List species and varieties by family
Centropagidae Giesbrecht, 1892 ( Diaptomoidea )
(1) Centropages Kröyer, 1849
Rem.: type: Centropages typicus Krøyer, 1849 . Total: 35 spp. (3 doubtful).

Diagnosis after Bradford-Grieve (1999 b, p.131) :
- As for the family definition.
- Urosomr 3-segmented in female.
- Genital segment female usually asymmetrical.
- Urosome male 5-segmented.
- Mx1 inner lobe 1 with 15 spines and setae; inner lobe 2 with 3 setae; inner lobe 3 with 3 setae; basipod 1 with 5 setae; endopod with 4+5 setae, segments 1 and 2 fused to basis; exopod with 9 setae, outer lobe 2 with 1 small seta, outer lobe 1 with 9 setae, 7 of them long.
- Exopod segment 3 of P2 - P4 with 3 outer-edge spines.
- Female P5 basipod 2 not ornamented, exopod segment 2 with a large, straight inner edge spine.
- Male P5 with basipod 2 unornamented; left exopod 2-segmented, segment 1 with 1 outer edge spine; segment 2with 3 outer edge spines and 1 small terminal spine; both endopods as in female; right exopod 3-segmented, in the form of a chela.

[1] Centropages abdominalis  Sato, 1913   (F,M)    [Figs]

[2] Centropages acutus  McKinnon & Dixon, 1994   (F,M)    [Figs]

[3] Centropages aegypticus  El-Sherbiny & Ueda, 2008   (F,M)    [Figs]

[4] Centropages alcocki  Sewell, 1912   (F,M)    [Figs]

Centropages arabicus    Cleve, 1904   (F,M)
Ref.: Cleve,1904 (p.371, figs.F,M)
Loc: Mer Arabe
Rem.: Cf. Centropages tenuiremis

[5] Centropages aucklandicus  Krämer, 1895   (F,M)    [Figs]

[6] Centropages australiensis  Fairbridge, 1944   (F,M)    [Figs]

[7] Centropages brachiatus  (Dana, 1849)   (F,M)    [Figs]

[8] Centropages bradyi  Wheeler, 1901   (F,M)    [Figs]

Centropages brevicaudatus    Brady, 1875   (F,M)
Ref.: Brady, 1918 (p.24, figs.F,M, Rem.)
Loc: Falkland Is., Kerguelen
Rem.: eau douce, saumâtre. Cf. Boeckella brevicaudata

[9] Centropages brevifurcus  Shen & Lee, 1963   (F,M)    [Figs]

[10] Centropages calaninus  (Dana, 1849)   (F,M)    [Figs]

[11] Centropages caribbeanensis  Park, 1970   (F,M)    [Figs]

[12] Centropages chierchiae  Giesbrecht, 1889   (F,M)    [Figs]

Centropages chilensis    Kröyer, 1849   
Ref.: Kröyer, 1849 (in Damkaer & Damkaer, 1979 (p.43); Giesbrecht & Schmeil, 1898 (p.55); Vervoort, 1964 b (p.308)
Loc: Chili
Rem.: Cf. Centropages brachiatus

Centropages discaudatus    Brady, 1899   
Ref.: Vervoort, 1964 b (301)
Loc: Nouvelle-Zélande
Rem.: Cf. Centropages aucklandicus

[13] Centropages dorsispinatus  Thompson & Scott, 1903   (F,M)    [Figs]

[14] Centropages elegans  Giesbrecht, 1895   (F,M)    [Figs]

[15] Centropages elongatus  Giesbrecht, 1896   (F,M)    [Figs]

[16] Centropages furcatus  (Dana, 1849)   (F,M)    [Figs]

Centropages furcatus carli    Früchtl, 1923   (M)
Syn.: C. furcatus : Carl, 1907 (p.8, figs.M)
Ref.: Früchtl, 1923 (p.452); 1924 b (p.45, figs.M); Sewell, 1932 (p.229, Rem.M); Vervoort, 1964 b (p.308)
Loc: Indonésie-Malaisie (Baie d'Amboine)

[17] Centropages gracilis  (Dana, 1849)   (F,M)    [Figs]

Centropages grimaldii    De Guerne, 1886   
Ref.: De Guerne, 1886 (p.277); Giesbrecht & Schmeil, 1898 (p.59)
Loc: G. de Finlande
Rem.: Cf. Limnocalanus grimaldii

[18] Centropages halinus  McKinnon & Kimmerer, 1988   (F,M)    [Figs]

