List species and varieties by family
Eucalanidae Giesbrecht, 1892 ( Eucalanoidea )
(1) Eucalanus Dana, 1852 ( Eucalaninae )
Rem.: Type: Calanus elongatus (Dana,1849).
Not all authors follow Geletin (1976) in his defenition of the genera Pareucalanus and Subeucalanus (Brodsky & al., 1983, p.203).
Fleminger (1973) splits up 17 spp. into 4 groups: 'subtenuis', 'pileatus', 'elongatus', 'attenuatus'.
17 spp. + 1 debated (E. pseudattenuatus ) + 2 to specify (E. muticus , E. peruanus ) + 3 very doubtful.
Bradford-Grieve (1994) rehabilitates the two genera Pareucalanus and Subeucalanus.
The genus Eucalanus comprises 6 spp. + 1 doubtful.

Diagnosis from Bradford-Grieve (1994, p.74) :
- As in the family definition.
- Urosome female 4-segmented, and posterior borders of pedigers 2-4 and urosome segment 1 without spines.
- Integumental organs located at Mx1 and Mx2 sites vary from 8-18 in number are not arranged in a rectangular pattern, posterior sets more lateral than the anterior sets. Mxp sites are in 4 or more pairs. Pediger segments 2 and 3 with 1 or 2 lateral sites ; pediger segment 4 lacking paired sites lying outside the dorsal quadrangle of hair sensilla ; abdominal segment 1-2 with at least 2 dorsal perforations.
- A2 endopod elongate, exopodal segments 1 and 2 separate.
- Md palp basis and exopod elongate ; endopod small, inserted just beyond midlength on basis which also bears 1 or 3 setae.
- Mx1 with 3 inner lobes bearing setae.
- P1 with a 2-segmented endopod and 3-segmented exopod.
P5 absent in female.
- P5 uniramous on both sides in male.

Eucalanus acus    Farran, 1929   (F)
Ref.: Farran, 1929 (p.218, figs.F); Sewell, 1948 (p.513, Rem.)
Rem.: Cf. Subeucalanus longiceps

Eucalanus atlanticus    Wolfenden, 1904   (M)
Ref.: Wolfenden, 1904 (p.113, figs.M); With, 1915 (p.48, Rem.); Candeias, 1926 (1929) (p.27, figs.M); Rose, 1933 a (p.69, figs.M, Rem.)
Rem.: Cf. Eucalanus elongatus (juv.)

Eucalanus attenuatus    Dana, 1849   (F,M)
Syn.: Calanus attenuatus Dana,1849
Ref.: Bradford-Grieve, 1994 (p.80, Rem.)
Rem.: Cf. Pareucalanus attenuatus

[1] Eucalanus bungii  Giesbrecht, 1892   (F,M)    [Figs]

Eucalanus bungii bungii    Johnson, 1938   (F,M)
Ref.: M.W Johnson, 1938 (p.167, figs.F,M); Vervoort, 1946 (p.91, Rem.); Sewell, 1948 (p.561, Rem.); Brodsky, 1950 (1967) (p.101, 102, figs.F, M); Mazza, 1967 (p.71, Rem.); Vidal, 1971 a (p.16, figs.F,M); Shih & al., 1971 (p.146, 204); Tanaka, 1956 (p.267, Rem.)
Ref. compl.: Harding, 1966 (p.17, 69); Dunbar & Harding, 1968 (p.319, 322); Omori, 1969 (p.5); Morris, 1970 (p.2300); Shih & Laubitz, 1978 (p.53); Van der Spoel & Heyman, 1983 (p.94, fig.58); Gordon & al., 1985 (p.91, 92); Sasaki & al., 1988 (tab.1); Coyle & al., 1990 (p.763); Hirakawa & al., 1990 (tab.3); Hattori, 1991 (tab.1, Appendix); Shih & Young, 1995 (p.69); Kotani & al., 1996 (tab.2); Park & Choi, 1997 (Appendix); Kosobokova & Hirche, 2000 (p.2029, tab.2)
Loc: Arct. (Fletcher's Ice Is.), Alaska (Auke Bay), off I. Reine Charlotte, Colombie Britannique, Arct. (bassin polaire) (rare), Japon, off Hokkaido, Corée, Mer de Chine E, Pacif. central N (subarct.)
N: 19
Lg.: (22) F: 8-6,5; M: 5,4-4,8
Rem.: Delalo (1968, p.137) signale cette forme en Mer Rouge et G. d'Aden ce qui nécessite confirmation. Cf. Eucalanus bungii.

