Calanoida ( Ordre )
    Epacteriscoidea ( Superfamille )
Epacteriscidae Fosshagen,1973; emend.: Fosshagen, Boxshall & Iliffe, 2001 ( Epacteriscoidea )
Ref.: Fosshagen, 1973 (p.146); Razouls, 1982 (p.595); Bowman & Abele, 1982 (p.9); Barr, 1984 (p.160, Rev., emend.); Fosshagen & Iliffe, 1985 (p.346, Rev.); Park, 1986 (p.192, Rem.); Huys & Boxshall, 1991 (p.406, 419); Razouls, 1993 (p.306); Bradford-Grieve, 1999 b (p.25, Def.); Jaume & Humphreys, 2001 (p.157, Genera key, Rem.); Ohtsuka & Huys, 2001 (p.461); Fosshagen & al., 2001 (p.246, Rem., Genera Key, Phylogeny); Boxshall & Halsey, 2004 (p.14; 49; 111: Def.; p. 112: key of genera); Vives & Shmeleva, 2007 (p.992); Fosshagen & Iliffe, 2007 (p.73, Rem.: p.89, Genera Key: p.91); Blanco-Bercial & al., 2011 (p.103, Fig.3, molecular biology, phylogeny); Bradford-Grieve & al., 2014 (p.507, 528: Rem. Table 1, fig.11, cladistic analysis)
Bradford-Grieve J.M., (2002 onwards). Key to calanoid copepod families. Version 1 : 2 oct 2002. http://www.crustacea.net/crustace/calanoida/index.htm
Rem.: Pour Fosshagen (1973), la position de cette famille est malaisée à situer. L'auteur l'inclut initialement dans la superfamille des Centropagina (Gurney,1931) mais souligne certains caractères qui les rapprochent des Arietellidae, des Ridgewayiidae, voire des Augaptilidae. Il apparait difficile de faire le partage entre convergences anatomiques dues au mode de vie et relations phylogénétiques.
Formes hyperbenthiques et de grottes sous-marines.
Genre type: Epacteriscus Fosshagen, 1973.
19 G.: Azygonectes, Balinella, Bofuriella, Bomburiella, Bunderia, Caiconectes, Cryptonectes, Edaxiella, Enantiosis, Enantronia, Enantronoides, Epacteriscus, Erebonectes, Erebonectoides, Gloinella, Iboyella, Miheptneria, Minnonectes, Oinella.
D'après Bradford-Grieve & al. (2014, p.528) les genres inclus dans cette famille sont: Balinella, Bofuriella, Bomburiella, Bunderia, Cryptonectes, Edaxiella, Enantiosis, Enantronia, Enantronoides, Epacteriscus, Erebonectes, Gloinella, Iboyella, Mikheptneria, Minnonectes, Oinella. Total: 15 genres.
Famille Epacteriscidae - Planche 1Issued from : J.M. Bradford-Grieve, G.A. Boxshall & L. Blanco-Bercial in Zool. J. Linn. Soc., 2014, 171. [p.525, Fig.11].
Strict consensus of one tree after one round of successive weighting. Clade numbers above the line. The out-group is Disseta palumbii Giesbrecht, 1889.

Famille Epacteriscidae - Planche 2Issued from : J.M. Bradford-Grieve, G.A. Boxshall & L. Blanco-Bercial in Zool. J. Linn. Soc., 2014, 171. [p.528, Table 7].
Spine and seta formulae of swimming legs in Epacteriscidae.
Roman numerals = robust setae; Arabic numerals = setae. Outer border setation listed in first in each segmental group, and separated by ;

Famille Epacteriscidae - Planche 3Issued from : G.A. Boxshall & S.H. Halsey in An Introduction to Copepod Diversity. The Ray Society, 2004, No 166, Part. I. [p.111].
Armature formula of swimming legs P1 to P4 and P5 female (f).

Nota: Female P5 comprising 3-segmented protopod retaining hoop-like vestige of praecoxa laterally, and 3-segmented rami; setation sometimes reduced.
- Male P5 asymmetrical; both legs typically with 3-segmented rami; exopod 2-segmented due to fusion of 2nd and 3rd segments fused on left side in Epacteriscus. Endopods similar to those of female with armature 0-1; 0-1; 2,2,2; slightly transformed in some genera. Exopods usually highly transformed; most complex structures usually present on left exopod; modified with a slender spiniform distal segment, long and curved on right side in Erebonectes and Enanttiosis.

Famille Epacteriscidae - Planche 4Issued from : G.A. Boxshall & S.H. Halsey in An Introduction to Copepod Diversity. The Ray Society, 2004, No 166, Part. I. [p.113, Fig.21].
Epacteriscidae. A, Erebonectes nesioticus habitus male; B, A1; C, A2; D, Epacteriscus rapax habitus female; E, P5. [Fosshagen & Iliffe, 1985: A; Huys & Boxshall, 1991: B-C, E; Fosshagen, 1973: D]

