Species Card of Copepod
Calanoida ( Order )
    Clausocalanoidea ( Superfamily )
        Euchaetidae ( Family )
            Euchaeta ( Genus )
Euchaeta media  Giesbrecht, 1888   (F,M)
Syn.: Euchäta media Giesbrecht, 1892 (p.246, 263, 772, figs.F);
Euchaeta diegensis Esterly, 1911 (p.323, figs.F); Sewell, 1948 (p.556); Brodsky, 1950 (1967) (p.202, figs.F, Rem.); ? De Decker & Mombeck, 1964 (p.13); Hernandez-Trujillo, 1991 (1993) (tab.I);
? E. acuta pacifica Esterly, 1911 (p.324, fig.M); Brodsky, 1950 (1967) (p.199, figs.M); Roe, 1972 b (p.528: Rem.);
Paraeuchaeta media : Bradford & al., 1983 (p.43, figs.F, M, Rem.); Errhif & al., 1997 (p.422); Chihara & Murano, 1997 (p.801, Pl.107,110: F,M);
Pareuchaeta media : Heinrich, 1990 (p.17)
Ref.:
Giesbrecht & Schmeil, 1898 (p.39, Rem. F); Esterly, 1905 (p.160, figs.F); A. Scott, 1909 (p.66, figs.F); Bigelow, 1926 (p.230); Farran, 1929 (p.208, 238); Sewell, 1929 (p.133, 149); Candeias, 1926 (1929) (p.32, 49, figs.F); Rose, 1933 a (p.116, figs.F); Farran, 1936 a (p.89); Mori, 1937 (1964) (p.46, figs.F); Sewell, 1947 (p.116); Brodsky, 1950 (1967) (p.201, figs.F); C.B. Wilson, 1950 (p.214, figs.M, Rem.F,M); Vervoort, 1957 (p.77, 84, Rem.); Tanaka, 1958 (p.330, figs.F); Grice, 1962 (p.201, figs.F); Vervoort, 1963 b (p.161, Rem.); Paiva, 1963 (p.43); Gaudy, 1963 (p.23); Owre & Foyo, 1967 (p.54, figs.F); Park, 1968 (p.551, Redescr.M, figs.F,M); Tanaka & Omori, 1968 (p.221); Shih & al., 1971 (p.39); Vives, 1972 (p.212, figs.F); Tanaka, 1973 (p.135, fig.F); Park, 1975 c (p.5, figs.F,M); 1978 (p.207, figs.F,M); Dawson & Knatz, 1980 (p.6, figs.F,M); Björnberg & al., 1981 (p.633, 635, figs.F,M); Gardner & Szabo, 1982 (p.264, figs.F,M); Roe, 1984 (p.357); Park, 1995 (p.24, Rem.F,M, figs.F,M); Mauchline, 1999 (n°182, p.8, figs.F,M); Bradford-Grieve & al., 1999 (p.880, 925, figs.F,M); G. Harding, 2004 (p.53, figs.F,M); Mulyadi, 2004 (p.72, figs.F,M, Rem.); Vives & Shmeleva, 2007 (p.651, figs.F,M, Rem.); Blanco-Bercial & al., 2011 (p.103, Table 1, Biol. mol, phylogeny)
Species Euchaeta media - Plate 1 of morphological figuresissued from : T. Park in Antarctic Res. Ser. Washington, 1978, 27. [p.209, Fig.67].
Female: A, forehead (lateral); B, C,D,E, distal end of metasome and genital segment (left side, right side, dorsal and ventral); F, outer lobe of Mx1; G, P1 (anterior); H, P2 (anterior).


Species Euchaeta media - Plate 2 of morphological figuresissued from : T. Park in Antarctic Res. Ser. Washington, 1978, 27. [p.210, Fig.68].
Male: A, forehead (lateral); B, last metasomal and genital segments (lateral); C, P5 (anterior); D, middle part of exopod of left P5 (anterior); E, middle part of exopod of left P5 (medial).


Species Euchaeta media - Plate 3 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1995, 29. [p.123, Fig.13].
Female: a, forehead (left side); b, urosome (left); c, d, e, f, genital somite (left, dorsal, right, ventral, respectively); g, outer lobe of Mx1.