[19] Centropages hamatus  (Lilljeborg, 1853)   (F,M)    [Figs]

[20] Centropages karachiensis  Haq & Fazal-Ur-Rehman, 1973   (F,M)    [Figs]

[21] Centropages kroyeri  Giesbrecht, 1892   (F,M)    [Figs]

Centropages kroyeri pontica    Karavaev, 1894   (F,M)
Ref.: Karavaev, 1894 (p.24, figs.F,M); Vervoort, 1964 b (p.309); Soler & al., 1988 (p.129, 130, Rem.)
Loc: Mer Noire
Rem.: Cf. Centropages ponticus

Centropages kroyeri var. Sydney    Dakin & Colefax, 1940   (F,M)
Ref.: Dakin & Colefax, 1940 (p.91, figs.F,M)
Loc: Australie (Nouvelle-Galles du Sud)
Lg.: (104) F: 1,7; M: 1,4
Rem.: Les auteurs suggèrent qu’il pourrait s’agir d’une espèce nouvelle, notamment par les caractères de la femelle.

Centropages lenucculari    Oliveira, 1945   
Ref.: Vervoort, 1965 (p.88, 89, Rem.)
Loc: Rio de Janeiro
Rem.: Cf. Centropages furcatus

[22] Centropages longicornis  Mori, 1932   (F)    [Figs]

[23] Centropages maigo  Ohtsuka, Itoh & Mizushima, 2005   (F,M)    [Figs]

Centropages mcmurrichi    Willey, 1920   (F,M)
Ref.: Willey, 1920 a (p.10, Rem. F); Sewell, 1948 (p.401, Rem.); Johnson, 1958 (p.274, Rem.); 1961 (p.317, Rem.); Vervoort, 1964 b (p.309); Cross & Small, 1967 (p.60); Kim & al., 1993 (p.269)
Loc: Alaska, Détroit de Géorgie, Baie d'Hudson, mers de Chine, Hong Kong, Corée S, Oregon
Rem.: Cf. Centropages abdominalis

[24] Centropages mohamedi  El-Sherbiny & Al-Aidaroos, 2015   (F,M)    [Figs]

[25] Centropages natalensis  Connell, 1981   (F,M)    [Figs]

Centropages notoceras    Cleve, 1904   (F,M)
Ref.: Cleve, 1904 (p.373, Descr.F, figs.F); Sewell, 1912 (p.360, Redescr.F,M, Rem.); 1932 (p.228, Rem.); Vervoort, 1964 b (p.308)
Loc: Mer Arabe, Ceylan, Chittagong
Rem.: Cf. Centropages dorsispinatus

[26] Centropages orsinii  Giesbrecht, 1889   (F,M)    [Figs]

Centropages pacificus    Chiba, 1956   (M)
Ref.: Bikini Atoll, Indien (I. Maurice), Afr. S (E) (in De Decker, 1964, p.15)
Lg.: (786) M: 1,75-1,63
Rem.: sp. douteuse. Cf. ? Centropages elongatus

[27] Centropages ponticus  Karavaev, 1894   (F,M)    [Figs]

[28] Centropages sinensis  Chen & Zhang, 1965   (F,M)    [Figs]

[29] Centropages spinosus  (Kriczagin, 1873)   (F,M)    [Figs]

Centropages tenuicornis    Dana, 1849   (F,M)
Syn.: Cyclopsina tenuicornis Dana,1849
Ref.: Giesbrecht & Schmeil, 1898 (p.58, Rem.); Vervoort, 1964 b (p.309)
Loc: Pacif.
Rem.: forme douteuse. Cf. ? Centropages calaninus

[30] Centropages tenuicornis Brady  Brady, 1914   (M)    [Figs]

[31] Centropages tenuiremis  Thompson & Scott, 1903   (F,M)    [Figs]

[32] Centropages trispinosus  Sewell, 1914   (F,M)    [Figs]

[33] Centropages typicus  Kröyer, 1849   (F,M)    [Figs]

Centropages typicus aucklandicus    Krämer, 1895   
Ref.: Vervoort, 1964 b (p.301, Rem.)
Loc: Nouvelle-Zélande (Hauraki Gulf)
Rem.: Cf. Centropages aucklandicus

[34] Centropages uedai  El-Sherbiny, 2011   (F,M)    [Figs]

[35] Centropages velificatus  (Oliveira, 1947)   (F,M)    [Figs]

[36] Centropages violaceus  (Claus, 1863)   (F,M)    [Figs]

