Species Card of Copepod
Calanoida ( Order )
    Arietelloidea ( Superfamily )
        Heterorhabdidae ( Family )
            Paraheterorhabdus ( Genus )
                Antirhabdus ( Sub-Genus )
Paraheterorhabdus (Antirhabdus) compactus  (Sars, 1900)   (F,M)
Syn.: Heterochaeta compacta Sars, 1900 (p.83, Descr.F, figs.F);
? Heterorhabdus brevicornis : Farran, 1926 (p.284, Rem.F);
Heterorhabdus compactus Sars, 1925 (p.226, figs.F,M); Farran, ? 1926 (p.284); 1929 (p.209, 267, Rem.); Rose, 1933 a (p.202, figs.F,M); Jespersen, 1934 (p.107); Lysholm & al., 1945 (p.35); Farran, 1948 e (n°16, p.3, figs.F,M); Brodsky, 1950 (1967) (p.348, figs.F,M); Grice, 1963 a (p.496); M.W. Johnson, 1963 (p.89, Table 1, 2); Tanaka, 1964 a (p.21, figs.F,M); Vidal, 1971 a (p.11, 22, 115, figs.F, M); Heptner, 1971 (p.154, figs.F,M); Harding, 1974 (p.141, tab.2, gut contents); Deevey & Brooks, 1977 (p.256, tab.2, Station "S"); Björnberg & al., 1981 (p.650, figs.F,M); Roe, 1984 (p.358); Wiebe & al., 1988 (tab.7); Hattori, 1991 (tab.1, Appendix); Razouls, 1994 (p.161, figs.F,M); Shih & Young, 1995 (p.70); Nishida & Ohtsuka, 1996 (p.620); Kosobokova & al., 1998 (tab.2); Bradford-Grieve & al., 1999 (p.883, 944, figs.F,M); Kosobokova & Hirche, 2000 (p.2029, tab.2); Yamaguchi & al., 2002 (p.1007, tab.1); Auel & Hagen, 2002 (p.1013, tab.2); Hopcroft & al., 2005 (p.198, table 2); Vives & Shmeleva, 2007 (p.301, figs.F,M, Rem.); Blachowiak-Samolyk & al., 2007 (p.2716, Table 2); Wishner & al., 2008 (p.163, Table 2, fig.8, oxycline); Gaard & al., 2008 (p.59, Table 1, N Mid-Atlantic Ridge); Galbraith, 2009 (pers. comm.); Homma & Yamaguchi, 2010 (p.965, Table 2)
Heterorhabdus (Paraheterorhabdus) compactus : Chihara & Murano, 1997 (p.819, Pl.122: F,M);
Ref. compl.: Damas & Koefoed, 1907 (p.400, tab.II); Wilson, 1942 a (p.190); Sewell, 1948 (p.349, 503, 512, 514, 567, 569); Fagetti, 1962 (p.30); De Decker & Mombeck, 1964 (p.12); Grice & Hulsemann, 1965 (p.224); Harding, 1966 (p.17, 66, 71); Grice & Hulsemann, 1967 (p.18); Dunbar & Harding, 1968 (p.319); Vinogradov, 1968 (1970) (p.201); Park, 1970 (p.477); Deevey, 1971 (p.224); Roe, 1972 (p.277, tabl.1, tabl.2); Björnberg, 1973 (p.345, 387); Vives & al., 1975 (p.50, tab.II, XII); Carter, 1977 (1978) (p.36); Vives, 1982 (p.293); Kosobokova, 1989 (p.26); Heinrich, 1990 (p.18); Mumm, 1993 (tab.1, fig.2); Richter, 1994 (tab.4.1a); Hanssen, 1997 (tab.3.1); Lapernat, 2000 (tabl.3, 4); Razouls & al., 2000 (p.343, Appendix); Kosobokova & Hirche, 2000 (p.2029, tab.2); Hop & al., 2006 (p.182, Table 4); Bucklin & al., 2010 (p.40, Table 1, Biol mol.);
Ref.:
Park, 2000 (p.85, figs.F,M, Rem.); Park & Ferrari, 2009 (p.143, Appendix 1); Blanco-Bercial & al., 2011 (p.103, Table 1, Biol. mol, phylogeny)
Species Paraheterorhabdus (Antirhabdus) compactus - Plate 1 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 2000, 31. [p.208, Fig.56].
Female: a, habitus (left side); b, c, urosme of a small specimen ( Prosome length = 1,96), left, dorsal, respectively; d, forehead (dorsal); e, f, genital somite (ventral, left, respectively); g, h, urosome of a large specimen (PL = 2,32), left, dorsal, respectively; i, left A2 (posterior).


Species Paraheterorhabdus (Antirhabdus) compactus - Plate 2 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 2000, 31. [p.209, Fig.57].
Female: a, left Md (posterior); b, masticatory edge of right Md (posterior); c, left Mx1 (posterior); d, left Mx2 (posterior); e, right Mxp (anterior); f, P1; g, P2; h, P3; i, P4. P1-4 = swimming legs 1-4 (anterior).


Species Paraheterorhabdus (Antirhabdus) compactus - Plate 3 of morphological figuresissued from : T. Park Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 2000, 31. [p.210, Fig.58].
Female: a, P5 (anterior).

