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List species and varieties by family |
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| Aetideidae Giesbrecht, 1892 ( Clausocalanoidea ) | |
| | | (1) Aetideopsis Sars, 1903 ( Aetideinae ) | |
| | Rem.: | type: Aetideopsis rostrata Sars,1903.
12 spp. + 1 indet. |
| | [1] Aetideopsis albatrossae Shih & Maclellan, 1981 (F, juv.M) [Figs] | |
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| | [2] Aetideopsis antarctica (Wolfenden, 1908) (F,M) [Figs] | |
|
| | [3] Aetideopsis armata (Boeck, 1872) (F,M) [Figs] | |
|
| | [4] Aetideopsis browsei McKinnon, Corley & Duggan, 2011 (F) [Figs] | |
|
| | [5] Aetideopsis carinata Bradford, 1969 (F,M) [Figs] | |
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| | [6] Aetideopsis cristata Tanaka, 1957 (F,M) [Figs] | |
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| | Aetideopsis divaricata Esterly, 1911 (F) | | Syn.: | ? A. trichechus Vervoort, 1949 (p.7) | | Ref.: | Esterly, 1911 (p.316, Descr.F, figs.F); Sewell, 1948 (p.555, 562); Vervoort, 1963 b (p.124, Rem.); Bradford & Jillett, 1980 (p.19); Markhaseva, 1996 (p.42, Rem.) | | Loc: | Californie (San Diego), Afr. SE (in De Decker & Mombeck, 1964 (p.11) | | Lg.: | (143) F: 3,36 | | Rem.: | Cf. Aetideopsis rostrata |
| | Aetideopsis divergens Tanaka, 1957 (F,M) | | Syn.: | A. rostrata : Tanaka & Omori, 1970 (p.112) | | Ref.: | Tanaka, 1957 a (p.40, figs.F,M); Bradford & Jillett, 1980 (p.19); Markhaseva, 1996 (p.42, Rem.) | | Loc: | Japon ( Izu) | | Lg.: | (39) F: 3,01; M: 3,13 | | Rem.: | Cf. Aetideopsis rostrata |
| | Aetideopsis inflata Park, 1978 (F) | | Ref.: | Park, 1978 (p.118, Descr.F, figs.F); Markhaseva, 1996 (p.42, Rem.) | | Loc: | Antarct. (Pacif.), Namibie (in Carola, 1993) | | Lg.: | (20) F: 3,64 | | Rem.: | Cf. Aetideopsis rostrata |
| | Aetideopsis magna Grice & Hulsemann, 1970 (M) | | Ref.: | Grice & Hulsemann, 1970 (p.188, Descr.M, figs.M); Bradford, 1976 a (p.9); Bradford & Jillett, 1980 (p.19, 74, Rem.) | | Loc: | off Woods Hole S | | Lg.: | (200) M: 4,56-4,2 | | Rem.: | ? Cf. Pseudotharybis magnus |
| | [7] Aetideopsis minor (Wolfenden, 1911) (F) [Figs] | |
|
| | Aetideopsis modesta With, 1915 (F) | | Syn.: | Chiridius modestus With, 1915 (p.82, Descr.F, figs.F) | | Ref.: | Vervoort, 1957 (p.53); Bradford & Jillett, 1980 (p.19); Markhaseva, 1996 (p.36, Rem.) | | Loc: | off Islande | | Lg.: | (7) F: 3,69 | | Rem.: | Cf. Aetideopsis minor |
| | [8] Aetideopsis multiserrata (Wolfenden, 1904) (F,M) [Figs] | |
|
| | Aetideopsis nasuta With, 1915 (F) | | Syn.: | Chiridius nasutus With, 1915 (p.81, Descr.F, figs.F) | | Ref.: | Vervoort, 1957 (p.53); 1963 b (p.123); Bradford, 1972 (p.38); Park, 1978 (p.111); Bradford & Jillett,1980 (p.19); Markhaseva, 1996 (p.37, Rem.) | | Loc: | Atlant. NE (off Ecosse W, off Islande S) | | Lg.: | (7) F: 2,88 | | Rem.: | Cf. Aetideopsis multiserrata |
| | Aetideopsis pacifica Esterly, 1911 (F) | | Ref.: | Esterly, 1911 (p.315, Descr.F, figs.F); Sewell, 1948 (p.555, 562); Vervoort, 1957 (p.53, Rem.); Bradford, 1972 (p.38); Bradford & Jillett, 1980 (p.19); Markhaseva, 1996 (p.37, Rem.) | | Loc: | San Diego (Californie) | | Lg.: | (143) F: 3,18 | | Rem.: | Cf. Aetideopsis multiserrata |
| | Aetideopsis rakuma Zvereva, 1977 (F) | | Ref.: | Zvereva, 1977 (p.6, Descr.F, figs.F); Bradford & Jillett, 1980 (p.19, 21) | | Loc: | Mer d'Okhotsk | | Rem.: | Cf. Bradyidius rakuma |
| | [9] Aetideopsis retusa Grice & Hulsemann, 1967 (F,M) [Figs] | |
|
| | [10] Aetideopsis rhinoceros McKinnon, Corley & Duggan, 2011 (F,M) [Figs] | |
|
| | [11] Aetideopsis rostrata Sars, 1903 (F,M) [Figs] | |
|
| | Aetideopsis trichechus Vervoort, 1949 (juv. F) | | Ref.: | Vervoort, 1949 (p.7, Descr.F, figs.F); 1957 (p.53, Rem.); 1963 b (p.123, Rem.); Tanaka & Omori, 1970 (p.114, Rem.); Bradford & Jillett, 1980 (p.19: comme trichecus ); Markhaseva, 1996 (p.42, Rem.) | | Loc: | Indien E, Pacif. | | Lg.: | (41) F: 3,38 | | Rem.: | Cf. Aetideopsis rostrata |
| | [12] Aetideopsis tumorosa Bradford, 1969 (F,M) [Figs] | |
|
| | [13] Aetideopsis sp. Bradford, 1972 (F) [Figs] | |
|
| | Aetideopsis sp. Unterüberbacher, 1964 (F) | | Ref.: | Unterüberbacher,1964 (p.21, figs.F) | | Loc: | Namibie (Walvis Bay) | | Rem.: | Cf. Aetideopsis carinata | | | (2) Aetideus Brady, 1883 ( Aetideinae ) | |
| | Rem.: | type: Pseudocalanus armatus Boeck,1872. 11 spp.
