Fiche d'espèce de Copépode
Calanoida ( Ordre )
    Clausocalanoidea ( Superfamille )
        Phaennidae ( Famille )
            Phaenna ( Genre )
Phaenna spinifera  Claus, 1863   (F,M)
Syn.: ? Onchocalanus latus Esterly, 1911 (p.326, figs.F); 1912 (tab.10); 1913 (p.183, figs.F); Brodsky, 1950 (1967) (p.235, figs.F);
Onchocalanus nudipes C.B. Wilson, 1942 a (p.199, Descr.F, figs.F, [fig.83: juv.M); 1950 (p.273); Fagetti, 1962 (p.23)
Ref.:
Claus, 1863 (p.189, figs.F,M); Giesbrecht, 1892 (p.293, 774, figs.F,M); Thompson, 1888 d (p.142); T. Scott, 1894 b (p.81, figs.M); Giesbrecht & Schmeil, 1898 (p.50, Rem. F,M); Thompson & Scott, 1903 (p.234, 246); Wolfenden, 1904 (p.111, 112); Farran, 1908 b (p.47) ; A. Scott, 1909 (p.80); Wolfenden, 1911 (p.285); With, 1915 (p.241, figs.F, Rem.F,M); Pesta, 1920 (p.514); Farran, 1926 (p.268); 1929 (p.209, 255); Sewell, 1929 (p.175); Rose, 1933 a (p.126, figs.F,M); Mori, 1937 (1964) (p.54, figs.F); Lysholm & al., 1945 (p.24); Vervoort, 1950 b (p.83, figs.F,M, juv.M, Rem.); Chiba, 1956 (p.32, figs.juv.F); Tanaka, 1960 a (p.85, figs. juv.F); Grice, 1962 (p.203, fig.F); Chen & Zhang, 1965 (p.57, figs.F,M); Vervoort, 1965 (p.25, Rem.); Owre & Foyo, 1967 (p.57, figs.F,M); Mazza, 1967 (p.174, figs.F,M, juv.); Park, 1968 (p.554); Vidal, 1968 (p.30, figs.F); Corral Estrada, 1970 (p.156; Bradford, 1970 a (p.355, figs.M); 1972 (p.40, fig.F); Razouls, 1972 (p.94, Annexe: p.52, figs.F,M); Bradford, 1973 (p.137, Rem.F,M, fig.2a); Marques, 1974 (p.15); 1976 (p.991); Björnberg & al., 1981 (p.635, figs.F,M); Zheng & al., 1982 (p.40, figs.F); Park, 1983 a (p.318, figs.F); Bradford & al., 1983 (p.70, figs.F,M); Roe, 1984 (p.357); Tanaka & Omori, 1992 (p.268); Mazzocchi & al., 1995 (p.188, figs.F, Rem.); Chihara & Murano, 1997 (p.854, Pl.137: F); Bradford-Grieve & al., 1999 (p.880, 928, 929, figs.F,M); Conway & al., 2003 (p.188, figs.F,M, Rem.); Boxshall & Halsey, 2004 (p.159, figs.F); Mulyadi, 2004 (p.187, figs.F, Rem.);Vives & Shmeleva, 2007 (p.687, figs.F,M, Rem.); Blanco-Bercial & al., 2011 (p.1, Table 103, molecular biology, phylogeny); Soh & al; (p.74, figs.F)
Espèce Phaenna spinifera - Planche 1 de figures morphologiquesissued from : J.M. Bradford, L. Haakonssen & J.B. Jillett in Mem. N.Z. Oceanogr. Inst., 1983, 90. [p.62, Fig.34].
Female: A, habitus (dorsal); B, idem (lateral right side); C, A1; D, A2; E, Md (mandibular palp); F, Md (cutting blade).


Espèce Phaenna spinifera - Planche 2 de figures morphologiquesissued from : J.M. Bradford, L. Haakonssen & J.B. Jillett in Mem. N.Z. Oceanogr. Inst., 1983, 90. [p.63, Fig.34 (legend p.62)].
Female: G, Mx1; H, Mx2; I, Mxp; J, P1; K, P2; L, P3; M, P4.


Espèce Phaenna spinifera - Planche 3 de figures morphologiquesissued from : J.M. Bradford, L. Haakonssen & J.B. Jillett in Mem. N.Z. Oceanogr. Inst., 1983, 90. [p.64, Fig.35 (legend p.65)].
Male: A, habitus (dorsal); B, idem (lateral left side); C, A1; D, A2; E, Md (mandibular palp); F, Mx1; G, Mx2.


