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Calanoida ( Ordre ) |
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Arietelloidea ( Superfamille ) |
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| Arietellidae Sars, 1902 ( Arietelloidea ) | | Ref.: | Sars, 1902 (1903) (p.123); Gurney, 1931 (p.85); Wilson, 1932 a (p.136); Rose, 1933 a (p.238); Brodsky, 1950 (1967) (p.393); Andronov, 1974 a (p.1005); Campaner, 1977 (p.811, Redef.); Björnberg & al., 1981 (p.650); Bowman & Abele, 1982 (p.9); Razouls, 1982 (p.526); Brodsky & al., 1983 (p.143, 145); Campaner, 1984 (p.102, 107, Rem.); Mauchline, 1988 (p.705, 706, 740: cuticular pores); Ohtsuka & Mitsuzumi, 1990 (p.96); Huys & Boxshall, 1991 (p.460); Razouls, 1993 (p.306); Ohtsuka & al., 1994 (p.105, Redef., Genera Key, cladistic analysis); Chihara & Murano, 1997 (p.719); Cuoc & al., 1997 (p.651, phylogeny); Bradford-Grieve & al., 1999 (p.881, 902, 903, 936, 937: Genera Key); Bradford-Grieve, 1999 b (p.27, Def.); Ohtsuka & Huys, 2001 (p.445, 461); Boxshall & Halsey, 2004 (p.13; 49; 62: Def.; p.64: Genera Key); Vives & Shmeleva, 2007 (p.146, part. Genera Key); Blanco-Bercial & al., 2011 (p.103, Table 1, Fig.2, 3, 4, Biol. mol, phylogeny)
Bradford-Grieve J.M., (2002 onwards). Key to calanoid copepod families. Version 1 : 2 oct 2002. http://www.crustacea.net/crustace/calanoida/index.htm  | | Rem.: | Ohtsuka, Boxshall & Roe (1994) redéfinissent la famille et établissent les relations phylétiques entre les genres (dont 3 nouveaux). Soit 12 genres: Arietellus, Campaneria, Crassarietellus, Metacalanalis, Metacalanus, Paramisophria, Paraugaptiloides, Paraugaptilus, Pilarella, Protoparamisophria, Sarsarietellus, Scutogerulus. Griceus est considéré comme incertae sedis et Rhapidophorus comme inquirendum. Ces auteurs ne retiennent pas les sous-familles mais montrent l'existence de 2 lignées. |  issued from : S. Ohtsuka, S. Nishida & R.J. Machida in J. Nat. Hist., 2005, 39 (27). [p.2510, Fig.14]. Cladogram of Arietellid genera. Main habitat and depth range provided only for hyperbenthic genera, exvluding two truly pelagic genera, Arietellus and Paraugaptilus. The phylogenetic relationship among arietellid genera two clades are recognized: the Metacalanus-group and the -group. |
 issued from : S. Ohtsuka, S. Nishida & R.J. Machida in J. Nat. Hist., 2005, 39 (27). [p.2512, Fig.15]. Cladogram depicting main habitats and colonization routes for arietellid genera and their presumed ancestrors. D: deep-water hyperbenthic: S: shallow-water hyperbenthic (including cavernicolous and epipelagic); P: deep-water planktonic. In the Metacalanus-Metacalanis-Pilarella-clade there are two possibilities for colonization routes (right: habitat transition from deep-sea to shallow-water hyperbenthos; left: habitat shift from deep-sea to shallow-water hyperbenthos occurred independently in Metacalanus and Pilarella. (o) Habitat shift from deep-sae hyperbenthic to shallow-sea hyperbenthos; (black o) habitat shift from shallow-water hyperbenthos to deep-sea hyperbenthos. |