On the basis of the females Bradford (1971) concluded that the only difference between Snelliaetideus Vervoort, 1949 and Aetidius Brady, 1883 was the absence of postero-lateral thoracic points. According to Roe (1975, p.305) the discovery of A. arcuatus male confirms Bradford’s opinion and SnelliaetideusAetideus. |
| | [1] Aetideus acutus Farran, 1929 (F,M) [Figs] | |
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| | [2] Aetideus arcuatus (Vervoort, 1949) (F,M) [Figs] | |
|
| | [3] Aetideus armatus (Boeck, 1872) (F,M) [Figs] | |
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| | Aetideus armatus Brady, 1883 (F,[non M]) | | Syn.: | Aetidius armatus Brady, 1883 (p.76, figs.F,M); Thompson, 1888 d (p.142) | | Ref.: | Sars, 1901 a (1903) (p.26: Rem.); Esterly, 1905 (p.145, figs.F); Lysholm, 1913 (p.5); Vervoort, 1957 (p.45, Rem.); Chiba & al., 1957 a (p.12) | | Loc: | G. de Guinée, Norvège, Médit. (Malte), Indien, mers de Chine, Californie | | Rem.: | [M] non identifiable; Cf. (part.) Aetideus armatus (F) (Boeck,1872) & ? Aetideus pseudarmatus (F), Aetideus bradyi (F) |
| | Aetideus atlanticus Calef & Grice, 1965 (F) | | Ref.: | Calef & Grice, 1965 (p.634, Descr.F, figs.F); Bradford, 1971 (p.32) | | Loc: | Brésil (NE) | | Rem.: | Cf. Paivella inaciae |
| | [4] Aetideus australis (Vervoort, 1957) (F,M) [Figs] | |
|
| | [5] Aetideus bradyi A. Scott, 1909 (F, ?M) [Figs] | |
|
| | [6] Aetideus divergens Bradford, 1971 (F,M) [Figs] | |
|
| | [7] Aetideus giesbrechti Cleve, 1904 (F,M) [Figs] | |
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| | Aetideus mediterraneus Steuer, 1910 (F) | | Ref.: | Steuer, 1910 a (p.23, Rem.F) | | Loc: | Adriatique (Dubrovnik) | | Rem.: | Espèce douteuse. ? Cf. Aetideus giesbrechti |
| | [8] Aetideus mexicanus Park, 1974 (F,M) [Figs] | |
|
| | [9] Aetideus pacificus Brodsky, 1950 (F,M) [Figs] | |
|
| | [10] Aetideus pseudarmatus Bradford, 1971 (F) [Figs] | |
|
| | Aetideus tenuirostris Wolfenden, 1903 | | Ref.: | Wolfenden, 1904 (p.116, Rem.); Vervoort, 1957 (p.45, Rem.) | | Rem.: | Cf. Aetideus armatus (Boeck,1872) |
| | [11] Aetideus truncatus Bradford, 1971 (F,M) [Figs] | |
|
| | Aetideus sp. Brodsky, 1955 (F) | | Ref.: | Brodsky, 1955 a (p.188, figs.F) | | Loc: | Kouriles-Kamtchatka | | Rem.: | Cf. Bradyidius sp. | | | (3) Azygokeras Koeller & Littlepage, 1976 | |
| | Rem.: | type: Azygokeras columbiae . 1 sp.: |
| | [1] Azygokeras columbiae Koeller & Littlepage, 1976 (F,M) [Figs] | |
| | | (4) Batheuchaeta Brodsky, 1950 ( Euchirellinae ) | |
| | Rem.: | type: Batheuchaeta lamellata Brodsky,1950. 9 spp.: |
| | [1] Batheuchaeta anomala Markhaseva, 1981 (F,M) [Figs] | |
|
| | [2] Batheuchaeta antarctica Markhaseva, 1986 (F) [Figs] | |
|
| | Batheuchaeta brodskyi Markhaseva, 1981 (F) | | Ref.: | Batheuchaeta gurjanova |