Espèce Phaenna spinifera - Planche 4 de figures morphologiquesissued from : J.M. Bradford, L. Haakonssen & J.B. Jillett in Mem. N.Z. Oceanogr. Inst., 1983, 90. [p.65, Fig.35].
Male: H, Mxp; I, P1; J, P2; K, P3; L, P4; M, P5.


Espèce Phaenna spinifera - Planche 5 de figures morphologiquesissued from : T. Park in Antarct. Res. Ser. Washington, 1983, 39 (5). [p.320, Fig.1].
Female (from southern coast of W Australia): a, habitus (dorsal); b, idem ( lateral left side); c, urosome (lateral left side); d, rostrum (anterior view); e, A2; f, Md; g, Mx1; h, Mx2.

Nota: 1st matasomal segment fully separate from head, 4th and 5th only partially fused close to ventral margin.


Espèce Phaenna spinifera - Planche 6 de figures morphologiquesissued from : T. Park in Antarct. Res. Ser. Washington, 1983, 39 (5). [p321, Fig.2].
Female: a, Mxp; b, P1 (anterior); c, P2 (posterior); d, P3 (posterior); e, P4 (posterior).


Espèce Phaenna spinifera - Planche 7 de figures morphologiquesissued from : J.M. Bradford in N.Z. Jl Mar. Freshw. Res., 1970, 4 (4). [p.355, Figs 24-25].
Male (off Kaikoura, New Zealand): 24, P3; 25, P5.
Scale bars represent 0.1 mm.


Espèce Phaenna spinifera - Planche 8 de figures morphologiquesIssued from: M.G. Mazzocchi, G. Zagami, A. Ianora, L. Guglielmo & J. Hure in Atlas of Marine Zooplankton Straits of Magellan. Copepods. L. Guglielmo & A. Ianora (Eds.), 1995. [p.189, Fig.3.33.1].
Female: A, habitus (dorsal); B, urosome (lateral left side).
Nota: head and 1st thoracic segment separated, 4th and 5th incompletely separated. Lateral thoracic somites 3 and 4 with a rather long sensillum on either side. Ventral surface of urosomal somites ornated with stout setae (this ornementation is fragile and can easily be lost in fixed specimens).


Espèce Phaenna spinifera - Planche 9 de figures morphologiquesIssued from: M.G. Mazzocchi, G. Zagami, A. Ianora, L. Guglielmo & J. Hure in Atlas of Marine Zooplankton Straits of Magellan. Copepods. L. Guglielmo & A. Ianora (Eds.), 1995. [p.190, Fig.3.33.2].
Female (SEM preparation): A, habitus (dorsal); B, cephalothorax (ventral ); C, forehead (frontal view); D, urosome (dorsal); E, idem (ventral).
Bars: A, B, D 0.100 mm; C, E 0.050 mm.


Espèce Phaenna spinifera - Planche 10 de figures morphologiquesissued from: Q.-c Chen & S.-z. Zhang in Studia Marina Sinica, 1965, 7. [Pl.17, 9-12].
Female (from E China Sea): 9, habitus (dorsal); 10, idem (lateral left side).

Male: 11, habitus (dorsal); 12, P5 (posterior).


Espèce Phaenna spinifera - Planche 11 de figures morphologiquesissued from : C.B. Wilson in Scientific Results of Cruise VII of the Carnegie during 1928-1929. Biology-I. Carnegie Inst. Wash. Publ. 536, 1942. [p.227, Fig.74]. As Onchocalanus nudipes.
Female: 74, Mx2.


Espèce Phaenna spinifera - Planche 12 de figures morphologiquesissued from : Z. Zheng, S. Li, S.J. Li & B. Chen in Marine planktonic copepods in Chinese waters. Shanghai Sc. Techn. Press, 1982 [p.40, Fig.23].
Female: a-b, habitus (dorsal and lateral, respectively); c, P3.
Scale bars in mm.


Espèce Phaenna spinifera - Planche 13 de figures morphologiquesissued from : T. Mori in The pelagic Copepoda from the neighbouring waters of Japan, 1937 (2nd edit., 1964). [Pl.27, Figs.9-15].
Female: 9, P2 (posterior); 10, P3 (posterior); 11, P4 (posterior); 12, Mx1; 13, P1; 14, habitus (dorsal); 15, A2.