 issued from : S. Ohtsuka, G.A. Boxshall & H.S.J. Roe in Bull. nat. Hist. Mus. Lond. (Zool.), 1994, 60 (2). [p.161, Fig.38]. Evolutionary trends in the structures of the female genital systems of the arietellid genera. A: Fusion of copulatory pores to form single pore, and anterolateral migration of both gonopores. B: Posterior migration of both gonopores and copulatory pores, and separation of copulatory pore from gonopore. C: Anterolateral migration of gonopores, and separation of copulatory pore from gonopore and their asymmetrical arrangement and enlargement. D: Lateral migration of both gonopores and copulatory pores, and separation of copulatory pore from gonopore. E: Lateral migration of both gonopores and copulatory pores. Pg: Paramisophria giselae; Pj: Paramisophria japonica; Pc: Paramisophria cluthae; Sa: Sarsarietellus abyssalis; Ap: Arietellus plumifer; M1: Metacalanus sp.1; M2: Metacalanus sp.2; Ps: Paraugaptilus similis; Ch: Crassarietellus huysi; Sp: Scutogerulus pelophilus. g : gonopore; c: copulatory pore. |
 issued from : S. Ohtsuka, G.A. Boxshall & H.S.J. Roe in Bull. nat. Hist. Mus. Lond. (Zool.), 1994, 60 (2). [p.163, Fig.39]. Schematic illustration of segmental fusion patterns and armature of female antennules of the arietellid genera. A: Crassarietellus; B: Sarsarietellus; C: Scutogerulus; D: Paramisophria; E: Arietellus; F: Paraugaptilus; G: Metacalanus; H: Pilarella. The number indicates the number of setae on each segment. Asterisk indicates segment without aesthetasc. |
 issued from : S. Ohtsuka, G.A. Boxshall & H.S.J. Roe in Bull. nat. Hist. Mus. Lond. (Zool.), 1994, 60 (2). [p.164, Fig.40]. Schematic illustration of segmental fusion patterns and armature of male antennules of the arietellid genera. A: Campaneria; B: Crassarietellus; C: Paramisophria; D: Arietellus; E: Paraugaptiloides; F: Paraugaptilus; G: Metacalanus. The number indicates the number of setae on each segment. Asterisk indicates a segment with no aesthetasc. |
 issued from :C. Cuoc, D. Defaye, M. Brunet, R. Notonier & J. Mazza in Mar. Biol., 1997, 129. [p.13, Fig.6]. Diagrammatic representation of main evolutionary trends in organization of female genital area in calanoïds for Arietellidae (modified after Huys & Boxshall, 1991 and Ohtsuka & al., 1994). | | | | | Arietellinae Brodsky, 1950 | | Ref.: | Brodsky, 1950 (1967) (p.394); Brodsky & al., 1983 (p.143) | | Rem.: | Brodsky crée cette sous-famille pour y inclure Arietellus, Paraugaptilus, Paramisophria et Scottula et considère Phyllopus comme le type d'une autre sous-famille (Phyllopinae ). Campaner (1977, p.811) considère celle-ci comme une famille : Phyllopidae (emend. Bowman & Abele, 1982, p.9 = Phyllopodidae = Nullosetigeridae ). Dans ces conditions la sous-famille établie par Brodsky n'a plus de raison d'être. | | | | (1) Arietellus Giesbrecht, 1892 | |