| | Batheuchaeta enormis Grice & Hulsemann, 1968 (F) | | Ref.: | Grice & Hulsemann, 1968 (p.323, Descr.F, figs.F, tab.2); Bradford & Jillett, 1980 (p.21); Vaupel Klein, 1984 a (p.50) | | Loc: | Pacif. SE (off Is. Juan Fernandez E) | | Rem.: | Cette forme n'appartient pas à ce genre. Cf.? (voir spp. inc. sedis) |
| | [3] Batheuchaeta gurjanovae (Brodsky, 1955) (F,M) [Figs] | |
|
| | [4] Batheuchaeta heptneri Markhaseva, 1981 (F) [Figs] | |
|
| | [5] Batheuchaeta lamellata Brodsky, 1950 (F,M) [Figs] | |
|
| | [6] Batheuchaeta peculiaris Markhaseva, 1983 (F,M) [Figs] | |
|
| | [7] Batheuchaeta pubescens Markhaseva, 1986 (F) [Figs] | |
|
| | [8] Batheuchaeta tuberculata Markhaseva, 1986 (F) [Figs] | |
| | | (5) Bradyetes Farran, 1905 | |
| | Rem.: | Type: Bradyetes inermis Farran,1905. This genus could be synonym of Pseudeuchaeta (in Sewell, 1947), which is undermined by Grice (1972 a)
6 spp. |
| | Bradyetes brevis Farran, 1936 (F) | | Ref.: | Farran, 1936 b (p.238, Descr.F, figs.F); Sewell, 1948 (p.495); Vervoort, 1952 b (n°43, p.3, figs.F); Markhaseva, 1996 (p.62, 226, Rem.) | | Loc: | Arct. (Groenland E), Mer de Barents, off Cap Cod (S) | | Rem.: | Cf. Jaschnovia brevis |
| | [1] Bradyetes curvicornis Markhaseva & Schulz, 2006 (F) [Figs] | |
|
| | Bradyetes florens Grice & Hulsemann, 1967 (F) | | Ref.: | Grice & Hulsemann, 1967 (p.23, Descr.F, figs.F); Bradford & Jillett, 1980 (p.21, Rem.) | | Loc: | Indien | | Rem.: | Cf. ? Bathypontiidae (voir spp. inc. sedis) |
| | [2] Bradyetes inermis Farran, 1905 (F,M) [Figs] | |
|
| | Bradyetes major Sars, 1907 (F) | | Ref.: | Sars, 1907 a (p.9, Rem.F) | | Rem.: | Cf. Pseudochirella fallax |
| | [3] Bradyetes matthei Johannessen, 1976 (F,M) [Figs] | |
|
| | [4] Bradyetes pacificus Ohtsuka, Boxshall & Shimomura, 2005 (F) [Figs] | |
|
| | [5] Bradyetes weddellanus Markhaseva & Schulz, 2006 (F) [Figs] | |
| | | (6) Bradyidius Giesbrecht, 1897 ( Aetideinae ) | |
| | Rem.: | Type: Bradyidius armatus Giesbrecht,1897. 16 spp. (of which 1 doubtful) + 2 unidentified.
Ten species are compared in Bradford, 1976 (table1, p.6).
Diagnosis after Vives & Shmeleva (2007, p.549) :
- Forehead without crest.
- Rostrum , strong, bifurcate, showing points being or no diverging.
- Cephalosome and 1st thoracic segment fused, segments thoracic 4 and 5 fused..
- Posterior corners of the prosome elongated ending in points.
- A1 as long as the prosome.
- Outer spines of the 2nd exopodal segment of P1 larger than the 1st segment. Endopod with 1 segment showing an outer lobe well developed.
- Endopod of P2 2-segmented, exopod 3-segmented.
- Basipodites of P1-P4 with generalemente groups of spinules on surfaces.
Male:
- Crest absent.
- Rostrum as female, but parallel rami..