Espèce Phaenna spinifera - Planche 14 de figures morphologiquesissued from : J.M. Bradford in Mem. N. Z. Oceonogr. Inst., 1972, 54. [p.42, Fig.9, (5)].
Female (from Kaikoura, New Zealand): 5, habitus (dorsal).
Scale bar: 1mm.


Espèce Phaenna spinifera - Planche 15 de figures morphologiquesissued from : G.D. Grice in Fish. Bull. Fish and Wildl. Ser., 1962, 61. [p.204, Pl.15, Fig.1].
Female (from equatorial Pacific): 1, habitus (doprsal).
Nota: Genital segment with a large protrusion on the right side from which arises a smaller spine-like process.


Espèce Phaenna spinifera - Planche 16 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.12, Figs.2, 3, 8, 36].
Female: 2, A2 (anterior surface); 3, Palp of Md (anterior surface); 8, Mxp (anterior surface); 36, Mx1 (posterior view).


Espèce Phaenna spinifera - Planche 17 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.12, Fig.4].
Female: 4, P3 (posterior view).


Espèce Phaenna spinifera - Planche 18 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.12, Figs.5, 35].
Male: 5, P5 (posterior view); 35, Mx1 ( (anterior view).
Ps = left leg; Pd = right leg.


Espèce Phaenna spinifera - Planche 19 de figures morphologiquesissued from : C. With in The Danish Ingolf-Expedition, Copepoda I, 1915, III, 4. [p.242, Text-fig. 79].
Female (from 51°00N, 11°43'W): Lateral outline of body).

Nota: In some of spexcimens the head possesses, dorsally, a conical protuberance.
Length of prosome: 2.42 mm, urosome: 0.48 mm.
The legs are scarcely different from Giesbrecht’s description. The outer margin of the 3rd exopodal segment of P1 has proximally to the middle a small incision and a glandular pore. On the anterior surface of the 3rd endoppodal segment of P2 and P3 a minute glandular pore, but in P4 none was seen. At the base of the outer spine in the exopodal segment 2, and at the base of the outer seta 1 and 3 of exopodal segment 3, glandular pores were found in P2, and in P3 and P4 at the base of the outer spine of exopodal 1. The endopodal segments 2-3, which are fused in P2, possess on the posterior surface two coronas of 8 and 6 long ; spines respectively; the endopodal segments 2 and 3 of P3 have 8 and 6 spines respectively ; the endopodal segment 2 of P4 has 6 long rather weak spines.

Male: Length of prosome: 1.89 mm, urosome: 0.37 mm. The prosome did not possess a lamelliform projection, as figured by Giesbrecht; no dorsal protuberance.


Espèce Phaenna spinifera - Planche 20 de figures morphologiquesissued from : C. With in The Danish Ingolf-Expedition, Copepoda I, 1915, III, 4. [Pl. VII, Fig.1, a-c].
Female: a, rostrum (anterior view); b, labrum (oral view); c, lamina labialis and serrula 6-dentata.

Nota: The very short labrum and the elevation in front of it are only indistinctly separated, and the latter is placed behind the insertion of the A1; the epistoma is covered by a number of longer and shorter setae; on the labrum proper a transverse group of fairly short setae; the marginal setae are only poorly developed. The oral surface (fig.b) (fig.b) has in front 4 longer and shorter oblique groups of shorter hairs converging towards the middle, and more posteriorly, probably composed of the 3 hinder, a longitudinally placed elongated group of short hairs; in the middle at least 3 groups of very short hairs or granules. The lamina labialis is only poorly developed (fig.c); in front of it a regular granular area; the area behind the lamina and on the labial lobes not studied in details.


Espèce Phaenna spinifera - Planche 21 de figures morphologiquesissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1967, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.23, Fig.91].
Female: 91, habitus (dorsal).


Espèce Phaenna spinifera - Planche 22 de figures morphologiquesissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1967, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.50, Figs.292, 293]. After Sars, 1925 and Giesbrecht, 1892, respectively.
Male: 292, prosome (dorsal); 293, P5.


Espèce Phaenna spinifera - Planche 23 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.37, Figs.20, 21].
Female: 20, forehead (ventral); 21, same (lateral).