| | Ref.: | &Giesbrecht, 1892 (p.66, 415); Giesbrecht & Schmeil, 1898 (p.124); Esterly, 1905 (p.189); Farran, 1908 b (p.81); Van Breemen, 1908 a (p.139, 238, 239); A. Scott, 1909 (p.142); Wolfenden, 1911 (p.329); Sewell, 1932 (p.329); Rose, 1933 a (p.238, clé spp.); Tanaka, 1964 c (p.231); Owre & Foyo, 1967 (p.90, clé spp.); Brodsky, 1950 (1967) (p.394, clé spp.); Razouls, 1982 (p.526); Gardner & Szabo, 1982 (p.395, Rem.); Mauchline, 1988 (p.705, 740); Razouls,1993 (p.306); Ohtsuka & al., 1994 (p.126, Redef.); Chihara Murano, 1997 (p.720); Mauchline, 1998 (p.71: F; p.74: M); Bradford-Grieve & al., 1999 (p.937: spp. key); Bradford-Grieve,1999 b (p.28, Déf.); Boxshall & Halsey, 2004 (p.65); Vives & Shmeleva, 2007 (p.147, spp. key) | | Rem.: | type: Arietellus setosus . 14 spp. + 1 indet.: | | Remarques sur les dimensions et le sex-ratio: | | La moyenne des dimensions des femelles est de 4,983 mm (n= 12; S= 0,668; Cv= 0,134) et de 4,872 mm pour les mâles (n= 6; S= 0,54; Cv= 0,111). Le rapport des longueurs (M/F) est de 0,967 ou 96,7 % (n= 6; S= 0,039; Cv= 0,040) | | | | (2) Campaneria Ohtsuka, Boxshall & Roe, 1994 | |
| | Ref.: | Ohtsuka & al., 1994 (p.119); Mauchline, 1998 (p.74: M); Bradford-Grieve,1999 b (p.34, Déf., Rem.); Bradford-Grieve, 2004 (p.287); Boxshall & Halsey, 2004 (p.64) | | | | (3) Crassarietellus Ohtsuka, Boxshall & Roe, 1994 | |
| | Ref.: | Ohtsuka & al., 1994 (p.109); Mauchline, 1998 (p.71: F); Bradford-Grieve,1999 b (p.34, Déf.); Bradford-Grieve, 2004 (p.287); Boxshall & Halsey, 2004 (p.64) | | Rem.: | type: Crassarietellus huysi. 1 sp. + 1 indet. | | | | (0) Griceus Ferrari & Markhaseva, 2000 | |
| | Ref.: | Ferrari & Markhaseva, 2000 (p.77); Bradford-Grieve, 2004 (p.287) | | Rem.: | Boxshall & Halsey, 2004 (p.64, Rem.: Genus incertae sedis).1 sp. | | | | (4) Metacalanalis Ohtsuka, Nishida & Machida, 2005 | |
| | Ref.: | Ohtsuka & al., 2005 (p.2484 | | Rem.: | Type: Metacalanalis hakuhoae. 1 sp. | | | | (5) Metacalanus Cleve, 1901 | |
| | Syn.: | Scottula (part.) | | Ref.: | Cleve, 1901 (p.43); A. Scott, 1909 (p.146); Sewell, 1932 (p.329); Tanaka, 1964 c (p.236); Razouls, 1982 (p.531); Alvarez, 1985 (p.190, 194, Rem.); Ohtsuka, 1985 (p.305); Mauchline, 1988 (p.706); Razouls, 1993 (p.306); Ohtsuka & al., 1994 (p.138, Redef.); Chihara & Murano, 1997 (p.719); Mauchline, 1998 (p.73: F; p.74: M); Bradford-Grieve,1999 b (p.35, Déf.); Bradford-Grieve, 2004 (p.287); Boxshall & Halsey, 2004 (p.64); Vives & Shmeleva, 2007 (p.155) | | Rem.: | type: Metacalanus aurivilli . 4 spp.+ 2 indet: | | Remarques sur les dimensions et le sex-ratio: | | La moyenne des longueurs des femelles est de 0,888 mm (n= 6; S= 0,227; Cv= 0,26) et de 0,738 mm pour les mâles (n= 5; S= 0,172; Cv= 0,23). Le rapport des longueurs (M/F) est de 0,836 ou 83,6 % (n= 5; S= 0,069; Cv= 0,083) | | | | (6) Paramisophria T. Scott, 1897 | |
| | Syn.: | Parapseudocyclops Campaner,1977 (p.812) | | Ref.: | T. Scott, 1897 f (p.147); Giesbrecht & Schmeil,1898 (p.123); Sars, 1902 (1903) (p.127); van Breemen, 1908 a (p.140); Rose, 1933 a (p.242); Brodsky, 1950 (1967) (p.394); Krishnaswamy, 1959 (p.546, Rem.: emend.); Tanaka, 1964 c (p.242); Razouls, 1982 (p.530); McKinnon & Kimmerer, 1985 (p.85, clé spp.); Ohtsuka, 1985 (p.296, 298, Rem.); Ohtsuka & Mitsuzumi, 1990 (p.97); Huys & Boxshall, 1991 (p.49, 325, 436); Ohtsuka & al.,1991 (p.793, 800, Rev.); Ohtsuka & al., 1993 (p.57, 65, Rem.: emend.); Ohtsuka & al., 1994 (p.158, figs., Rem.); Chihara & Murano, 1997 (p.719); Mauchline, 1998 (p.68); Bradford-Grieve,1999 b (p.36, Déf.); Bradford-Grieve, 2004 (p.287); Boxshall & Halsey, 2004 (p.65); Vives & Shmeleva, 2007 (p.156, spp. key) | | Rem.: | type: Paramisophria cluthae . Ohtsuka & al, 1991, p.800, signalent 11 espèces: 7 Indo-Pacifique et 4 Atlantique, certaines non encore décrites à ce jour). 