (p.69); Deevey, 1971 (p.224); Gueredrat, 1971 (p.300, fig.3, 10, Table 1, 2); Roe, 1972 (p.277, tabl.1, tabl.2); 1972 a (p.327); Brodsky, 1972 (P.256); Binet & al., 1972 (p.55, 68); Corral Estrada & Pereiro Muñoz, 1974 (tab.I); Harding, 1974 (p.141, Table 2, gut contents); [Patel, 1975 (p.659)]; Vives & al., 1975 (p.35, tab.II, IV); Deevey & Brooks, 1977 (p.256, tab.2, Station "S"); [Timonin & Voronina, 1977 (p.286, fig.4)]; [Frontier, 1977 a (p.14)]; Dessier, 1979 (p.204); [Stephen & Iyer, 1979 (p.228, tab.1, 3, 4, figs.3, 4)]; Vaissière & Séguin, 1980 (p.23, tab.1); Sreekumaran Nair & al., 1981 (p.493), Vives, 1982 (p.290); Kovalev & Shmeleva, 1982 (p.82); ? De Decker, 1984 (p.315, 362: carte); Cummings, 1984 (p.163, Table 2); [Guangshan & Honglin, 1984 (p.118, tab.)]; Kinzer & Schulz, 1985 (tab.4); Brenning, 1985 a (p.28, Table 2); Wishner & Allison, 1986 (tab.2); [Longhurst, 1985 (tab.2)]; [Petipa & Borichenko, 1985 (tab.1,2)]; [Renon, 1987 (tab.2)]; Lozano Soldevilla & al., 1988 (p.57); Suarez & al., 1990 (tab.2); Suarez Gasca, 1991 (tab.2); Suarez, 1992 (App.1); Ashjian & Wishner, 1993 (p.483, abundance, species group distributions); Seguin & al., 1993 (p.23); [Shih & Young, 1995 (p.69)]; Webber & Roff, 1995 (tab.1); Suarez-Morales & Gasca, 1997 (p.1525); Go & al., 1997 (tab.1, as corrutus); Alvarez-Cadena & al., 1998 (tab.2,3); Suarez-Morales, 1998 (p.345, Table 1); Suarez-Morales & Gasca, 1998 a (p.109); Mauchline, 1998 (tab.8, 23, 30); Neumann-Leitao & al., 1999 (p.153, tab.2); Lapernat, 2000 (tabl.3, 4); [Fernandez-Alamo & al., 2000 (p.1139, Appendix)]; Lopez-Salgado & al., 2000 (tab.1); Lo & al., 2001 (1139, tab.I); Fransz & Gonzalez, 2001 (p.255, tab.1); Sameoto & al., 2002 (p.13); Beaugrand & al., 2002 (p.179, figs.5, 6); [Hwang & al., 2003 (p.193, tab.2)]; [Kazmi, 2004 (p.229, Rem.: p.232]); [Yin & al., 2004 (p.3)]; [Smith & Madhupratap, 2005 (p.214, tab.6)]; Valdés & al., 2007 (p.103: tab.1); Jitlang & al., 2008 (p.65, Table 1); McKinnon & al., 2008 (p.844: Tab.I); Neumann-Leitao & al., 2008 (p.799: Tab.II, fig.6); Morales-Ramirez & Suarez-Morales, 2008 (p.513); Fernandes, 2008 (p.465, Tabl.2); Wishner & al., 2008 (p.163, Table 2, fig.8, oxycline); Gaard & al., 2008 (p.59, Table 1, N Mid-Atlantic Ridge); ? Pagano, 2009 (p.116); C.-Y. Lee & al., 2009 (p.151, Tab.2); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical/tropical); Goetze & Ohman, 2010 (p.2110, Table 1); Dias & al., 2010 (p.230, Table 1); Mazzocchi & Di Capua, 2010 (p.425); Medellin-Mora & Navas S., 2010 (p.265, Tab. 2); W.-B. Chang & al., 2010 (p.735, Table 2, abundance); Xu & Gao, 2011 (p.514, figs.3, 4, Table 2: optimal salinity); Salah S. & al., 2011 (Tableau 1) | | | | NZ: | 14 + 1 doubtful | | | | | | | | | | | | | Loc: | | | ? Afr. S (E), Namibie, off Tristan da Cunha E, Angola, Baia Farta, Congo, Atlant. (central équatorial), G. de Guinée, Ivorian shelf, Dakar, Is. du Cap Vert, off Mauritanie-Is. du Cap Vert NW, Maroc-Mauritanie, Cap Ghir, Canaries, off Madère, off Cap Finisterre W, Açores, Brésil (off Macaé, Cabo Frio, off Natal, Mouth of