Espèce Phaenna spinifera - Planche 24 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.37, Figs.17, 19].
Female: 17, urosome (ventral); 19, thoracic segments 4-5 and urosomal segment 1-2 (lateral).


Espèce Phaenna spinifera - Planche 25 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.12, Fig.37].
Female: 37, Md (cutting edge of gnathobase).


Espèce Phaenna spinifera - Planche 26 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.12, Fig.6].
Male: 6, A1 (ventral view).


Espèce Phaenna spinifera - Planche 27 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.12, Fig.1].
Male: 1, Mxp (anterior view).


Espèce Phaenna spinifera - Planche 28 de figures morphologiquesissued from : C.B. Wilson in Scientific Results of Cruise VII of the Carnegie during 1928-1929. Biology-I. Carnegie Inst. Wash. Publ., 1942, n° 536. [p.227, Figs.71-83]. As Onchocalanus nudipes.
Female (from Pacific): 71, habitus (dorsal); 72, A2; 73, Mx1; 74, Mx2; 75, Mxp; 76, P1; 77, endopod of P1 (enlarged); 78, P1 from another specimen; 79, P2; 80, P2 from another specimen); 81, P3; 82, P4; 83, P5.

Nota: Probably fig.83 is the P5 from juvenile male and an error in the legend (p.226), or an intersex form. After the author numerpous stages of development were taken in the eastern, central, and western parts of the tropical Pacific. No adult males were obtained, althrough some of the younger stages were probably undeveloped males. See in Vervoort's (1950 b, p.83) commentaries concerning the synonymy and the redescription of this species.


Espèce Phaenna spinifera - Planche 29 de figures morphologiquesissued from : W. Vervoort in Proc. K. ned. Akad. Wet., 1950, LIII, 1. [p.84, Fig.1].
Female (from ''Carnegie'' collection, 1°32'S, 82°16'W): a, habitus (dorsal view); b, urosome (lateral view from left side; c, left Md (mandibular palpus).

Nota: Urosome very short, 4-segmented, the 4th (anal) segment completely telescoped into the preceding. Proportional length of segments and caudal rami 44 : 19 : 15 : 4 : 18 = 100. Urosomal segments with some scattered hair-like scales of characteristic appearance.


Espèce Phaenna spinifera - Planche 30 de figures morphologiquesissued from : W. Vervoort in Proc. K. ned. Akad. Wet., 1950, LIII, 1. [p.85, Fig.2].
Female: a, urosome (dorsal); b, same (ventral); c, head (lateral); d, A2 (posterior surface).

Nota: Genital segment ventrally with a broadly rounded genital flap, covering the orifices of 2 small, scarcely visible receptacles.

Rem.: After Vervvort (1950, p.86) Wilson's drawings of the A2, Mx1, Mx2 and Mxp all are very inaccurate with regards to the number of setae on the various lobes and segments.


Espèce Phaenna spinifera - Planche 31 de figures morphologiquesissued from : W. Vervoort in Proc. K. ned. Akad. Wet., 1950, LIII, 1. [p.85, Fig.3].
Female: a, left Md (manducatory plate, interior surface); b, left Mx1 (posterior surface); c, left Mxp (anterior surface).

Nota: Mx1 small; 1st inner lobe elongated, carrying 6 strong, more or less plumose setiform spines and 4 smaller setae, one of which is very small and delicate; 2nd inner lobe with 1 single seta; 3rd inner lobe with 3 setae; 1st outer lobe with 9 strong, plumose setae; exopod with 5 setae; 2nd basal segment with 3 setae; endopod with 6 setae, the jointing between the endopodal joints is indistinct, but apparently there is 1 seta on the 1st, 1 on the 2nd and 4 on the 3rd (terminal ) joint.


Espèce Phaenna spinifera - Planche 32 de figures morphologiquesissued from : W. Vervoort in Proc. K. ned. Akad. Wet., 1950, LIII, 1. [p.87, Fig.4, b].
Female: b, right Mx2 (interior surface).

Nota: Mx2 strongly developed. 1st basal segment carries 4 lobes, on the 1st lobe there are 4 setae of about equal length and 1 smaller seta, on the 2nd lobe there are 3 equal setae, on the 3rd 2 subequal setae, and on the 4th 1 strong, spinulose seta and a much smaller and shorter seta. The denticles on the last mentioned strong seta on the 4th lobe are minute. 2nd basal segment terminating into a single (5th) lobe, which carries a strong, curved, spiniform seta (this seta is much longer than the other setae, there are rather coarse denticles along the convex border, from the apex down to about half its length), in addition there are 2 muvch smaller setae on the 5th lobe. The segmentation of the endopod is very indistinct; there are 6 brush-shaped sensory appendages and 2 much longer, filiform appendages (the extremities of these sensory filaments are coverd by a mucous layer. There is no seta along the external margin of Mx2.