Famille Epacteriscidae - Planche 5Issued from : J.M. Bradford-Grieve in NIWA Biodiv. Memoir 111, 1999 [p.26]. Armature formula of the swimming legs P1 to P5.
(1) Azygonectes Fosshagen & Iliffe, 2004
Ref.: Fosshagen & Iliffe, 2004 (p.119); 2007 (p.80: emendation, 91); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 2 spp.
Bradford-Grieve & al. (2014, p.530) considèrent ce genre , après analyse phylogénétique, comme de placement incertain dans cette famille.
(2) Balinella Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen & al., 2001 (p.292, 316); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.91); Bradford-Grieve & al., 2014 (p.507, 529: Rem., Table 1, fig. 9, 10, 11)
Rem.: Total: 1 sp.
(3) Bofuriella Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen & al., 2001 (p.286, 316); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.85: emendation, p.92)
Rem.: 3 spp.:
(4) Bomburiella Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen & al., 2001 (p.282, 316); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.92); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 1 sp.
(5) Bunderia Jaume & Humphreys, 2001
Ref.: Jaume & Humphreys (p.158, 168); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.92); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Toyal: 1 sp. :
(6) Caiconectes Fosshagen & Iliffe, 2007
Ref.: Fosshagen & Iliffe, 2007 (p.74, 91); Bradford-Grieve & al., 2014 (p.507, 530, Table 1, fig. 9, 10, 11).
Rem.: Total: 1 sp.
Bradford-Grieve & al. (2014, p.530) considèrent ce genre , après analyse phylogénétique, comme de place incertaine dans cette famille.
(7) Cryptonectes Fosshagen & Iliffe, 2004
Ref.: Fosshagen & Iliffe, 2004 (p.122); 2007 (p.91); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 1 sp.:
(8) Edaxiella Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen & al., 2001 (p.247, 316); Fosshagen & Iliffe, 2007 (p.92); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 1 sp.:
(9) Enantiosis Barr,1984; emend. Fosshagen, Boxshall & Iliffe, 2001
Ref.: Barr, 1984 (p.161); Fosshagen & Iliffe, 1985 (p.346); Huys & Boxshall, 1991 (p.62, 63); Razouls, 1993 (p.306); Mauchline, 1998 (p.70: F; p.73: M; comme Enantiosus); Bradford-Grieve, 1999 b (p.26, Def.); Fosshagen & al., 2001 (p.258, 316); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.92); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 7 spp.:
Remarques sur les dimensions et le sex-ratio:
The mean female size is 1.328 mm (n = 11, SD = 0.115), and the mean male size is 1.235 mm (n = 13, SD = 0.085). The size ratio male: female is 0.923.
(10) Enantronia Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen & al., 2001 (p.251, 316); Vives & Shmeleva, 2007 (p.993); Fosshagen & Iliffe, 2007 (p.91); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 1 sp.:
Genre Enantronia - Planche 1Issued from : F. Vives & A.A. Shmeleva in Fauna Iberica, 26, 2007. [p. 994, Fig. 573, A-D]. Enantronia canariensis Female.
A, habitus (dorsal view); B, urosome (lateral view); C, Mxp; D, P1.
(11) Enantronoides Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen & al., 2001 (p.255, 316); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.91); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 1 sp.:
(12) Epacteriscus Fosshagen,1973; emend. Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen, 1973 (p.146); Huys & Boxshall, 1991 (p.53); Razouls, 1993 (p.306); Mauchline, 1998 (p.68); Bradford-Grieve, 1999 b (p.26, Déf.); Fosshagen & al., 2001 (p.276, 316, Rem.); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.91); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: type: Epacteriscus rapax. Total: 3 spp.
Genre Epacteriscus - Planche 1Issued from : G.A. Boxshall & S.H. Halsey in The Ray Society, 2004. [p.113, Fig. 21, D, E]. Epacteriscus rapax Female.
D, habitus (lateral view); E, P5.
(13) Erebonectes Fosshagen,1985; emend. Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen & Iliffe, 1985 (p.346); Huys & Boxshall, 1991(p.50, 51); Razouls, 1993 (p.306); Mauchline, 1998 (p.67); Bradford-Grieve, 1999 b (p.26, Déf.); Fosshagen & al., 2001 (p.291, 316, Rem.); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.91); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 1 sp.:
Genre Erebonectes - Planche 1Issued from : G. A. Boxshall & S.H. Halsey in The Ray Society, 2004. [p.179, Fig. 21, A-C]. Erebonectes nesioticus Male.
A, habitus (dorsal view); B, A1; C, A2.
(14) Erebonectoides Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen & al., 2001 (p.291, 316); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.91); Bradford-Grieve, 2014 (p.507, 530, fig.9, 10, 11)
Rem.: 1 sp.
Bradford-Grieve & al. (2014, p.530) considèrent ce genre, après analyse phylogénétique, comme de place incertaine dans cette famille.
(15) Gloinella Fosshagen, 2001
Ref.: Fosshagen & al., 2001 (p.301, 316); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2007 (p.92); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 1 sp.:
(16) Iboyella Boxshall & Jaume, 2003
Ref.: Boxshall & Jaume, 2003 (p.86); Fosshagen & Iliffe, 2007 (p.92); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 1 sp.
(17) Miheptneria Andronov, 2007
Ref.: Andronov, 2007 (p.672); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Type: Miheptneria abyssalis. Total: 1sp.
(18) Minnonectes Fosshagen & Iliffe, 2004
Ref.: Fosshagen & Iliffe, 2004 (p.128: male diagnosis); 2007 (p.88: female diagnosis, p.92); Bradford-Grieve & al., 2014 (p.507, 529: Rem., Table 1, Fig. 9, 10, 11)
Rem.: Total: 1 sp.:
(19) Oinella Fosshagen, Boxshall & Iliffe, 2001
Ref.: Fosshagen & al., 2001 (p.297, 316); Bradford-Grieve, 2004 (p.287); Fosshagen & Iliffe, 2004 (p.132); 2007 (p.91); Bradford-Grieve & al., 2014 (p.507, 522: Rem., Table 1, fig. 9, 10, 11, cladistic analysis)
Rem.: Total: 1 sp.:

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Razouls C., Desreumaux N., Kouwenberg J. et de Bovée F., 2005-2024. - Biodiversité des Copépodes planctoniques marins (morphologie, répartition géographique et données biologiques). Sorbonne Université, CNRS. Disponible sur http://copepodes.obs-banyuls.fr [Accédé le 20 avril 2024]

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