Species Card of Copepod
Calanoida ( Order )
    Clausocalanoidea ( Superfamily )
        Euchaetidae ( Family )
            Paraeuchaeta ( Genus )
Paraeuchaeta hanseni  (With, 1915)   (F,M)
Syn.: Euchaete Hansenii With, 1915 (p.181, Descr.F, figs.F);
Euchaete sarsi (M): With, 1915 (p.178, fig.M, no F);
Pareuchaeta Hanseni : Sars, 1925 (p.115, figs.F); Rose, 1929 (p.25); 1933 a (p.120, figs.F); Jespersen, 1934 (p.80);
Paraeuchaeta withi Sewell, 1947 (p.131, figs.M); 1948 (p.331, 521, 527, 532); Silas, 1972 (p.646); Euchaeta withi : Owre & Foyo, 1967 (p.57, figs.M, Rem.);
Pareuchaeta withi : Tanaka & Omori, 1967 (p.247); 1968 (p.258, figs.M);
Pareuchaeta hanseni : C.B. Wilson, 1950 (p.282); Tanaka, 1958 (p.361, figs.F, no M); Tanaka & Omori, 1967 (p.246); 1968 (p.237, figs.F); Vaissière & Séguin, 1980 (p.23, tab.1); Kovalev & Shmeleva, 1982 (p.83); Vives, 1982 (p.291); Lozano Soldevilla & al., 1988 (p.58); Mauchline, 1998 (tab.42);
Euchaeta hansenii : Owre & Foyo, 1967 (p.53, figs.F,M); Park, 1975 c (p.17, figs.F,M, Rem.); 1978 (p.246, figs.F,M, Rem.);
Euchaeta hanseni : Grice & Hulsemann, 1967 (p.15); 1968 (tab.2); Deevey & Brooks, 1977 (p.256, tab.2, Station "S"); Mauchline, 1992 a (p.2, 3); 1999 (n°182, p.8, figs.F,M)
Ref.:
Sewell, 1947 (p.121, Rem.); Bradford & al., 1983 (p.39, figs.F,M, Rem., as hansenii); Park, 1994 (p.322); 1995 (p.57, Rem.F,M, figs.F,M); Chihara & Murano, 1997 (p.800, Pl.108,114: F,M); Bradford-Grieve & al., 1999 (p.880, 925, figs.F,M); Vives & Shmeleva, 2007 (p.663, figs.F,M, Rem.)
Species Paraeuchaeta hanseni - Plate 1 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1995, 29. [p.161, Fig.51].
Female: a, forehead (left side); b, urosome (left); c, genital somite (left); d, outer lobe of Mx1; e, P1 (anterior); f, P2 (anterior).
Nota: Similar in habitus to P. comosa but distinguishable from it.
Laterally, rostrum relatively short and thick, pointing obliquely forward in parallel to anterodorsal margin of forehead. Genital somite with a low dorsal hump just anterior to middle. Genital flange with emarginate ventral margin and produced distally into an extraordinary large lobe. Posterior edge of genital field produced into a relatively small but high conical process followed by short and convex posterior margin of genital prominence, which is about 1/2 length of posterior ventral wall of somite. A low ridge is found close to anterodorsal margin on each side of genital somite. Cephalosomal appendages including Mx1 as in P. malayensis.

Male: g, forehead (left); h, exopod of P1 (anterior); i, P2 (anterior); j, distal exopodal segments of left 5th leg (anterior).


Species Paraeuchaeta hanseni - Plate 2 of morphological figuresissued from : T. Park in Antarctic Res. Ser. Washington, 1978, 27. [p.247, Fig.95]. As Euchaeta hansenii.
Female: forehead (lateral); B, C, D, E, distal end of metasome and genital segment (left side, right side, dorsal, and ventral, respectively); F, outer lobe of Mx1; G, P1; H, P2; P1-2: legs (anterior).


Species Paraeuchaeta hanseni - Plate 3 of morphological figuresissued from : T. Park in Antarctic Res. Ser. Washington, 1978, 27. [p.248, Fig.96]. As Euchaeta hansenii.
Male: A, forehead (lateral); B, C, distal end of metasome and genital segment (dorsal and left side, respectively); D, P1; E, P2; F, P5; G, H, I, distal part of exopod of left P5 (anterior, medial, and lateral, rspectively). P1-5: legs (anterior).

Nota from Bradford & al. (p.42):
- P1 exopod male: Aa minute; Bb < 1/2 BC; Cc > 1/2 BC.
- P2 exopod male: Aa = 1/3 AB; Bb = 1/3 BC; Cc = 1/3 CD; Dd = 1/5 CD. (see code of lengths outer spines in the Genus' figure, or in Paraeuchaeta sp. A).