Espèce Phaenna spinifera - Planche 33 de figures morphologiquesissued from : W. Vervoort in Proc. K. ned. Akad. Wet., 1950, LIII, 1. [p.90, Fig.5].
Female: a, left P1; b, left P2; c, left P3; d, left P4
Of all legs the posterior surface has been figured. b, c, d at the same scale; a: magnified twofold from the others.

After vervoort (1950, p.86) the figures of P3 and P4 apparently have been changed in Wilson's paper (fig.81 represents P4, fig.82 the P3.


Espèce Phaenna spinifera - Planche 34 de figures morphologiquesissued from : W. Vervoort in Proc. K. ned. Akad. Wet., 1950, LIII, 1. [p.90, Fig.6, b].
Male: b, adult habitus (dorsal aspect).


Espèce Phaenna spinifera - Planche 35 de figures morphologiquesissued from : W. Vervoort in Proc. K. ned. Akad. Wet., 1950, LIII, 1. [p.90, Fig.6, a].
Male 5th copepodid stage: a, urosome (dorsal view).


Espèce Phaenna spinifera - Planche 36 de figures morphologiquesissued from : W. Vervoort in Proc. K. ned. Akad. Wet., 1950, LIII, 1. [p.87, Fig.4, a].
Male 5th copepodid stage: a, P5 (anterior surface).


Espèce Phaenna spinifera - Planche 37 de figures morphologiquesissued from : Mulyadi in Published by Res. Center Biol., Indonesia Inst. Sci. Bogor, 2004. [p.188, Fig.107].
Female (from East Indonesian Seas): a, habitus (dorsal); b, urosome (dorsal); c, P1; d, P2.

Male: e, habitus (dorsal); f, P5.

Nota Female: Exopod segment 2 armed with transverse rows of acicular spines on posterior surface. P5 absent.


Espèce Phaenna spinifera - Planche 38 de figures morphologiquesIssued from : O. Tanaka in Publ. Seto Mar. Biol. Lab., 1960, VIII (1). [p.85, Fig.1].
Immature female (from Sagami, middle Japan): a, habitus (dorsal); b, distal segments of Mx2; c, P1; d, P2.
Nota: Body size = 1.81 mm.


Espèce Phaenna spinifera - Planche 39 de figures morphologiquesIssued from : H.Y. Soh, S.Y. Moon & J.H. Wi in Invertebrate Fauna of Korea (eds) Incheon: NIBR, 2013, 21 (28). [p.76, Fig.43].
Female (from Korean waters): A-B, habitus (dorsal and lateral, respectively); C, forehead (ventral); D, urosome (ventral).
Scale bars: A, B = 400 µm; C = 200 µm; D = 100 µm.


Espèce Phaenna spinifera - Planche 40 de figures morphologiquesIssued from : H.Y. Soh, S.Y. Moon & J.H. Wi in Invertebrate Fauna of Korea (eds) Incheon: NIBR, 2013, 21 (28). [p.77, Fig.44].
Female: A, P1; B, P2; C, P3; D, P4.
Scale bars: 100 µm.