Male: h, forehead (left); i, j, exopod of left 5th leg (anterior, medial, respectively).


Species Euchaeta media - Plate 4 of morphological figuresIssued from : O. Tanaka in Publs Seto mar. Biol. Lab., 1958329, Fig.62].
Female: h, habitus (dorsal); i, forehead (lateral); j, last thoracic segment and urosomal segments 1 and 2 (lateral right side); k, outer margin of exopod of P2.


Species Euchaeta media - Plate 5 of morphological figuresissued from : J.M. Bradford, L. Haakonssen & J.B. Jillett in Mem. N.Z. Oceanogr. Inst., 1983, 90. [p.45, Fig.23]. As Paraeuchaeta media.
Female: A, habitus (lateral left side); B-D, genital segment (dorsal, left lateral, right lateral, respectively); E, exopod of P1; F, exopod segment 3 of P2.

Male: G, habitus (lateral left side); H, P5; I, terminal part of left P5 exopod.


Species Euchaeta media - Plate 6 of morphological figuresissued from : A. Scott in Siboga-Expedition, 1909, XIX a. [Plate XX, Figs.10-18].
Female (from Banda Sea): 10, habitus (dorsal); 11, forehead (lateral); 12, last thoracic and genital segments (left side); 13, genital segment (dorsal); 14, A1; 15, Mxp (end hair); 16, P1; 17, P2; 18, part of terminal spine of exopodite of P3.


Species Euchaeta media - Plate 7 of morphological figuresissued from : T. Mori in The pelagic Copepoda from the neighbouring waters of Japan, 1937 (2nd edit., 1964). [Pl.19, Figs.9-13].
Female: 9, habitus (dorsal; with eggs); 10, P2; 11, last thoracic and genital segmens (lateral); 12, forehead (lateral); 13, genital area.


Species Euchaeta media - Plate 8 of morphological figuresissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1911, 6 (14). [Pl.28, Fig.37].
Female (from San Diego Region): 37, forehead (lateral).


Species Euchaeta media - Plate 9 of morphological figuresissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1911, 6 (14). [Pl.29, Figs.49, 55].
Female: 49, part of last thoracic segment and genital segment (lateral, left side); 55, genital segment (ventral).
Nota: For Esterly, the urosome and genital segment resembles Euchaeta media Giesbrecht, but the genital convexity and the protuberances about the orifice are different; the right side of the segment as seen in outline is more irregular, with a smaller hnob-like protrusion near the posterior margin of the segment.


Species Euchaeta media - Plate 10 of morphological figuresissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1911, 6 (14). [Pl.31, Fig.92].
Female: 92, part of exopod of P2.
Nota: In exopod of P2, the middle spine of the outer margin reaches two-thirds of the distance from its base to the base of the third outer marginal spine (in Euchaeta media) it is about one-third as long).


Species Euchaeta media - Plate 11 of morphological figuresissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1911, 6 (14). [Pl.32, Fig.115]. As Euchaeta acuta var. pacifica. With doubt.
Male (from San Diego Region): 115, P5 (the left leg is at left).
Nota: P5 resemble those of Euchaeta acuta (see in Giesbrecht, 1892, Pl.16, fig.18, 21 (not 19)), but the process on the 2nd segment of exopodite of the left leg is not pyramidal and pointed, and in addition the endopodite of the right leg is broadened at the distal end and carries a flap-like appendage.


Species Euchaeta media - Plate 12 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.37, Figs.39, 40]. As Euchäta media.
Female: 39-40, urosome (lateral and ventral, respectively).


Species Euchaeta media - Plate 13 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.16, Fig.13]. As Euchäta media.
Female: 13, exopodal segments 2 and 3 of P2 (posterior view).