Tocantins, off Amazone), I. Barbade, Mer des Antilles, Caribbean Colombia, Yucatan, Jamaïque, Yucatan (Baie de l'Ascension), E Costa Rica, G. du Mexique, Caraïbes, Louisiana (off Grand Isle), Floride, off Bermuda, Mer des Sargasses, Station "S" (32°10'N, 64°30'W), off C. Hatteras, off E Cape Cod, G. du Maine, off Woods Hole, NouveIle-Ecosse, off Nova Scotia E, Féroé, Irlande W., Mer du Nord, Baie Ibéro-marocaine, Médit. (Mer d'Alboran, Marseille, Tyrrhénienne, Chenal Liguro-provençal, S Adriatic Sea), ? Mer Rouge, ? [Natal, Madagascar (Nosy Bé), Mer Arabe, Pakistan, India (Saurashtra coast, off Cochin, equatorial Indian Ocean, Madras), E India, Bay of Bengal, W Malay Peninsula (Andaman Sea), W Australia, Indonesian Seas (Bali Sea, Flores Sea, Banda Sea (Ambon Bay), S Celebes Sea: Manado Bay), Philippines, Viet-Nam (Baie de Cauda), Golfe du Tonkin, Hainan (Sanya Bay), mers de Chine (Yellow Sea, East China Sea, South China Sea), Taiwan, Okinawa, Corée S, Japon, Mer de Béring, Pacif. W (équatorial), Australie (Grande Barrière, Nouvelle-Galles du Sud), North West Cape), Nouvelle-Zélande, Pacif. (equatorial, Subarct.), off Hawaii S, G. de Californie, G. de Tehuantepec, Amér. centrale W, Clipperton Is., off Pérou, off Galapagos, off Pâques Is.] | | | | N: | 189 ? | | | | Lg.: | | | (14) F: 3,6-3,1; M: 2,6-2,4; [(28) F: 3,65-2,9; M: 2,9-2,45]; (54) F: 3,47; M: 2,7; [(92) F: 3,37]; [(101) F: 3,4-2,79; M: 2,95-2,69]; (131) F: 3,8-2,9; [(135) F: 3,5]; (180) F: 3,25-3,85; M: 2,57-2,7; (199) F: 4,16-2,81; M: 3,68-2,72; (237) F: 3,5; M: 2,5; [(290) F: 2,65-3,35; M: 2,6-2,8]; [(332) F: 3-2,6; M: 3,8-3,6]; [(333) F: 3,55-2,84; M: 2,63-2,5]; [(334) F: 3,6; M: 2,68]; (335) F: 3,45; [(530) F: 3,2]; [(795) F: 3,2)]; (1122) F: 3,3; M: 2,52; {F: 2,81-4,16; M: 2,40-3,68} | | | | Rem.: | meso-bathypelagic, sometimes near the surface. Overall Depth Range in Sargasso Sea: 0-2000 m. 1000-2000 m (Harding, 1974).
There are any confusions between R. cornutus and R. rostrifrons by the authors.
According to Bowman (1971), this species would be restricted to the Atlantic and R. rostrifrons to the Indo-Pacific (authors, locality records and dimensions between brackets). The indication of this tropical species in the North Sea appears exceptional.
Lo & al. (2001) report this species near Taiwan, as well as Rhincalanus rostrifrons, and Mulyadi (2004) in the Indonesian Seas. W. Zhang (pers. comm., nov. 2006) confirms the presence of R. cornutus in his samples from the Chinese Seas.
For Itoh (1970 a, fig.2, from co-ordonates) the Itoh's index value of the mandibular gnathobase = 240 (perhaps R. rostrifrons). | | | Last update : 07/05/2013 | |
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 Any use of this site for a publication will be mentioned with the following reference :
Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2012. - Diversity and Geographic Distribution of Marine Planktonic Copepods. Available at http://copepodes.obs-banyuls.fr/en
[Accessed May 22, 2013] © copyright 2005-2012 CNRS, UPMC
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