Male: b, urosome (dorsal); c, P5 (with exopods omitted), anterior; d, e, f, exopod of right P5 (anterior, posterior, tilted clockwise, respectively); g, h, exopod of left P5 (anterior, tilted counterclockwise, respectively).)


Species Paraheterorhabdus (Antirhabdus) compactus - Plate 4 of morphological figuresissued from : O. Tanaka in Publs Seto Mar. Biol. Lab., 1964, XII (1). [p.22, Fig.184]. As Heterorhabdus compactus.
Female: a, habitus (dorsal); b, last thoracic segment and urosome (lateral left side); c, Mxp; d, P5.
Nota: The urosome segments and furca are in the proportional lengths as 47:9:9:13:22 = 100. A1 extends to themiddle of the last thoracic segment.

Male: e, urosome (dorsal); f, right P5; g, left P5. Nota: The lateral margin of the 2nd urosomal segment is furnished with two groups of short hairs on the right side.


Species Paraheterorhabdus (Antirhabdus) compactus - Plate 5 of morphological figuresIssued from : G.O. Sars in Résult. Camp. Scient. Prince Albert I, 69, pls.1-127 (1924). [Pl.LXII, figs.1-8]. As Heterorhabdus compactus.
Female: 1, habitus (dorsal); 2, idem (lateral left side); 3, right Md (biting edge); 4, left Md (biting edge); 5, P5; 6, anal segment and caudal rami.

Male: left A1; 8, P5.


Species Paraheterorhabdus (Antirhabdus) compactus - Plate 6 of morphological figuresIssued from : K.A. Brodskii in Calanoida of the Far Eastern Seas and Polar Basin of the USSR. Opred. Fauna SSSR, 1950, 35 (Israel Program for Scientific Translations, Jerusalem, 1967) [p.348, Fig.244]. As Heterorhabdus compactus.
Female (from central Arctic): habitus (dorsal and lateral right side); R, rostrum; MLe, Md (biting edge, left side view); S1, P1; S5, P5.

Male (from NW pacif.): S5, P5 (Le = left leg).


Species Paraheterorhabdus (Antirhabdus) compactus - Plate 7 of morphological figuresissued from : M.V. Heptner in Trudy. Inst. Okeanol., 1971, 92. [p.155, Fig.38]. As Heterorhabdus compactus.
Female & Male (from Kuril-kamchatka Trench).

Compl. Ref.:
Kosobokova & Hopcroft, 2010 (p.96, Table 1, fig.7); Kosobokova & al., 2011 (p.29, Table 2, figs.4, 8, Rem.: Arctic Basins)
NZ: 17

Distribution map of Paraheterorhabdus (Antirhabdus) compactus by geographical zones
Loc:
Antarct. (Atlant. SW, Pacif. SW), sub-Antarct. (Atlant. SW), Afr. S (E), Atlant. S, off Is. du Cap Vert NW, Maroc-Mauritanie, Canaries, off Madère, off Portugal, Baie Ibéro-marocaine, off Cap Finisterre W, Açores, Atlant. SW, Brésil, Mer des Antilles, G. du Mexique, Mer des Sargasses, off Bermuda: Station ‘’ S’’ (32°10’N, 64°30’W), off E Cape Cod, Détr. de Davis S, Fram Strait, Greenland Sea, S Iceland, Arct. (Mer de Barents, Mer de Laptev, Nansen Basin, Amundsen Basin, Makarov Basin, , Fletcher's Ice Is., Canadian abyssal plain, Lomonosov Ridge, Mer de Chukchi, Canada Basin), ? G. de Gascogne, Gibraltar W, Natal, Madagascar (Nosy Bé), Mer Arabe, Indien S, Indo-Malaisie, mers de Chine (South China Sea), Japon, Nansei Is., Kouriles, Bering Sea (Aleutian Basin), Station Knot, off British Columbia, off Californie N, Guaymas Basin, Pacif. SE (tropic.), off I. de Pâques SE, Chili N
N: 58
Lg.:
(1) F: 2,3; (22) F: 3,4-3; M: 3,4-2,9; (23) F: 3,5-2,9; M: 3,6-2,95; (35) F: 3,2; M: 2,88; (121) M: 2,59; (134) F: 3,35; (199) F: 2,58-2,28; M: 2,13-1,98; (377) F: 3,29; M: 2,98; (808) F: 2,9-2,36; (824) F: 3,32-2,24; M: 3,2-2,08; {F: 2,24-3,50; M: 1,98-3,60}
Rem.: Bathypelagic-Abyssalc (500-8500 m).
Sampling depth (Antarct., sub-Antarct.) : 250-1000 m.
(meso-bathypelagic in Roe, 1972; 1984). Overall Depth Range in Sargasso Sea: 500-2000 m (Deevey & Brooks, 1977, Station "S"); 2000-1000 m (Harding, 1974).
According to Park (2000, p.88), this species could constitute varieties or sub-species.
Last update : 26/03/2013
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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2013. - Diversity and Geographic Distribution of Marine Planktonic Copepods. Available at http://copepodes.obs-banyuls.fr/en 
[Accessed June 18, 2013]

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