Ref. compl.:
Cleve, 1904 a (p.194); Pearson, 1906 (p.21, Rem.); Wilson, 1942 a (p.202); 1950 (p.285); Yamazi, 1958 (p.149, Rem.); Fagetti, 1962 (p.22); Gaudy, 1962 (p.93, 99, Rem.: p.107); Duran, 1963 (p.16); V.N. Greze, 1963 a (tabl.2); Gaudy, 1963 (p.23, Rem.); De Decker & Mombeck, 1964 (p.13); Shmeleva, 1965 b (p.1350, lengths-volume -weight relation); Mazza, 1966 (p.70); Pavlova, 1966 (p.43); Ehrhardt, 1967 (p.738, geographic distribution, Rem.); Fleminger, 1967 a (tabl.1); Delalo, 1968 (p.138); Evans, 1968 (p.14); Berdugo & Kimor, 1968 (p.448); Dowidar & El-Maghraby, 1970 (p.269); Morris, 1970 (p.2300); Deevey, 1971 (p.224); Bainbridge, 1972 (p.61, Appendix Table III: occurrence); Apostolopoulou, 1972 (p.327, 349); Binet & al., 1972 (p.55); Roe, 1972 (p.277, tabl.1, tabl.2); Björnberg, 1973 (p.330, 388); Corral Estrada & Pereiro Muñoz, 1974 (tab.I); Vives & al., 1975 (p.42, tab.II); Deevey & Brooks, 1977 (p.256, tab.2, Station "S"); Carter, 1977 (1978) (p.35); Dessier, 1979 (p.205); Vaissière & Séguin, 1980 (p.23, tab.3); Porumb, 1980 (p.168); Kovalev & Shmeleva, 1982 (p.83); Vives, 1982 (p.291); Dessier, 1983 (p.89, Tableau 1, Rem., %); Turner & Dagg, 1983 (p.16, 22); De Decker, 1984 (p.316); Brenning, 1984 (p.3, Rem.); Scotto di Carlo & al., 1984 (p.1044); Cummings, 1984 (p.163, Table 2); Guangshan & Honglin, 1984 (p.118, tab.); Almeida Prado Por, 1985 (p.250); Brenning, 1985 a (p.28, Table 2); Brinton & al., 1986 (p.228, Table 1); Madhupratap & Haridas, 1986 (p.105, tab.1); Jimenez-Perez & Lara-Lara, 1988; Lozano Soldevilla & al., 1988 (p.58); Hernandez-Trujillo, 1989 a (tab.1); Cervantes-Duarte & Hernandez-Trujillo, 1989 (tab.3); Heinrich, 1990 (p.17); Suarez & al., 1990 (tab.2); Suarez & Gasca, 1991 (tab.2); Hernandez-Trujillo, 1991 (1993) (tab.I); Suarez, 1992 (App.1); Seguin & al., 1993 (p.23); Shih & Young, 1995 (p.71); Suarez-Morales & Gasca, 1997 (p.1525); Hure & Krsinic, 1998 (p.101); Gilabert & Moreno, 1998 (tab.1, 2); Suarez-Morales & Gasca, 1998 a (p.110); Smith S. & al., 1998 (p.2369, Table 6, moonsoon effects); Neumann-Leitao & al., 1999 (p.153, tab.2); Siokou-Frangou, 1999 (p.476); Lapernat, 2000 (tabl.3, 4); Razouls & al., 2000 (p.343, Appendix); El-Sherif & Aboul Ezz, 2000 (p.61, Table 3: occurrence); Moraitou-Apostolopoulou & al., 2000 (tab.I); Lopez-Salgado & al., 2000 (tab.1); Holmes, 2001 (p.56, Rem.); Hidalgo & Escribano, 2001 (p.159, tab.2); Sameoto & al., 2002 (p.13); Beaugrand & al., 2002 (p.179, figs.5, 6); Vukanic, 2003 (139, tab.1); Hwang & al., 2003 (p.193, tab.2); Shimode & Shirayama, 2004 (tab.2); Hsiao & al., 2004 (p.326, tab.1); Gallienne & al., 2004 (p.5, tab.3); Lo & al., 2004 (p.89, tab.1); Zuo & al., 2006 (p.162: tab.1); Lavaniegos & Jiménez-Pérez, 2006 (p.151, tab.2, 4, Rem.); Dur & al., 2007 (p.197, Table IV); Khelifi-Touhami & al., 2007 (p.327, Table 1); Jitlang & al., 2008 (p.65, Table 1); Ayon & al., 2008 (p.238, Table 4: Peruvian samples); Cabal & al., 2008 (289, Table 1); Neumann-Leitao & al., 2008 (p.799: Tab.II, fig.6); Morales-Ramirez Suarez-Morales, 2008 (p.521); Gaard & al., 2008 (p.59, Table 1, N Mid-Atlantic Ridge); Galbraith, 2009 (pers. comm.); Williamson & McGowan, 2010 (p.273, Table III, Pacific central gyres: N & S); Hernandez-Trujillo & al., 2010 (p.913, Table 2); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical/tropical); Hidalgo & al., 2010 (p.2089, Table 2); Mazzocchi & Di Capua, 2010 (p.426); Medellin-Mora & Navas S., 2010 (p.265, Tab. 2); Hsiao S.H. & al., 2011 (p.475, Appendix I); Selifonova, 2011 a (p.77, Table 1, alien species in Black Sea); Tutasi & al., 2011 (p.791, Table 2, abundance distribution vs La Niña event); Shiganova & al., 2012 (p.61, Table 4); Uysal & Shmeleva, 2012 (p.909, Table I); Salah S. & al., 2012 (p.155, Tableau 1); Dorgham & al., 2012 (p.473, Table 4: abundance vs season); Palomares-Garcia & al., 2013 (p.1009, Table I, V, abundance vs environmental factors); in CalCOFI regional list (MDO, Nov. 2013; M. Ohman, comm. pers.); Hirai & al., 2013 (p.1, Table I, molecular marker); Lidvanov & al., 2013 (p.290, Table 2, % composition)
NZ: 21