Species Euchaeta media - Plate 14 of morphological figuresissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1905, 2 (4). [p.161, Fig.25];
Female (from San Diego Region): a, habitus (lateral), with egg sac; b, forehead (lateral); c-d, genital segment (dorsal and right lateral side, respectively); e, P2; f, Mx1 (Le.1 = 1st lobe of outer margin; Li.1 = 1st lobe of inner margin; Ri.2 and 3 = fused 2nd and 3rd segments of endopod, bristles not shown); g, caudal rami (Si = inner marginal bristle).
Nota: A1 extend a little beyond the posterior border of the genital segment.1st outer marginal lobe of Mx1 with 8 bristles, 2nd basal with 3, fused 2nd and 3rd segments of the endopodite with 4. Outer border of 1st segment of exopodite of P1 concave; outer border of 3rd segment of P2 and its outer bristles different than in the following pair.


Species Euchaeta media - Plate 15 of morphological figuresissued from : G.D. Grice in Fish. Bull. Fish and Wildl. Ser., 1962, 61. [p.200, Pl.13, Figs.10-14].
Female (from 01°10'S, 134°57'W): 10, genital segment (dorsal); 11, same (ventral); 12, same (left side); 13, same (right side); 14, P2.
Characteristic swelling on the left side and the knob-like protrusion on the right side of the genital segment.
The middle spine on the outer edge of the 3rd exopodal segment of P2 is elongate and the spine on the 2nd exopodal segment reaches beyond the base of the proximal spine on the 3rd segment.


Species Euchaeta media - Plate 16 of morphological figuresissued from : J. Mauchline in ICES Ident. Leafl. Plankton, 1999, N°182. [p.3, Fig.2: 4a-4b].
Female (Northeast Atlantic): 4a, genital double-somite (right side); 4b, same (dorsal).


Species Euchaeta media - Plate 17 of morphological figuresissued from : J. Mauchline in ICES Ident. Leafl. Plankton, 1999, N°182. [p.4, Fig.3: 4c].
Male (Northeast Atlantic): 4c, terminal two segments, exopodal segments of left P5.


Species Euchaeta media - Plate 18 of morphological figuresissued from : G. Harding in Key to the adullt pelagic calanoid copepods found over the continental shelf of the Canadian Atlantic coast. Bedford Inst. Oceanogr., Dartmouth, Nova Scotia, 2004. [p.53].
Female & Male.


Species Euchaeta media - Plate 19 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.26, Fig.123].
Female: 123, forehead (lateral).


Species Euchaeta media - Plate 20 of morphological figuresissued from : T. Park in Smiths. Contr. Zool., 1975, 196. [p.6, Fig.2].
Female (G. of Mexico): a, forehead (lateral); b, genital segment (ventral).

Male: c, exopod of left P5 (anterior).


Species Euchaeta media - Plate 21 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. - Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.16, Fig.36]. As Euchäta spinosa.
Female: 36, outer margin of Mx2.


Species Euchaeta media - Plate 22 of morphological figuresissued from : O. Tanaka in I O B C Handbook, 1973, IV. [p.148, Fig.9: 4].
Female (from Indian Ocean): 4, genital segment (lateral).

Nota: Genital segment asymmetrical, shorter than the following three segments together. The right side with a low rounded process near the distal margin.


Species Euchaeta media - Plate 23 of morphological figuresissued from : Mulyadi in Published by Res. Center Biol., Indonesia Inst. Sci. Bogor, 2004. [p.73, Fig.41].
Female (from 07°29'S, 121°05'E): a, habitus (dorsal); b, posterior part of last thoracic segment and genital complex (dorsal); c, same (lateral left side); d, forehead (lateral); e, P1; f, P2; g, left caudal ramus (dorsal).

Male: h, habitus (dorsal); i, forehead (lateral); j, P5; k, exopod of left P5.