Eucalanus bungii californicus    Johnson, 1938   (F,M)
Syn.: Eucalanus bungii : Davis, 1949 (p.16, figs.F,M)
Ref.: M. W. Johnson, 1938 (p.169, figs.F); Vervoort, 1946 (p.91, Rem.); Sewell, 1948 (p.561, Rem.); Mazza, 1967 (p.71, Rem.); Morris, 1970 (p.2300); Shih & al., 1971 (p.147); Hernandez-Trujillo, 1991 (1993) (tab.I); Van der Spoel & Heyman, 1983 (p.94, fig.58)
Loc: Détr. Juan de Fuca, I. Malcolm
Rem.: Cf. Eucalanus californicus

[2] Eucalanus californicus  Johnson, 1938   (F,M)    [Figs]

Eucalanus crassus    Giesbrecht, 1888   (F,M)
Syn.: Eucalanus oculanus Marukawa, 1921 (p.11); Brodsky, 1950 (1967) (p.105)
Ref.: Bradford-Grieve, 1994 (p.87)
Rem.: Cf. Subeucalanus crassus

Eucalanus danai    Lubbock, 1860   
Syn.: Calanus danai Lubbock,1860
Ref.: Giesbrecht, 1892 (p.132); Giesbrecht & Schmeil, 1898 (p.22, Rem.)
Loc: G. de Guinée
Rem.: sp. douteuse

Eucalanus dentatus    A. Scott, 1909   (F)
Ref.: A. Scott, 1909 (p.22, figs.F); Geletin, 1976 (p.84); Fleminger, 1973 (p.978, 999, figs.F)
Loc: Indonésie-Malaisie
Rem.: Cf. Subeucalanus dentatus

[3] Eucalanus elongatus  (Dana, 1849)   (F,M)    [Figs]

Eucalanus elongatus bungii    Giesbrecht, 1892   
Ref.: Giesbrecht, 1892 (p.149); Vervoort, 1946 (p.91, Rem.); Davis, 1949 (p.16)
Rem.: Cf. Eucalanus bungii ou californicus

Eucalanus elongatus hyalinus    Claus, 1866   (F,M)
Syn.: Calanella hyalina Claus,1866; Eucalanus spinifer T. Scott,1894 b ; Eucalanus atlanticus Wolfenden,1904
Ref.: Giesbrecht, 1892 (p.131, 149, figs.F,M, Rem.); M.W. Johnson, 1938 (p.167, 169, fig.F); Vervoort, 1946 (p.91, 92, Rem.); 1957 (p.32, Rem.); 1963 b (Rem., p.88); Sewell, 1948 (p.561, Rem.); Davis, 1949 (p.16); Marques, 1968 (p.395); Corral Estrada, 1970 (p.72, Rem.); Mazzocchi & al., 1995 (p.116, figs.F, Rem.)
Rem.: Cf. Eucalanus hyalinus

Eucalanus elongatus inermis    Giesbrecht, 1892   
Ref.: Giesbrecht, 1892 (p.149); Vervoort, 1946 (p.91, 93, Rem.); 1963 b (p.89, Rem.); Sewell, 1948 (p.561, Rem.); Davis, 1949 (p.16)
Rem.: Cf. Eucalanus inermis

Eucalanus giesbrechti    Mori, 1937   (F,M juv.)
Syn.: Eucalanus elongatus bungii Giesbrecht,1892 (part.); Eucalanus elongatus (F): Esterly,1905 (p.131,figs.F,M)
Ref.: Mori, 1937 (1964) (p.22, figs.F, juv.M); Davis, 1949 (p.17, Rem.); Vervoort, 1957 (p.32)
Rem.: Cf. Eucalanus californicus

[4] Eucalanus hyalinus  (Claus, 1866)   (F,M)    [Figs]

[5] Eucalanus inermis  Giesbrecht, 1892   (F,M)    [Figs]