16 spp. + 2 indet. : | | Remarques sur les dimensions et le sex-ratio: | | La moyenne des dimensions des femelles est de 1,509 mm (n= 10; S= 0,524; Cv= 0,347) et de 1,249 mm pour les mâles (n= 10; S= 0,293; Cv= 0,235). Le rapport des longueurs (M/F) est de 0,889 ou 88,9 % (n= 9; S= 0,075; Cv= 0,084). |  issued from : M. Carola, C. Razouls & J.L. Pretus in Vie Milieu, 1995, 45 (2). [p.150, Table III B]. Distribution géographique des espèces du genre Paramisophria connues en 1995. |
 issued from : M. Carola, C. Razouls & J.L. Pretus in Vie Milieu, 1995, 45 (2). [p.150, Table III A]. Liste des rapports chez les adultes femelles du gentre connues en 1995.. Pr = prosome (mesuré du bord antérieur de la tête au bord postérieur du dernier segment thoracique; dans l'axe sagittal). Ur = urosome (mesuré du bord antérieur du segment génital à l'extrémité distale des rames caudales). L = longueur maximale; w = largeur maximale. |
 issued from : M. Carola, C. Razouls & J.L. Pretus in Vie Milieu, 1995, 45 (2). [p.150, Table IV]. Characteristics of some appendages of the Paramisophria |
 issued from : D. Jaume, J.E. Cartes & G.A. Boxshall in Contr. Zool., 2000, 68 (4). [p.220, Table III]. Caractères permettant de différencier les 13 espèces de Paramisophria. Tous les caractères se réfèrent aux adultes. A1: antennule; art: articulation; end-: endopod segment; exp-: exopod segment; seg.: segment. |
 issued from : D. Jaume, J.E. Cartes & G.A. Boxshall in Contr. Zool., 2000, 68 (4). [p.24, A]. Cladogramme (l'un des plus parcimonieux) résultant d'une analyse cladistique pour les espècre de Paramisophria. A, arbre pour la valeur f = 516. Les branches ombrées se réfèrent à l'habitat ancestral. | | | | | Parapseudocyclops Campaner, 1977 | |
| | Ref.: | Campaner, 1977 (p.812); Razouls, 1982 (p.531); Campaner, 1984 (p.107, Rem.); Ohtsuka & al., 1991 (p.793, 800, Rem.) | | Rem.: | Cf. Paramisophria | | | | (7) Paraugaptiloides Ohtsuka, Boxshall & Roe, 1994 | |
| | Ref.: | Ohtsuka & al., 1994 (p.120); Mauchline, 1998 (p.74: M); Bradford-Grieve,1999 b (p.37, Déf.); Bradford-Grieve, 2004 (p.287); Boxshall & Halsey, 2004 (p.65); Ohtsuka & al., 2005 (p.2497: emend part.) | | Rem.: | Type: Paraugaptiloides magnus (Bradford,1974). 3 spp.: | | | | (8) Paraugaptilus Wolfenden, 1904 | |