Species Paraeuchaeta hanseni - Plate 4 of morphological figuresissued from : O. Tanaka & M. Omori in Publs Seto Mar. Biol. Lab., 1968, XVI (4). [p.259, Fig.22]. As Pareuchaeta withi.
Male: A, forehead (lateral); B, last thoracic segment and genital segment (lateral left side); C, distal segments of left P5; D, E, serrated lamella.
Nota: The urosome segments and furca are in the proportional lengths as 15:32:27:18:3:5 = 100. Prosome and urosome are in the proportional lengths as 70 to 30.


Species Paraeuchaeta hanseni - Plate 5 of morphological figuresIssued from : O. Tanaka in Publs Seto mar. Biol. Lab., 1958, VI (3). [p.362, Fig.78 a-f]. As Pareuchaeta hanseni].
Female: a, habitus (dorsal); b, forehead (lateral); c, last thoracic segment and genital segment (lateral left side); d, genital complex (ventral); e, exopod of P1; f, exopod of P2.
Nota: The firstthree urosomal segments in 0.01 mm are: 106:56:56.


Species Paraeuchaeta hanseni - Plate 6 of morphological figuresissued from : J.M. Bradford, L. Haakonssen & J.B. Jillett in Mem. N.Z. Oceanogr. Inst., 1983, 90. [p.41, Fig.20].
Female: A, habitus (lateral left side); B, genital segment (dorsal); C, idem (lateral left side); D, exopod of P1; E, exopod segment 3 of P2.

Nota: The external spine of P1 exopod segment 1 is longer in the south-west Pacific specimen than in the original description.
- P1 Exopod female: Aa = 1/2 AB; Bb ≤ 3/4 BC; Cc ≤ BC.
- P2 exopod female: Aa < AB; Bb ≤ 1/2 BC ; Cc = 3/4 CD; Dd = 1/5 CD.


Species Paraeuchaeta hanseni - Plate 7 of morphological figuresIssued from : G.O. Sars in Résult. Camp. Scient. Prince Albert I, 69, pls.1-127 (1924). [Pl.XXXI, figs.15-18]. As Pareuchaeta hanseni.
Female: habitus (lateral left side); 16, forehead (lateral); 17, posterior part cephalothorax and genital gegment (lateral left side); 18, genital complex (ventral).


Species Paraeuchaeta hanseni - Plate 8 of morphological figuresissued from : R.B.S. Sewell in The John Murray Expedition, 1933-34, Scientific Reports, VIII (1), 1947. [p.132, Fig.30]. As Paraeuchaeta withi.
Male (from central part of Arabian Sea): A, P2; B, clasping organ of left P5; C, idem (another view).
Nota: The proportional lengths of the cephalothorax and abdomen as 67 to 33. In P1 the marginal spine on 1st segment of the exopod is absent.

Nota: With (1915, p.182) points out that this species is nearly related to P. sarsi (Farran), but is easily distinguished by the structure of the genital somite, by 9 setae in the outer lobe of Mx1.

This form seems to be identical with the male of P. hanseni described by Park (1975 c, p.17).


Species Paraeuchaeta hanseni - Plate 9 of morphological figuresissued from : J. Mauchline in ICES Ident. Leafl. Plankton, 1999, N°182. [p.3, Fig.2: 15a]. As Euchaeta hanseni.
Female (Northeast Atlantic): 15a, genital double-somite (left side).


Species Paraeuchaeta hanseni - Plate 10 of morphological figuresissued from : J. Mauchline in ICES Ident. Leafl. Plankton, 1999, N°182. [p.4, Fig.3: 15b]. As Euchaeta hanseni.
Male (Northeast Atlantic): 15b, terminal two segments, exopodal segments of left P5.


Species Paraeuchaeta hanseni - Plate 11 of morphological figuresissued from : C. With in The Danish Ingolf-Expedition, 1915, III (4). [p.181, Text-Fig.52]. As Euchaeta Hansenii.
Female (from 60°00N, 10°35'W): a, genital segment (lateral); b, genital area (ventral).


Species Paraeuchaeta hanseni - Plate 12 of morphological figuresissued from : C. With in The Danish Ingolf-Expedition, 1915, III (4). [Pl.VI, 7b]. As Euchaete Sarsi.
Male (from 61°30'N, 17°08'W): 7b, 2nd and 3rd exopodal segments of left P5.


Species Paraeuchaeta hanseni - Plate 13 of morphological figuresissued from : T. Park in Smiths. Contr. Zool., 1975, 196. [p.17, Fig.14]. As Euchaeta hansenii.
Female (G. of Mexico): a, forehead (lateral); b, genital segment (lateral).
gf = genital flange.

Male: c, forehead (lateral); d, exopod of left P5 (anterior); e, same (lateral); f, same (medial).
dp = digitiform process.
The male described is believed to represent the hitherto unknown male of E. hansenii, for it usually found along with the female of that species and is the closet to it in body size.