A1 whether short and not reaching end of the 2nd thoracic segment or exceeding the caudal rami;
- P5 unramous or biramous; in the first instance both rami are equal, in the second instance, endopod has only 1 segment (rarement 2) and generally styliform. |
| | [1] Bradyidius angustus (Tanaka, 1957) (M) [Figs] | |
|
| | [2] Bradyidius armatus Giesbrecht, 1897 (F,M) [Figs] | |
|
| | [3] Bradyidius arnoldi Fleminger, 1957 (F,M) [Figs] | |
|
| | Bradyidius bradyi Sars, 1884 (F,M) | | Syn.: | Pseudocalanus armatus : Brady,1878 (p.46); Bradyidius armatus Giesbrecht, 1897;Undinopsis bradyi Sars, 1884; 1902 (1903) (p.32) | | Ref.: | Matthews, 1964 (p.18); Markhaseva, 1996 (p.70) | | Rem.: | Cf. Bradyidius armatus |
| | Bradyidius brevispinus Bradford, 1969 (F) | | Ref.: | Bradford, 1969 b (p.479, Descr.F, figs.F); 1976 a (p.9); Bradford & Jillett, 1980 (p.21,74) | | Loc: | Nouvelle-Zélande (NE) | | Rem.: | Cf. Pseudotharybis brevispinus |
| | [4] Bradyidius capax Bradford-Grieve, 2003 (F) [Figs] | |
|
| | [5] Bradyidius curtus Markhaseva, 1993 (F) [Figs] | |
|
| | Bradyidius dentatus Bradford, 1969 (M) | | Ref.: | Bradford, 1969 (p.481, Descr.M, figs.M); 1976 a (p.9); Bradford & Jillett, 1980 (p.74, figs.M) | | Loc: | Nouvelle-Zélande (NE) | | Rem.: | Cf. Pseudotharybis dentatus |
| | [6] Bradyidius hirsutus Bradford, 1976 (F,M) [Figs] | |
|
| | [7] Bradyidius luluae Grice, 1972 (F,M) [Figs] | |
|
| | [8] Bradyidius pacificus (Brodsky, 1950) (F,M) [Figs] | |
|
| | [9] Bradyidius plinioi Campaner, 1978 (F,M) [Figs] | |
|
| | Bradyidius plinioi minor Campaner, 1978 (F) | | Rem.: | Cf. Bradyidius plinioi |
| | Bradyidius plinioi plinioi Campaner, 1978 | | Rem.: | Cf. Bradyidius plinioi |
| | [10] Bradyidius rakuma (Zvereva, 1977) (F) [Figs] | |
|
| | Bradyidius robustus Bradford, 1969 (F,M) | | Ref.: | Bradford, 1969 b (p.476, figs.F,M); Bradford, 1976 a (p.9); Bradford & Jillett, 1980 (p.21,74) | | Rem.: | Cf. Pseudotharybis robustus |
| | [11] Bradyidius saanichi Park, 1966 (F,M) [Figs] | |
|
| | [12] Bradyidius similis (Sars, 1902) (F,M) [Figs] | |
|
| | Bradyidius spinatus Bradford, 1969 (F,M) | | Ref.: | Bradford, 1969 b (p.484 : lapsus calami ); 1976 a (p.9); Bradford & Jillett, 1980 (p.21) | | Rem.: | Cf. Parabradyidius spinibasis |
| | Bradyidius spinibasis Bradford, 1969 | | Syn.: | Bradyidius spinatus Bradford, 1969 b (p.484) | | Ref.: | Bradford, 1969 b (p.481, figs. juv.F,M); Bradford & Jillett, 1980 (p.21, 74, figs.F,M) | | Loc: | Nouvelle-Zélande (N) | | Rem.: | Cf. Parabradyidius spinibasis |
| | [13] Bradyidius spinifer Bradford, 1969 (F,M) [Figs] | |
|
| | [14] Bradyidius styliformis Othman & Greenwood, 1987 (F,M) [Figs] | |
|
| | [15] Bradyidius subarmatus Markhaseva, 1993 (F,M) [Figs] | |
|
| | [16] Bradyidius tropicus Wolfenden, 1905 (F) | |
|
| | Bradyidius sp. Bradford, 1972 (F) | | Ref.: | Bradford, 1972 (p.38, figs.F) | | Lg.: | (116) F: 2,5 | | Rem.: | Cf. Bradyidius spinifer |
| | [17] Bradyidius sp. Brodsky, 1955 (F) | |
|
| | [18] Bradyidius sp. Greenwood, 1977 (F) [Figs] | |
| | | Bryaxis Sars, 1902 ( Aetideinae ) | |
| | Rem.: | Genusname established by Boeck,1872, but not published by this author. Cf. Comantenna (part.) & Paracomantenna (part.) | | | | Bryaxis brevicornis Sars, 1902 (F) | | Ref.: | Sars, 1902 (1903) (p.35, Descr.F, figs.F) | | Loc: | Norvège, Mer de Barents, Irlande | | Rem.: | Cf. Comantenna brevicornis | | | | Bryaxis minor Farran, 1905 (F) | | Ref.: | Farran, 1905 (p.32, Descr.F, figs.F); Bradford & Jillett, 1980 (p.29) | | Loc: | Atlant. NE | | Rem.: | Cf. Paracomantenna minor | | | | (7) Chiridiella G.O. Sars, 1907 | |
| | Rem.: | Type: Chiridiella macrodactyla Sars,1907. 18 spp.:
Diagnosis from Deevey (1974), after Bradford (1971) to, include 16 species..
- Body usually short and thick, with anterior part rather swollen.
- Forehead rounded, no trace of rostrum.
- Head and 1st pedigerous segment separate, 4th and 5th thoracic segments fused.
)- Last metasomal segment rounded.
- Urosome usually very short.
- Genital segment female, strongly protuberant ventrally.
- Caudal rami short.
- A1 of 23 or 24 segments.
- A2 female rather short, exopodite longer than endopodite; male slightly larger than exopodite.