| | Ref.: | Wolfenden, 1904 (p.123); Van Breemen, 1908 a (p.138); A. Scott, 1909 (p.144); Wolfenden, 1911 (p.353); Sars, 1925 (p.335); Sewell, 1932 (p.329); Rose, 1933 a (p.241); Tanaka, 1964 c (p.234); Brodsky, 1950 (1967) (p.397); Deevey, 1973 (p.247, Rev, clé spp.); Razouls, 1982 (p.529); Campaner, 1984 (p.107); Razouls, 1993 (p.306); Ohtsuka & al., 1994 (p.143, 158, 160, Redef.); Chihara & Murano, 1997 (p.721); Mauchline, 1998 (p.71: F; p.74: M); Bradford-Grieve,1999 b (p.38, Déf.); Boxshall & Halsey, 2004 (p.65); Vives & Shmeleva, 2007 (p.166) | | Rem.: | type: Paraugaptilus buchani . 6 spp.: | | Remarques sur les dimensions et le sex-ratio: | | La moyenne des longueurs des femelles est de 3,220 mm (n= 5; S= 0,215; Cv= 0,067) et de 3,310 mm pour les mâles (n= 5; S= 0,796; Cv= 0,24). Le rapport des longueurs (M/F) est de 0,912 ou 91,2 % (n= 4; S= 0,012; Cv= 0,013) | | | | (9) Pilarella Alvarez, 1985 | |
| | Ref.: | Alvarez, 1985 (p.189); Razouls, 1993 (p.306); Ohtsuka & al., 1994 (p.156, Redef.); Mauchline, 1998 (p.73: F); Bradford-Grieve,1999 b (p.40, déf.); Bradford-Grieve, 2004 (p.287); Boxshall & Halsey, 2004 (p.64) | | Rem.: | 1 sp.: | | | | (10) Protoparamisophria Ohtsuka, Nishida & Machida, 2005 | |
| | Ref.: | Ohtsuka & al., 2005 (p.2491) | | Rem.: | Type: Protoparamisophria biforaminis. 1 sp. | | | | (0) Rhapidophorus Edwards, 1891 | |
| | Ref.: | C.L. Edwards, 1891 (p.84); Giesbrecht, 1892 (p.423); Wilson, 1932 a (p.543, non M :'F'); Fosshagen, 1968 a (p.61, 62, Rem.); Campaner, 1977 (p.817, Rem.); Razouls, 1982 (p.531); Campaner, 1984 (p.107, Rem.); Ohtsuka & al., 1991 (p.801, Rem.); Razouls, 1993 (p.306); Ohtsuka & al., 1994 (p.131, Rem.); Mauchline, 1998 (p.68: M); | | Rem.: | Pour Fosshagen (1968 a, p.62) la description de la (F) devrait permettre de savoir si le genre Paramisophria n'est pas un synonyme de ce genre. Boxshall & Halsey, 2004 (p.64: Genus inquirendum) 1sp.: | | | | (11) Sarsarietellus Campaner, 1984 | |
| | Syn.: | Scottula (part.) Sars, 1905 c (p.22); 1925 (p.338, figs.F,M) | | Ref.: | Campaner, 1984 (p.107, Rem.); Razouls, 1993 (p.306); Ohtsuka & al., 1994 (p.155, 158, 160, Redef.); Mauchline, 1998 (p.68: F; p.76: M); Bradford-Grieve, 2004 (p.287); Boxshall & Halsey, 2004 (p.65); Ohtsuka & al., 2005 (p.2504, emend. part.) | | Rem.: | type: Scottula abyssalis Sars,1905. 3 spp.: | | | | Scottula Sars, 1902 | | Ref.: | Sars, 1902 (1903) (p.124); Van Breemen, 1908 a (p.142); A. Scott, 1909 (p.146); Rose,1933 a (p.243); Tanaka, 1964 c (p.242); Brodsky, 1950 (1967) (p.394); Razouls, 1982 (p.532); Campaner, 1984 (p.102, 107, Rem.); Ohtsuka, 1985 (p.305); Mauchline, 1988 (p.706) | | Rem.: | Cf. Metacalanus (part.) & Sarsarietellus (part.) | | | | (12) Scutogerulus Bradford, 1969 | |
| | Ref.: | Bradford, 1969 b (p.495); Razouls, 1982 (p.531); Campaner, 1984 (p.107); Razouls, 1993 (p.306); Ohtsuka & al., 1994 (p.152, 160, Redef.); Mauchline, 1998 (p.68: F); Bradford-Grieve,1999 b (p.39, Déf.); Bradford-Grieve, 2004 (p.287); Boxshall & Halsey, 2004 (p.64), Ohtsuka & Boxshall, 2004 (p.54: Rem.) | | Rem.: | 2 spp.: | | | | |
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 Toute utilisation de ce site pour une publication sera mentionnée avec la référence suivante :
Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2012. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Disponible sur http://copepodes.obs-banyuls.fr
[Accédé le 24 mai 2013] © copyright 2005-2012 CNRS, UPMC
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