Eucalanus langae    Fleminger, 1973   (F,M)
Ref.: Fleminger, 1973 (p.978, 990, 996, figs.)
Rem.: Cf. Pareucalanus langae

Eucalanus longiceps    Matthews, 1925   (F,M)
Syn.: Eucalanus acus Farran,1929 (p.218, figs.F); ? E. attenuatus : With, 1915 (p.52); Subeucalanus longiceps : Geletin, 1976 (p.91)
Ref.: Bradford-Grieve, 1994 (p.87)
Rem.: Cf. Subeucalanus longiceps

Eucalanus monachus    Giesbrecht, 1888   (F,M)
Syn.: Eucalanus vadicola F. Dahl,1894; non E. monachus : Dakin & Colefax, 1940
Ref.: Bradford-Grieve, 1994 (p.88)
Rem.: Cf. Subeucalanus monachus

Eucalanus mucronatus    Giesbrecht, 1888   (F,M)
Syn.: non E. mucronatus : Mori, 1937 (1964) (p.25, figs.F,M)
Ref.: Bradford-Grieve, 1994 (p.88)
Rem.: Cf. Subeucalanus mucronatus

[6] Eucalanus muticus  C.B. Wilson, 1950   (F,M)    [Figs]

Eucalanus oculanus    Marukawa, 1921   (F,M)
Ref.: Brodsky, 1950 (1967) (p.105, figs.F,M)
Rem.: Mori, 1937 (1964) (p.24) considère l'espèce comme synonyme de Eucalanus crassus (juv.). Cf. Subeucalanus crassus

Eucalanus parki    Fleminger, 1973   (F,M)
Syn.: Eucalanus attenuatus Park, 1968 (p.533, grandes formes); E. quadrisetosus Geletin, 1973 (p.1786: M)
Ref.: Bradford-Grieve, 1994 (p.80, Rem.)
Rem.: Cf. Pareucalanus parki

Eucalanus peruanus    Volkov, 1971   (F,M)
Ref.: Volkov, 1971 (p.141, figs.F,M)
Rem.: Cf. Pareucalanus peruanus

Eucalanus pileatus    Giesbrecht, 1888   (F,M)
Syn.: Eucalanus monachus : Dakin & Colefax,1940; E. saravatae Oliveira, 1945; E. subcrassus-pileatus : Deevey, 1960 (p.16); Björnberg, 1963 (p.20)
Ref.: Bradford-Grieve, 1994 (p.88)
Rem.: Cf. Subeucalanus pileatus

Eucalanus pseudattenuatus    Sewell, 1947   (F,M)
Syn.: Eucalanus pseudoattenuatus : Brodsky, 1962 c (p.112, figs.F,M); Chen & Zhang, 1965 (p.34, figs.F); Deevey, 1971 (p.224); Sazhina, 1985 a (p.491, tab.1)
Ref.: Sewell, 1947 (p.47, figs.F); Rose, 1956 (p.459); Kasturirangan, 1963 (p.17, 19, fig.F); Vervoort, 1963 b (p.90); Geletin, 1973 (p.1783, 1786, figs.); 1976 (p.83); Fleminger, 1973 (p.967, 990, 992); Chen Q-c, 1980 (p.794); Brodsky & al., 1983 (p.208); Zheng Zhong & al., 1984 (1989) (p.229, figs.F,M); Shih & Young, 1995 (p.69); Wong & al, 1998 (tab.2); Mauchline, 1998 (tab.30)
Lg.: (290) F: 3,75-3,8; (864) F: 4-4,2
Rem.: Cf. Pareucalanus attenuatus et sewelli

Eucalanus quadrisetosus    Geletin, 1973   (M)
Ref.: Geletin, 1973 (p.1786, figs.M, non F); Brodsky & al., 1983 (p.212, Rem.)
Rem.: Cf. Paereucalanus parki

Eucalanus saravatae    Oliveira, 1945   (F)
Ref.: Oliveira, 1945 a (p.456, fig.F); Björnberg & al., 1981 (p.604, Rem.)
Rem.: Cf. Eucalanus pileatus

Eucalanus setiger    Brady, 1883   (F,M)
Ref.: Brady, 1883 (p.39,figs.F,M); Thompson, 1888 d (p.139); Giesbrecht, 1892 (p.132); T. Scott, 1894 b (p.30); Giesbrecht & Schmeil, 1898 (p.22)
Rem.: Cf. ? Eucalanus subtenuis ou Subeucalanus crassus