- Md females very large and strongly toothed, palp comparatively short; mandibular blade male reduced and probably useless, palp larger.
- Mx1 with endopodal part of palp little developed in female; reatly reduced in male.
- Mx2 female varyiing in structure from 5 or 6 lobes with spines and setae to 2 or 3 highly modified lobes with hooks and pincers; much reduced in male.
- Mxp with 2nd segment often elongate, terminal part very short.
- P1 with both rami 1-segmented.
- P2-P4 with 2- or 3-segmented exopods, 1- to 3-segmented endopods.
- P5 lacking in female.
- Male P5 as in Aetideidae, 1-segmented endopods larger on left leg. |
| | [1] Chiridiella abyssalis Brodsky, 1950 (F,M) [Figs] | |
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| | [2] Chiridiella atlantica Wolfenden, 1911 (F) [Figs] | |
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| | [3] Chiridiella bichela Deevey, 1974 (F, juv.M) [Figs] | |
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| | [4] Chiridiella bispinosa Park, 1970 (F) [Figs] | |
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| | [5] Chiridiella brachydactyla Sars, 1907 (F,?M) [Figs] | |
|
| | [6] Chiridiella brooksi Deevey, 1974 (F, juv.M) [Figs] | |
|
| | [7] Chiridiella chainae Grice, 1969 (F) [Figs] | |
|
| | [8] Chiridiella gibba Deevey, 1974 (F, juv.M) [Figs] | |
|
| | [9] Chiridiella kuniae Deevey, 1974 (F, juv.M) [Figs] | |
|
| | [10] Chiridiella macrodactyla Sars, 1907 (F) [Figs] | |
|
| | [11] Chiridiella megadactyla Bradford, 1971 (F) [Figs] | |
|
| | [12] Chiridiella ovata Deevey, 1974 (F) [Figs] | |
|
| | [13] Chiridiella pacifica Brodsky, 1950 (F,M) [Figs] | |
|
| | Chiridiella reducta Brodsky, 1950 (F,M) | | Ref.: | Brodsky, 1950 (1967) (p.194, figs.F,[non M], Rem.); Bradford, 1971 b (p.21, Rem.); Vidal, 1971 a (p.17, figs.F, [non M]); Markhaseva, 1983 (p.120, Rem.M) | | Loc: | Arct. (bassin central), Mer de Beaufort E | | Lg.: | (22) F: 2,4; M: 2,7; {F: 2,40; M: 2,70} | | Rem.: | Cf. Chiridiella abyssalis (M). Pour Markhaseva (1996, p.104) la femelle appartiendrait partiellement à Chiridiella pacifica : Cf. Chiridiella reductella |
| | [14] Chiridiella reductella Markhaseva, 1996 (F) [Figs] | |
|
| | [15] Chiridiella sarsi Markhaseva, 1983 (F) [Figs] | |
|
| | [16] Chiridiella smoki Markhaseva, 1983 (F) [Figs] | |
|
| | [17] Chiridiella subaequalis Grice & Hulsemann, 1965 (F) [Figs] | |
|
| | [18] Chiridiella trihamata Deevey, 1974 (juv. F) [Figs] | |
| | | (8) Chiridius Giesbrecht, 1892 ( Aetideinae ) | |
| | Rem.: | Type: Chiridius poppei Giesbrecht,1892. Not all authors admit the synonymy between Chiridius et Pseudaetideus. 8 spp. : |
| | Chiridius armatus Boeck, 1872 (F,M) | | Syn.: | Euchaeta armata Boeck, 1872; Pseudaetideus armatus Wolfenden, 1904 (p.115); van Breemen, 1908 a (p.32); Brodsky, 1950 (1967) (p.157); Vervoort, 1952 c (n°44, p.3); 1963 b (p.125); non C. armatus : Sars, 1900 (p.64) | | Ref.: | Sars, 1901 a (1903) (p.27, figs.F,M); Markhaseva, 1996 (p.27, Rem.) | | Rem.: | Cf. Aetideopsis armata |
| | Chiridius brevicaudatus Sars, 1907 (F) | | Ref.: | Sars, 1907 a (p.7) | | Loc: | Canaries S, G. du Mexique, Féroé, Indien W, Philippines, Chili | | Rem.: | Cf. Gaidius brevicaudatus |
| | Chiridius brevispinus Sars, 1900 (F) | | Syn.: | Gaidius major Wolfenden,1904 (p.114, figs.F) | | Ref.: | Sars, 1900 (p.68, Descr.F, figs.F); Markhaseva, 1996 (p.187, Rem.) | | Loc: | Arct. | | Rem.: | Cf. Gaidius brevispinus |
| | Chiridius carnosus Tanaka, 1957 (M) | | Ref.: | Tanaka, 1957 a (p.55, Descr.M, figs.M); Bradford & Jillett, 1980 (p.23); Markhaseva, 1996 (p.119, Rem.) | | Loc: | Japon (Sagami) | | Rem.: | Cf. Chiridius pacificus |
| | [1] Chiridius gracilis Farran, 1908 (F,M) [Figs] | |
|
| | [2] Chiridius longispinus Tanaka, 1957 (M) [Figs] | |
|
| | [3] Chiridius mexicanus Park, 1975 (F) [Figs] | |
|