Centropages yamadai    Mori, 1934   (F,M)
Ref.: Mori, 1937 (1964) (p.59, figs.F,M); Tanaka, 1963 (p.11, figs.F,M); Vervoort, 1964 b (p.309); Kasahara & al, 1974 (p.170, fig. O); Hirota & Uno, 1977 (p.77, fig.O)
Loc: Corée, Japon, G. Arabe (in Michel & Herring, 1984)
Lg.: (26) F: 1,5; M: 1,41-1,32; (91) F: 1,7-1,5; M: 1,54-1,27
Rem.: Cf. Centropages tenuiremis
(2) Gippslandia Bayly & Arnott, 1969
Rem.: type: Gippslandia estuarina Bayly & Arnott, 1969. Total: 1 sp.

[1] Gippslandia estuarina  Bayly & Arnott, 1969   (F,M)    [Figs]
(3) Isias Boeck, 1864
Rem.: Type: Isias clavipes Boeck, 1865. Total: 4 spp.

Diagnosis after Bradford-Grieve (1999 b, p.145) :
- As for the family definition.
- Exopod segment 3 of P2 - P4 with 3 outer edge spines.
- Female P5 basipod 2 ornamented with a large distal spine, exopod segment 2 with a medium, curved inner edge spine, exopod segments 2 and 3 may be fused; endopods 1-segmented, with a variable number of setae.
- Male P5 with basipod 2 usually ornamented asymmetrically with large spines or projections; left exopod 2-segmented, segment 1 with 1 outer edge spine, segment 2 with a variable number of spines; endopods reduced or absent; right exopod 2-segmented, segment 1 with 1 outer edge spine, segment 2 may or may not be expanded with 3 outer edge spines and 1 terminal spine.

Isias bonnieri    Canu, 1888   (F,M, Juv.)
Ref.: Canu, 1888 b (p.228, figs.F,M, juv.); Giesbrecht & Schmeil, 1898 (p.62)
Rem.: Cf. Isias clavipes

[1] Isias clavipes  Boeck, 1864   (F,M)    [Figs]

[2] Isias cochinensis   Pillai P. Parameswaran, 1975   (F,M)    [Figs]

[3] Isias tropica  Sewell, 1924   (F,M)    [Figs]

[4] Isias uncipes  Bayly, 1964   (F,M)    [Figs]
(4) Parathalassius Dussart, 1986
Rem.: Type: Parathalassius fagesi Dussart, 1986. Total: 1 sp.
In brackish water.

Definition from Bradford-Grieve (1999 b, p.147) :
- As for the family definition.
- Urosome female 4-segmented.
- Urosome male 5-segmented.
- A1 24-segmented.
- Left A1 male 24-segmented; 22-segmented on the right.
- P1-P4 endopod 3-segmented, with segment 3 supplied with external setae.
- P5 male and female biramous, each ramus 3-segmented.
- P5 male hardly modified compared with other swimming legs.

This genus is nearest ro the freshwater genera Pseudoboeckella and Hemiboeckella but the primitive character of P5 seems to make this genus the most primitive all known of Centropagidae (Dussart, 1986)

[1] Parathalassius fagesi  Dussart, 1986   (F,M)    [Figs]
(5) Sinocalanus Burckhardt, 1913
Rem.: Fresh-water and in estuaries.
Total: 3 spp.
Characteristics of the genus (After J.J. Orsi, T.E. Bowman, D.C. Marelli & A. Hutchinson, 1983, p.358-359) : 1- Pediger 5 with spine on each posterior corner. 2- Caudal rami without surface spinules. 3- [A2 exopod longer than endopod.] Exopod of A2 longer than endopod 4- A2 distal endopod segment not longer than wide. 5- A2 exopod 2nd segment not distally bilobed. 6- Mxp 2nd basipod segment with modified short setae.

Nota: L.A. Bledzki informs us (29.08.2015) the error in the diagnosis from Orsi & al. (1983, p.358). 7- P5 female with 2nd exopod segment without outer spine ; 1st endopod segment without inner seta. 8- Right P5 male with 1st basipod segment withoutmedial process ; 2nd basipod segment with medial process (except in S. solstitialis).

[1] Sinocalanus doerrii  (Brehm, 1909)   (F,M)    [Figs]

[2] Sinocalanus sinensis  (Poppe, 1889)   (F,M)    [Figs]

[3] Sinocalanus tenellus  Kikuchi, 1928   (F,M)    [Figs]

 Any use of this site for a publication will be mentioned with the following reference :

Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2021. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed November 28, 2021]

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