Compl. Ref.:
Cleve, 1904 a (p.190); Sewell, 1948 (p.391); Ahlstrom & Thrailkill, 1963 (p.57, Table 5, abundance); Grice, 1963 a (p.495); Gaudy, 1963 (p.23, Rem.); Unterüberbacher, 1964 (p.23); De Decker & Mombeck, 1964 (p.12); Mazza, 1966 (p.70); 1967 (p.331); Fleminger, 1967 a (tabl.1); Morris, 1970 (p.2300); Park, 1970 (p.476); Gueredrat, 1971 (p.300, fig.2, 10, Table 1, 2); Deevey, 1971 (p.224); Roe, 1972 (p.277, tabl.1, tabl.2); 1972 b (p.528); Harding, 1974 (p.141, tab.2, gut contents); Vives & al., 1975 (p.42, tab.II); Tranter, 1977 (p.596); Deevey & Brooks, 1977 (p.256, tab.2, Station "S"); Carter, 1977 (1978) (p.35); Dessier, 1979 (p.205); Hayward, 1980 (p.295, Table 2, vertical distribution, feeding); Vives, 1982 (p.291); De Decker, 1984 (p.316); Guangshan & Honglin, 1984 (p.118, tab.); Cummings, 1984 (p.163, Table 2); Longhurst, 1985 (tab.2); Madhupratap & Haridas, 1986 (p.105, tab.1); Rudyakov, 1986 (tab.1); Ambler & Miller, 1987 (tab.2, 3, 4, 5); Lozano Soldevilla & al., 1988 (p.58); Jimenez-Perez & Lara-Lara, 1988; Hernandez-Trujillo, 1989 a (tab.1); Cervantes-Duarte & Hernandez-Trujillo, 1989 (tab.3); Suarez & al., 1990 (tab.2); Suarez & Gasca, 1991 (tab.2); Hernandez-Trujillo, 1991 (1993) (tab.I); Suarez, 1992 (App.1); Hattori, 1991 (tab.1, Appendix); Mauchline, 1992 a (p.2); Shih & Young, 1995 (p.70); Go & al., 1997 (tab.1); Alvarez-Cadena & al., 1998 (tab.1,2,3,4); Suarez-Morales, 1998 (p.345, Table 1); Suarez-Morales & Gasca, 1998 a (p.109); Mauchline, 1998 (tab.42); Lavaniegos & Gonzalez-Navarro, 1999 (p.239, Appx.1); Lapernat, 2000 (tabl.3, 4); Haury & al., 2000 (p.69, Table 1, Fig.7); Lopez-Salgado & al., 2000 (tab.1); Rebstock, 2001 (tab.2); Holmes, 2001 (p.52); Beaugrand & al., 2002 (p.179, figs.5, 6); Rebstock, 2002 (p.71, Table 3, 5, 6, Figs.2, 3: climatic variability); Hsiao & al., 2004 (p.326, tab.1); Pusch & al., 2004 (251, tab.3); Lo & al., 2004 (p.89, tab.1); Berasategui & al., 2005 (p.313, fig.2); Lavaniegos & Jiménez-Pérez, 2006 (p.143, tab.2, 3, Rem.); Mackas & al., 2006 (L22S07, Table 2); Dur & al., 2007 (p.197, Table IV); Neumann-Leitao & al., 2008 (p.799: Tab.II, fig.6); Morales-Ramirez & Suarez-Morales, 2008 (p.520); Fernandes, 2008 (p.465, Tabl.2); C.-Y. Lee & al., 2009 (p.151, Tab.2); Galbraith, 2009 (pers. comm.); Williamson & McGowan, 2010 (p.273, Table 3, Pacific central gyres: N and S); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical/tropical); Hidalgo & al., 2010 (p.2089, Table 2): Medellin-Mora & Navas S., 2010 (p.265, Tab. 2); Hsiao S.H. & al., 2011 (p.475, Appendix I)
NZ: 18