| | Chiridius modestus With, 1915 (F) | | Ref.: | With, 1915 (p.82, Descr.F, figs.F); Markhaseva, 1996 (p.32, Rem.) | | Loc: | Atlant. N | | Rem.: | Cf. Aetideopsis minor |
| | [4] Chiridius molestus Tanaka, 1957 (F,M) [Figs] | |
|
| | Chiridius nasutus With, 1915 (F) | | Ref.: | With, 1915 (p.81, Descr.F, figs.F); Tanaka, 1957 a (p.37); Vervoort, 1963 b (p.123, Rem.); Bradford & Jillett, 1980 (p.19); Markhaseva, 1996 (p.32) | | Loc: | Atlant. N | | Rem.: | Cf. Aetideopsis multiserrata |
| | [5] Chiridius obtusifrons Sars, 1902 (F,M) [Figs] | |
|
| | [6] Chiridius pacificus Brodsky, 1950 (F,M) [Figs] | |
|
| | [7] Chiridius polaris Wolfenden, 1911 (F,M) [Figs] | |
|
| | [8] Chiridius poppei Giesbrecht, 1892 (F,M) [Figs] | |
|
| | Chiridius subantarcticus Park, 1978 (F) | | Ref.: | Park, 1978 (p.125, figs.F); Markhaseva, 1996 (p.119, Rem.) | | Loc: | Antarct., sub-Antarct. (Pacif.) | | Lg.: | (20) F: 3,44; 3,36 | | Rem.: | Cf. Chiridius polaris |
| | Chiridius subgracilis Park, 1975 (F,M) | | Ref.: | Park, 1975 b (p.285, figs.F,M); Bradford & Jillett, 1980 (p.23, figs.F,M); Mazzocchi & al., 1995 (p.143, figs.F,M, Rem.); Markhaseva, 1996 (p.113, 114, Rem.); Boxshall & Halsey, 2004 (p.59: fig.M) | | Loc: | G. du Mexique, Mer de Tasman, Nouvelle-Zélande, Détr. de Magellan | | Rem.: | mésopélagique . Les mâles de C. molestus et subgracilis semblent identiques, mais ni Park ni Bradford & Jillett ne font de cette espèce un synonyme junior de la première. Cf. Chiridius molestus |
| | Chiridius tenuispinus Sars, 1900 (F,M) | | Syn.: | Gaidius boreale Wolfenden, 1902 (p.365); G. borealis Wolfenden, 1904 (p.131); Gaetanus tenuispinus : Park, 1975 a (p.11,30, figs.F,M) | | Ref.: | Sars, 1900 (p.67, Descr.F, figs.F); Vervoort, 1957 (p.58) | | Loc: | Arct., Antarct., G. du Mexique | | Rem.: | Cf. Gaetanus tenuispinus |
| | Chiridius vanhöffeni Wolfenden, 1904 (M) | | Syn.: | Pseudocalanus armatus : Vanhöffen,1897 | | Ref.: | Wolfenden, 1904 (p.117, figs.M); van Breemen, 1908 a (p.35, Rem.) | | Lg.: | (229) M: 3,1 | | Rem.: | Cf. Chiridius obtusifrons |
| | Chiridius sp. Tanaka, 1937 (Juv. M) | | Ref.: | Tanaka, 1937 (p.256, figs. juv.5M) | | Loc: | Japon (Suruga Bay) | | Lg.: | (128) juv.M: 2,05 | | Rem.: | Cf. Chiridius gracilis | | | (9) Chirundina Giesbrecht, 1895 ( Euchirellinae ) | |
| | Rem.: | Type: Chirundina streetsii Giesbrecht,1895. 3 spp. + 1 doubtful: |
| | Chirundina abyssalis With, 1915 (F) | | Syn.: | Chirudina abyssalis With, 1915 (p.147, Descr.F, figs.F) | | Loc: | Atlant. N | | Rem.: | Cf. Pseudochirella obtusa |
| | [1] Chirundina alaskaensis (Tanaka, 1969) (F) [Figs] | |
|
| | Chirundina angulata Sars, 1905 (F) | | Ref.: | Sars, 1905 b (p.13) | | Rem.: | Cf. Undeuchaeta major |
| | Chirundina antarctica Wolfenden, 1911 (F) | | Ref.: | Wolfenden, 1911 (p.243, Descr.F, figs.F); Sewell, 1948 (p.574); Park, 1978 (p.176, Rem.); Bradford & Jillett, 1980 (p.27) | | Loc: | Antarct. (continent, secteur Indien) | | Lg.: | (10) F: 6 | | Rem.: | Espèce considérée comme douteuse par certains auteurs. |
| | [2] Chirundina indica Sewell, 1929 (F,M) [Figs] | |
|
| | Chirundina magna Wolfenden, 1911 (F,M) | | Ref.: | Wolfenden, 1911 (p.241, Descr.F, figs.F); Park, 1978 (p.176, Rem.); Bradford & Jillett, 1980 (p.66, Rem.); Markhaseva, 1996 (p.135, Rem.) | | Loc: | Atlant. S | | Rem.: | Cf. Chirundinella magna |
| | Chirundina parvispina Farran, 1908 (juv. M) | | Syn.: | Gaidius parvispinus Farran, 1908 b (p.34, Descr. juv.M, non F, figs. juv.M, non F); non C. parvispina : With, 1915 (p.151, figs.juv., M); Rose, 1933 a (p.107, figs.M); | | Ref.: | Bradford, 1980 (p.66); Vives, 1982 (p.291); Markhaseva, 1996 (p.275) | | Loc: | G. de Gascogne, off Maroc-Mauritanie | | Lg.: | (7) M: 5,28; (24) juv.M: 4,9 | | Rem.: | Cf. Pseudochirella cryptospina |
| | [3] Chirundina streetsii Giesbrecht, 1895 (F,M) [Figs] | |
| | | (10) Chirundinella Tanaka, 1957 ( Euchirellinae ) | |
| | Rem.: | Diagnose after Tanaka (1957, p.197): Female
- Cephalothorax elongate, ovate.