Eucalanus sewelli    Fleminger, 1973   (F,M)
Syn.: E. attenuatus : Vervoort, 1946 (part., p.95); Sewell, 1947 (part., p.39, fig. 7B); Brodsky, 1962 (p.113, fig.14); Ramirez, 1969 (p.44); Björnberg, 1973 (p.226)
Ref.: Bradford-Grieve, 1994 (p.80, 82, Rem.)
Rem.: Cf. Pareucalanus sewelli

[7] Eucalanus spinifer  T. Scott, 1894   (F,M)    [Figs]

Eucalanus subcrassus    Giesbrecht, 1888   (F,M)
Ref.: Bradford-Grieve, 1994 (p.88)
Rem.: Cf. Subeucalanus subcrassus

Eucalanus subtenuis    Giesbrecht, 1888   (F,M)
Ref.: Giesbrecht, 1892 (p.132, 150, figs.F,M); Sars, 1925 (p.21); Farran, 1929 (p.218); Vervoort, 1946 (p.106); 1963 b (p.91); Carvalho, 1952 a (p.141, figs.M); Fukase, 1957 (p.17, figs.F,M, Rem.); Marques, 1968 (p.396); 1974 (p.12); 1982 (p.753); Fleminger, 1973 (p.969, 978, 979, 982, 999, figs.F); 1975 (p.395); Geletin, 1976 (p.83, 91, Rem.); Greenwood, 1976 (p.16); Van der Spoel & Heyman, 1983 (p.40, fig.52)
Rem.: Cf. Subeucalanus subtenuis

Eucalanus subtenuis japonica    Fukase, 1957   (F,M)
Syn.: Eucalanus subtenuis : Tanaka,1935; Eucalanus mucronatus : Mori, 1937 (1964) (p.25, figs.F,M)
Ref.: Fukase, 1957 (p.19, figs.F,M, Rem.); Grice, 1962 (p.181, figs.F,M, Rem.)
Loc: Japon
Rem.: Cf. Subeucalanus subtenuis japonica

Eucalanus vadicola    Dahl, 1894   (F,M)
Ref.: Dahl, 1894 c (p.20, Rem. F,M); Giesbrecht & Schmeil, 1898 (p.21); Björnberg, 1963 (p.21, Rem.)
Loc: Estuaire de l'Amazone
Rem.: Cf. Subeucalanus monachus

Eucalanus sp.    Candeias, 1926   (? M juv)
Ref.: Candeias, 1926 (1929) (p.28, figs.juv)
Loc: Portugal

Eucalanus sp.    Tanaka, 1935   (F,M)
Ref.: Tanaka, 1935 (p.146, figs.F,M)
Loc: Japon (Sagami Bay)
Lg.: (389) F: 6,19-5,72; M: 4,67-4,5; {F: 5,72-6,19; M: 4,5-5,72}
Rem.: Cette forme nécessite confirmation.
(2) Pareucalanus Geletin, 1976 ( Eucalaninae )
Rem.: type: Paracalanus attenuatus Dana,1849. Total: 6 spp. + 1 undetermined.
Brodsky & al., 1983 (p.203) do not acknowledge the creation of this genus. The genus is confirmed by Bradford-Grieve (1994)

Diagnosis from Bradford-Grieve (1994, p.80) :
- As in the family definition.
- Urosome female 3-segmented, and posterior borders of pedigers 2-4 and urosome segment 1 without spines.
- Integumental organs located at M11 and Mx2 with a total of 10 sites forming a rectangle and arranged anterior to posterior in 2 bilateral sets of 1 pair, 1 single and 1 pair of sites ; Mxp site has 6 pairs ; pedigers segments 2 and 3 in lateral view with 3 sites ; pediger segment 4 lacking paired sites lying outside of the dorsal quadrangle of hair sensilla ; urosome segment 4-5 with at least 3 dorsal sites.
- A2 endopod elongate, exopod segments 1 and 2 separate.
- Md palp basis and exopod elongate ; endopod small, inserted just at midlength on basis which also bears 2 or 4 setae.
- Mx1 with 3 inner lobes bearing setae.
- P1 with a 2-segmented endopod and 3-segmented exopod.
- P5 female absent.
- P5 male uniramous on both sides with right leg not extending beyond the distal part of left leg segment 2.