Species Paraeuchaeta hanseni - Plate 14 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1967, 1, Crustacea, 1: Copepoda. [p.53, Figs.320-321]. As Euchaeta hansenii].
Female (from Florida Current): 320, outer lobe of Mx1; 321, P2.


Species Paraeuchaeta hanseni - Plate 15 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1967, 1, Crustacea, 1: Copepoda. [p56, Figs.351-353]. As Euchaeta withi.
Male (from 23°40'N, 83°17'W): 351, habitus (lateral); ; 352, P5; 353, terminal segment of left P5.

Nota: The specimen male was collected from 800-900 m. With (1915) descibed it as the male of Euchaete Sarsi but expressed doubt as belonging to ''E. sarsi'' or another species of similar size nearly related to it.
E. withi (Sewell, 1947), known exclusively from the male, seems to be identical with the male of E. hansenii described.


Species Paraeuchaeta hanseni - Plate 16 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1967, 1, Crustacea, 1: Copepoda. [p57]. As Euchaeta withi.
Comparative bodily proportions males.
Gerda specimen: sample with the Isaacs-Kidd Mid-Water Trawl from 23°40'N, 83°17'W by Owre & Foyo (1964)


Species Paraeuchaeta hanseni - Plate 17 of morphological figuresParaeuchaeta hansenii Female:
1 - See key to species Groups and independent species of Paraeuchaeta (p.30): malayensis species Group.
2 - Outer spine of 2nd exopodal segment (or the 2nd of the first 2 exopodal segments forming a proximal, compound segment) of P1 normally developed (Fig.51-h).
3 - Outer lobe of Mx1 with 7 large distal setae extending laterad and 2 medium-sized, densely plumose proximal setae extending proximad along lateral side of appendage (Fig.51-d).
4 - Genital segment without conspicuous conical processes on left side at anterior base of genital prominence (Fig.51-c).
5 - Laterally, rostrum spiniform (Fig.51-a).
6 - Laterally, posterior ventral wall of genital somite very short (Fig.51-c).
7 - Laterally, posterior margin of genital prominence shorter than posterior ventral wall of genital somite (Fig.51-c).

Compl. Ref.:
Sewell, 1948 (p.501, 521, 524, 527, 530); Grice, 1963 a (p.495); De Decker & Mombeck, 1964 (p.13); Gueredrat & Friess, 1971 (p.187, fig.2); Roe, 1972 (p.277, tabl.1, tabl.2); 1994 (p.320); Lapernat, 2000 (tabl.3, 4); Razouls & al., 2000 (p.343, Appendix); Shimode & al., 2005 (p.113 + poster); Morales-Ramirez & Suarez-Morales, 2008 (p.520); Park & Ferrari, 2009 (p.143, Table 5, Appendix 1, as Paraeuchaeta hansenii, biogeography); Mazzocchi & Di Capua, 2010 (p.425); Teuber & al., 2013 (p.1, Table 2, respiration rates);
NZ: 15

Distribution map of Paraeuchaeta hanseni by geographical zones
Loc:
sub-Antarct. ( S Australia), South Africa (E), off NW Cape Verde Is., off Morocco-Mauritania, Canary Is., SW Atlant., Barbados Is., Caribbean Sea, G. of Mexico, Florida, off Bermuda: Station "S" (32°10'N, 64°30'W), Azores, Davis Strait, Norwegian Sea, off Rockall Is., NW Scotland, Medit. W (Ligurian Sea, Tyrrhenian Sea), Arabian Sea, Indian, Madagascar, Bay of Bengal, Indonesia-Malaysia, Philippines, China Seas (South China sea, East China Sea), Japan, Tasman Sea, off British Columbia, California, Central America, W Costa Rica, Pacif. (equatorial), SE Pacif., off Peru, off Juan Fernandez Is.
N: 28
Lg.:
(1) F: 9,4; (3) F: 9,7-8,1; M: 8,9-8,1; (7) F: 8,9; (11) M: 8,4; (19) F: ?5,41-9,25; M: 8,08-8,58; (63) F: 9,99-8,65; M: 8,24; 8,45-8,36; (70) F: 9,1-9; M: 8,5; (99) F: 8,7; (140) M: 8,4; (199) F: 9,4-8,32; {F: 8,10-9,99; M: 8,08-8,90}
Rem.: bathypelagic (2000-3000 m), 625-950 m (in Roe, 1972).
Sampling depth (sub-Antarct.) : 0-2000 m. Sargasso Sea: Deevey & Brooks, 1977, station "S").
Park (1995, p.58) found this species in the Atlantic between 47°N and 36°S; in the eastern and central Pacific between 38°N and 47°S; in the western Pacific including the East and South China seas, Malay Archipelago, and Tasman Sea between 35°N and 50°S; in the Bay of Bengal and the Indian Ocean between 10°N and 25°S.
Last update : 27/01/2016
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