- Head and 1st pediger segment separate, 4th and 5th pediger segments fused.
- Head with a median crest.
- Rostrum one-pointed.
- Abdomen 4-segmented in female.
- Outer marginal seta of caudal rami short; appendicular seta very long.
- A1 24-segmented, segments 8-9 fused, 24-25 separte.
- A2 exopod longer than endopod.
- Md with a small endopod, slightly longer than half the length of exopod; biting part robust.
- Mx1 with 4 setae on 2nd basal segment; 10 setae on endopod.
- Mx2 and Mxp resemble those of Pseudochirella
- P1 exopod 3-segmented, endopod 2-segmented.
P3 and P4 each exopod and endopod 3-segmented.
- P4 with a row of bristles on inner margin of coxa.
P5 female absent.
- Male unknown.
Male describes by Vervoort 1949 as Pseudochirella magna. |
| | Chirundinella cara Tanaka, 1957 (F) | | Ref.: | Tanaka, 1953 (p.131: nom. nud.); 1957 b (p.197, Descr.F, figs.F); Grice & Hülsemann, 1967 (p.23, fig.F, Rem.) | | Loc: | Japon (Izu), Indien W | | Rem.: | Cf. Chirundinella magna |
| | [1] Chirundinella magna (Wolfenden, 1911) (F,M) [Figs] | |
| | | (11) Comantenna Wilson, 1924 ( Aetideinae ) | |
| | Rem.: | type: Bryaxis brevicornis . 5 spp. : |
| | [1] Comantenna brevicornis (Sars, 1902) (F,M) [Figs] | |
|
| | [2] Comantenna crassa Bradford, 1969 (F) [Figs] | |
|
| | [3] Comantenna curtisetosa Alvarez, 1986 (F) [Figs] | |
|
| | [4] Comantenna gesinae Schulz, 2002 (F) [Figs] | |
|
| | [5] Comantenna recurvata Grice & Hülsemann, 1970 (F) [Figs] | |
| | | (12) Crassantenna Cole, Green, Croft & Rawlins, 1972 ( Aetideinae ) | |
| | Rem.: | Type: Crassantenna comosa. The swimming feet are not intact in the two species being the origin of this genus. Total: 2 spp.
Diagnosis from Bradford (1969, p.493) :
Female:
- head and 1st thoracic segment fused (with fusion line visible dorsally), 4yj and 5th thoracic segments fused.
- Posterior borders of prosome up-yurned.
- Genital segment slightly swollen anteriorly;
- Small single rostrum.
- A1 24-segmented, reaching at least thoracic segment 3, and covered with annulate setae.
- A2 with thick squat endopod, 2 setae on segment 2 of exopod; segment 1 of exopod with or without 2 setae.
- Md palp large with small endopod.
- Mx1, inner lobe 1 with 12 spines and setae.
- Mx2 with lobe 5 enlarged.
- Mxp basipod 1 short and thick with no sensory appendage distally.
- P1 endopod 1-segmented.
- P2 endopod 2-segmented, all other rami 3-segmented. External spines on exopods large. Terminal spines on exopods bear numerous sharp teeth. Exopod segments where external spine inserted not produced into spines externally.
- P5 absent.
- Male unknown. |
| | [1] Crassantenna comosa Bradford, 1969 (F) [Figs] | |
|
| | [2] Crassantenna mimorostrata Bradford, 1969 (F) [Figs] | |
| | | Derjuginia Jaschnov, 1947 ( Aetideinae ) | |
| | Rem.: | Cf. Jaschnovia | | | | Derjuginia tolli Linko, 1913 (F,M) | | Syn.: | Scolecithrix tolli Linko, 1913; non. D. tolli : Johnson, 1963 (p.95, figs.F, M) | | Ref.: | Brodsky, 1950 (1967) (p.157, figs.F,M); Shih & al., 1971 (p.201); Vidal, 1971 a (p.17,25,114, figs.F,M); Markhaseva, 1980 a (p.65, figs.F,M) | | Loc: | Arct. | | Rem.: | Cf. Jaschnovia tolli |
| | Euaetideus Sars, 1925 ( Aetideinae ) | |
| | Rem.: | type: Aetideus giesbrechti Clève,1904. Cf. Aetideus | | | | (13) Euchirella Giesbrecht, 1888 ( Euchirellinae ) | |
| | Rem.: | Type: Undina messinensis Claus,1863. 24 spp.
