Species Card of Copepod
Calanoida ( Order )
    Arietelloidea ( Superfamily )
        Heterorhabdidae ( Family )
            Heterorhabdus ( Genus )
Heterorhabdus clausi  (Giesbrecht, 1889)   (F,M)
Syn.: Heterochäta clausii Giesbrecht, 1889; 1892 (p.372, 382, 773, figs.F,M);
Heterorhabdus (Heterorhabdus) clausii : Vervoort, 1965 (p.121, Rem.)
Ref.:
Giesbrecht & Schmeil, 1898 (p.115, Rem. F,M); Thompson & Scott, 1903 (p.235, 249); Esterly, 1905 (p.185, fig.M; A. Scott, 1909 (p.130, Rem.); Wolfenden, 1911 (p.303); Sewell, 1932 (p.299, figs.F); Rose, 1933 a (p.204, figs.F,M); Lysholm & al., 1945 (p.36); Brodsky, 1950 (1967) (p.353, figs.F,M); C.B. Wilson, 1950 (p.239, fig.F); Vervoort, 1957 (p.134); Gardner & Szabo, 1982 (p.364, figs.F,M); Bradford-Grieve & al., 1999 (p.883, 944, figs.F,M, as clausii); Bradford-Grieve,1999 b (p.79, figs.F,M, Rem., figs.174, 191); Park, 2000 (p.121, figs.F,M, Rem.); Vives & Shmeleva, 2007 (p.310, figs.F,M, Rem.)
Species Heterorhabdus clausi - Plate 1 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 2000, 31. [p.240, Fig.88].
Female: a, habitus (left side); b, c, urosome (left, dorsal, respectively); d, e, genital somite (left, right, respectively). gfl = genital flange.


Species Heterorhabdus clausi - Plate 2 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 2000, 31. [p.241, Fig.89].
Female: a, genital somite with operculum open (left side); b, genital somite (ventral); c, genital somite with operculum open (right).

Male: d, P5 (anterior); e, basipod of left P5 (anterior).


Species Heterorhabdus clausi - Plate 3 of morphological figuresIssued from : R.B.S. Sewell in Mem. Indian Mus., 1932, X (continued). [p.301, Fig.98, c-d].
Female (from off SE Sri Lanka): c, exopod of P2; d, exopod of P3.


Species Heterorhabdus clausi - Plate 4 of morphological figuresissued from : C.O. Esterly in Univ. Calif. Publs. Zool., 1905, 2 (4). [p.185, Fig.39].
Male (from San Diego Region): P5.

Nota: 2nd basal of the right P5 with a long lamellar process, the 2nd segment of the exopod with a shorter projection on the inner border, 3rd relatively longer ( especially on the left P5).


Species Heterorhabdus clausi - Plate 5 of morphological figuresissued from : C.B. Wilson in Bull. U.S. natn Mus., 100, 14 (4). [Pl.11, Fig.120].
Female (from off Honshu, Japan and Philippines): 120, P5.


Species Heterorhabdus clausi - Plate 6 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf. 20, Fig.28 ]. As Heterochäta clausii.
Female: 28, P5 (endopod and exopod).


Species Heterorhabdus clausi - Plate 7 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf. 20, Fig.2]. As Heterochäta clausii.
Male: 2, A1 (distal segments).


Species Heterorhabdus clausi - Plate 8 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf. 20, Figs.37, 38]. As Heterochäta clausii.
Male: 37, left P5 (anterior view); 38, right P5 (anterior view).
B2 = basipod 2 (= basis); Ri = endopod; Re = exopod.

Compl. Ref.:
Sewell, 1948 (p.503, 521, 547, 558); De Decker & Mombeck, 1964 (p.12); Grice & Hulsemann, 1965 (p.224); 1967 (p.18); Fleminger, 1967 a (tabl.1); Deevey, 1971 (p.224); Deevey & Brooks, 1977 (p.256, tab.2, Station "S"); Vives, 1982 (p.293); Guangshan & Honglin, 1984 (p.118, tab.); Lozano Soldevilla & al., 1988 (p.59); Suarez-Morales & Gasca, 1998 a (p.109); Razouls & al., 2000 (p.343, Appendix); Holmes, 2001 (p.16); Beaugrand & al., 2002 (p.179, figs.5, 6); Lo & al., 2004 (p.89, tab.1); Galbraith, 2009 (pers. comm.)
NZ: 19

Distribution map of Heterorhabdus clausi by geographical zones
Loc:
sub-Antarct. (Tasmanie S), Afr. S (E), off Tristan da Cunha E, G. de Guinée, Canaries, Brésil, Mer des Sargasses, off Bermuda (Station "S"), off Irlande SW, off Ecosse E, Groenland, Féroé, W Medit. (Ligurian Sea in Mazzocchi & Di Capua, 2010; Minicoy Is., Indien, Indonésie-Malaisie, Philippines, mers de Chine (East China Sea), Taiwan (N: Mienhua Canyon), Japon (off Honshu), Pacif. W (équatorial), Pacif. tropical central & E (N-S), off British Columbia, Californie, San Diego, off Galapagos E & W, Tasmanie S
N: 24
Lg.:
(16) F: 2,65-2,6; M: 2,41-2,2; (22) F: 2,4; M: 2,5-2,2; (46) F: 2,4; M: 2,4-2,2; (142) M: 2,5-2; (824) F: 2,64-2,32; M: 2,4-2,24; {F: 2,32-2,65; M: 2,00-2,50}
Rem.: Abyssalis" Group.
bathypelagic. Overall Depth Range in Sargasso Sea: 0-2000 m (Deevey & Brooks, 1977, Station "S");
Sampling depth (sub-Antarct.) : 0-750 m.
Last update : 22/11/2012
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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2012. - Diversity and Geographic Distribution of Marine Planktonic Copepods. Available at http://copepodes.obs-banyuls.fr/en 
[Accessed May 19, 2013]

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