Distribution map of Euchaeta media by geographical zones
Species Euchaeta media - Distribution map 3issued from : L. Haury, C. Fey, C. Newland & A. Genin in Progress in Oceanography, 2000, 45 [p.83, Fig.7].
Vertical distribution of Euchaeta media female from day and night tows taken over (solid lines) and away from (dashed lines) the seamount Fieberling Guyot (32°25'N, 127°47'W).
The horizontal line at 500 m marks the approximate depth of the summit plain. Note the reduction in abundance at night over the seamount.
Species Euchaeta media - Distribution map 4issued from : G.A. Rebstock in Global change Biology, 2002, 8. [p.77, Fig.2 a].
Climatic regime shifts and decadal-scale variability in calanoid copepod populations off southern California (31°-35°N, 117°-122°W, from years 1950 to 2000.
Cumulative sums of nonseasonal anomalies from the long-term means of copepod abundance .
A negative slope indicates a period of below-average anomalies; a positive slope indicates a period of above-average anomalies. Abrupt changes in slope indicate step changes. Step changes are marked with arrows (downward -pointing for decreases).
The October 1966 cruise (prior to the increase in sampling depth), March 1976 cruise (prior to the 1976-77 climatic regime shift), and October 1988 cruise (prior to the hypothesized 1989 climatic regime shift) are marked with vertical lines.
Species Euchaeta media - Distribution map 5issued from : T.L. Hayward in Mar. Biol., 1980, 58. [p.299, Fig.2].
Plots of day (open circle) and night (filled circle) depth distributions of Euchaeta media in the North Pacific central gyre (main sampling location: 25°N, 155°W with drogue), September 1968.
Loc:
Afr. S (E), Namibie, Congo, G. de Guinée, Is. du Cap Vert, off Mauritanie-Is. du Cap Vert NW, Maroc-Mauritanie, Great meteor Seamount, Canaries, off Madère, Argentine, Brésil, off Amazone, Caraïbes, Mer des Antilles, Caribbean Colombia, Yucatan, G. du Mexique, Floride, off Bermuda: Station "S" (32°10'N, 64°30'W), off E Cape Cod, G. du Maine (rare), off Rockall Is., Portugal, off Cap Finisterre W, Baie Ibéro-marocaine, Médit. (baie d'Alger), Mer Arabe, Natal, Indien, Indien S (convergence subtropicale), Is. Nicobar (Nankauri Harbour), Bay of Bengal, Australie W, Indonesia-Malaysia, Flores Sea, Philippines, mers de Chine (East China Sea, South China Sea), Taiwan, Kuroshio Current, Okinawa, Japon, off Sanriku, California, W Baja California, G. de Californie, off Guatemala W, W Costa Rica, Pacif. W (equatorial), Pacific central gyres: N and S, Australia (Grande Barrière), Nouvelle-Zélande, Bikini, Hawaii, off Hawaii NE, Pacif. central subtropical N, Costa Rica, Pacif. SE (tropic.), off Chili, Chili (N-S)
N: 111
Lg.:
(3) F: 4,58-3,64; M: 3,88-3,56; (5) F: 3,75; (9) F: 4,1-3,3; M: 3,7-3,3; (14) F: 4-3,3; M: 3,55-3,05; (19) F: 4,24-3,6; M: 3,74-3,42; (20) F: 4,52-3,72; M: 4,2-3,36; (22) F: 4; 3,75-3,3; 4,22; M: 4-3,5; (34) F: 3,8; (47) F: 3,5; (63) F: 4,64-4,12; M: 3,86-3,61; (72) F: 4,51-4,03; M: 4,03-3,79; (73) F: 4,21; (91) F: ± 3,5; (98) F: 4,82-3,7; M: 3,94-3,39; (99) F: 4,32-4; M: 3,63; (101) F: 3,52; (142) F: 3,3; (143) F: 4,22; (199) F: 4,26-3,8; M: 3,88-3,57; (244) F: 3,6; (432) F: 3,77-3,68; (1122) F: 4,5; M: 4,02; {F: 3,30-4,82; M: 3,05-4,20}
Rem.: epi- bathypelagic. Considered as vetical migrator. Sargasso Sea: 0-1000 m (Deevey & Brooks, 1977, station "S"); 2000-1000 m (Harding, 1974).
For Vervoort (1963 b, p.161) this species is closely related E. acuta Giesbrecht, 1892. For Park (1995, p.22) this species belongs to ‘’acuta ’’ Group. The two species can be separated by the following characters: E. media : 1- Posterolateral thoracic border slightly produced and triangular in outline. 2- Tubercle on the left side of the genital somite rounded. On the right side a slight widening of the outline, terminating into a rounded protuberance just behind the middle of the genital somite. 3- A1 reach slightly beyond the end of the thorax. 4- Appendicular seta of the caudal rami lengthened and thickened at its base; remaining setae straight, as long as the urosome. (Cf the same characters in E. acuta.
Last update : 13/05/2013
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