[1] Pareucalanus attenuatus  (Dana, 1849)   (F,M)    [Figs]

[2] Pareucalanus langae  (Fleminger, 1973)   (F,M)    [Figs]

[3] Pareucalanus parki  (Fleminger, 1973)   (F,M)    [Figs]

[4] Pareucalanus peruanus  (Volkov, 1971)   (F,M)

[5] Pareucalanus sewelli  (Fleminger, 1973)   (F,M)    [Figs]

[6] Pareucalanus smithae  Prusova, 2007   (F,M)    [Figs]

[7] Pareucalanus sp.  Goetze & Ohman, 2010       [Figs]
Rhincalanus Dana, 1852
Rem.: Cf. Rhincalanidae.
(3) Subeucalanus Geletin, 1976
Rem.: type: Eucalanus subtenuis Giesbrecht,1888. Total: 9 spp. + 3 undet.
Brodsky & al. (1983, p.203) do not acknowledge the creation of this genus. Bradford-Grieve (1994) affirms the validity of the genus.

Diagnosis from Bradford-Grieve (1994, p.69) :
- As in the family definition.
- Female urosome 3-segmented, and posterior borders of pedigers 2-4 and urosome segment 1 without spines.
- Integumental organs located at Mxp site in 2 pairs ; pediger segments 3 and 4 with a single mid-dorsal site lying between the anterior 2 hair sensilla ; pediger segment 4 with 1 pair of sites lateral to the quadrangle formed by 4 hair sensilla.
- A2 endopod elongate, exopod segments 1 and 2 fused.
- Md palp basis and exopod elongate ; endopod small, inserted on the terminal one third or quarter of basis which also bears 2 or 3 setae.
- Mx1 with 2 inner lobes bearing setae, 1 lobe absent.
- P1 with a 2-segmented endopod and 3-segmented exopod.
- P5 female absent.
- P5 male uniramous with right leg absent.

[1] Subeucalanus crassus  (Giesbrecht, 1888)   (F,M)    [Figs]

[2] Subeucalanus dentatus  (A. Scott, 1909)   (F)    [Figs]

[3] Subeucalanus flemingeri  Prusova, Al-Yamani & Al-Mutairi, 2001   (F,M)    [Figs]

[4] Subeucalanus longiceps  (Matthews, 1925)   (F,M)    [Figs]

[5] Subeucalanus monachus  (Giesbrecht, 1888)   (F,M)    [Figs]

[6] Subeucalanus mucronatus  (Giesbrecht, 1888)   (F,M)    [Figs]

[7] Subeucalanus pileatus  (Giesbrecht, 1888)   (F,M)    [Figs]

[8] Subeucalanus subcrassus  (Giesbrecht, 1888)   (F,M)    [Figs]

[9] Subeucalanus subtenuis  (Giesbrecht, 1888)   (F,M)    [Figs]

Subeucalanus subtenuis japonica    Fukase, 1957   (F,M)
Syn.: Eucalanus subtenuis japonica Fukase, 1957 (p.19, figs.F,M, Rem.); Tanaka, 1960 (p.21, Rem.); Grice, 1962 (p.181, figs.F,M, Rem.); Vervoort, 1963 b (p.92, Rem.); Eucalanus subtenuis : Tanaka, 1935 a ; Eucalanus mucronatus : Mori, 1937 (1964) (p.25, figs.F,M); Chihara & Murano, 1997 (p.789, Pl.99 part.,101: F,M);
Loc: Mer de Chine E, Japon, Pacif. équatorial
Lg.: (91) F: ± 3; M: 2,9-2,7; (101) F: 3,12-2,4; M: 2,52-2,45; {F: 2,40-3,12; M: 2,45-2,90}

[10] Subeucalanus sp.  Goetze & Ohman, 2010       [Figs]

[11] Subeucalanus sp.2  Goetze & Ohman, 2010       [Figs]

[12] Subeucalanus sp.3  Goetze & Ohman, 2010       [Figs]

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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2021. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed July 26, 2021]

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