Carte de distribution de Phaenna spinifera par zones géographiques
Espèce Phaenna spinifera - Carte de distribution 4
Loc:
Cosmopolite (tropical et temperate): South Africa (S & E), Atlant., G. of Guinea, Brazil (off Natal), Caribbean Sea, Caribbean Colombia, Yucatan, G. of Mexico, Florida, off Bermuda: Station "S" (32°10'N, 64°30'W), Long Island, off E Nova Scotia, G. of Guinea, off NW Cape Verde Is., off Morocco-Mauritania, Cap Ghir (Morocco), Canary Is., off Madeira, Azores, Bay of Biscay, off W Cabo Finisterre, off W Ireland, Ibero-moroccan Bay, Medit. (Alboran Sea, Gulf of Annaba, Banyuls, G. of Lion, Marseille, Tyrrhenian Sea, Napoli, Messina, Malta, Adriatic Sea, Ionian Sea, Aegean Sea, Lebanon Basin, Alexandria, Black Sea), Sharm El-Sheikh, Safaga, Red Sea, Arabian Sea, Natal, Mascarene Basin, Rodrigues Is. - Seychelles, off SW Sri Lanka, Bay of Bengal, Flores Sea, Celebes Sea, Banda Sea, Ambon Bay (SW Ceram Is.), China Seas (East China Sea, South China Sea), Korea, Taiwan (SW, E, N: Mienhua Canyon), Japan (Tanabe Bay), Vancouver Is., Pacif. (W equatorial), Pacific (central gyres: N & S), BaJa California (Bahia Magdalena, W), Gulf of California, W Costa Rica, Galapagos-Ecuador, Peru, Chile (N-S, Mejillones Peninsula); Also sub-Antarct. (Pacif. SE), Straits of Magellan (Pacific area)
N: 182
Lg.:
(7) F: 2,9; M: 2,26; (9) F: 2,3; 2,05; M: 2,5; (15) F: 2,22-2,2; (16) F: 2,3-2,13; (35) F: 2,23; M: 2,1; (36) F: 2,88-3,02; (38) F: 2,34; M: 2,22; (46) F: 2,1-1,8; M: 1,8; (57) F: 2,25; (72) F: 2,4-1,84; M: 1,94; (101) F: 1,9-1,6; (116) F: 2,2; (180) F: 1,82; (187) F: 2,12; (199) F: 2,2-1,98; M: 1,98; (207) F: 1,94-1,89; (231) M: 2,5; (290) F: 1,9-2; M: 1,9-1,95; (313) M: 2,5; (333) F: 1,5; (335) F: 1,71; (447) F: 2,07; M: 1,89; (449) F: 2,1-1,8; M: 1,8; (867) F: 1,6-2,73; (991) F: 1,5-2,9; M: 1,8-2,5; (1023) F: 1,89; (1111) F: 2,16; (1122) F: 2,1; (1174) F: 2,45; {F: 1,50-3,02; M: 1,80-2,50}
Rem.: épi-mésopélagique. After Pearson (1906, p.21): vertical range down to 2900 m.
Sampling depth (sub-Antarct.) : 250 m.
With (1915) signale la présence d’une petite protubérance dorsale sur le cephalosome de certains individus. Caractéristique observée également au large de la côte algérienne (Kerzabi, 1999, comm. pers.).
Voir aussi les remarques en anglais
Dernière mise à jour : 18/10/2016

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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2017. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Disponible sur http://copepodes.obs-banyuls.fr 
[Accédé le 26 avril 2017]

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