The bathymetric distribution of these forms shows generally a very wide amplitude.
For Tanaka (1957, p/176) the genus is characterized by the head usually separates from 1st thoracic segment; head with or without a crest; last thoracic segments fused; lateral corners of last thoracic segment rounded; rostrum usually present; abdomen 4-segmented in female; genital segment often asymmetrical: caudal rami about as long as wide; A1 23-segmented; endopod of A2 shorter than half the length of exopod; 2nd basal segment of Mx1 long; exopod of P1 2-segmented; exopod of P2 to P4 3-segmented; endopod of P1 and P2 1-segmented; endopod of P3 and P4 3-segmented; coxa of female P4 with spines on the inner margin; P5 absent in female; male P5 biramous on the right; left leg with a rudimentary endopod.
For Vervoort (1957, p.63) there is a great deal of confusion concerning the limitations of the genera Euchirella and Pseudochirella, representatives of which resemble each other in many respects. The menbers of the genus Euchirella (known in 1957), all have a reduced number of setae on the 2nd basal segment and on the endopod of Mx1 (2-3 and 4-6 respectively) ; in Pseudochirella this number is increased (5 and 14-16 respectively). The endopod of A2 is reduced in nearly all Euchirella’s (it is less than half the length of the exopod in many species and about half that length in some others ; it is, however, much less reduced in Pseudochirella. Both Euchirella and Pseudochirella are characterized by the presence of a spine or spinules on the posterior surface of the 1st basal segment of P4. Euchirella brevis Sars (1905), approaches genera Gaetanus by the presence of thin, tube like spinules ; all other species have stronger spinules or a single strong spine. Ann additional difficulty in both genera is the scarcity of adult males and the obstacles encountered when trying to identity both sexes of a species. |
| | Euchirella acadiana Willey, 1919 (F) | | Ref.: | Willey, 1918 (1919) (p.190, figs.F) | | Loc: | Canada (Acadie) | | Lg.: | F: 6,25 | | Rem.: | Cf. Euchirella truncata |
| | Euchirella acuta Tanaka & Omori, 1969 (F,M) | | Ref.: | Tanaka & Omori, 1969 (p.34, figs.F,M) | | Loc: | Pacif. | | Lg.: | F: 4,78; 4,7; M: 3,89-3,85 | | Rem.: | Cf. Euchirella unispina |
| | [1] Euchirella amoena Giesbrecht, 1888 (F,M) [Figs] | |
|
| | Euchirella areata Tanaka, 1957 (F) | | Ref.: | Tanaka, 1957 b (p.186, Descr.F, figs.F); Vaupel Klein, 1984 a (p.36) | | Loc: | Japon (Sagami) | | Rem.: | Cf. Euchirella bella |
| | Euchirella atlantica Wolfenden, 1904 (F) | | Syn.: | E. curticauda atlantica Wolfenden, 1904 (p.116) | | Ref.: | Wolfenden, 1905 a (p.20: Rem.F); van Breemen, 1908 a (p.48, Rem.) | | Rem.: | Cf. Euchirella curticauda |
| | [2] Euchirella bella Giesbrecht, 1888 (F,M) [Figs] | |
|
| | Euchirella bella indica Wolfenden, 1906 | | Ref.: | Wolfenden, 1905 (1906) (p.1006, Descr.F, figs.F); Sewell, 1948 (p.531); Vaupel Klein, 1984 a (p.36) | | Rem.: | Cf. Euchirella bella |
| | [3] Euchirella bitumida With, 1915 (F,M) [Figs] | |
|
| | Euchirella brevis Sars, 1905 (F) | | Ref.: | Wolfenden, 1905 (p.18, figs.F, Rem.); Grice, 1962 a (p.194, Rem.); Madhupratap & Haridas, 1986 (p.105, tab.1) | | Rem.: | Cf. Euchirella amoena |
| | Euchirella carinata Wolfenden, 1902 (F,M) | | Ref.: | Wolfenden, 1902 (p.366, Descr.M); 1904 (p.115, Descr.F) | | Rem.: | Cf. Chirundina streetsii |
| | [4] Euchirella curticauda Giesbrecht, 1888 (F,M) [Figs] | |
|
| | Euchirella curticauda atlantica Wolfenden, 1904 (F) | | Ref.: | Wolfenden, 1904 (p.116); Pearson, 1906 (p.16) | | Rem.: | Cf. Euchirella curticauda |
| | Euchirella dentata A. Scott, 1909 (F) | | Ref.: | A. Scott, 1909 (p.59,Descr.F,figs.F); Bradford & Jillett, 1980 (p.66) | | Loc: | Manipa Strait | | Rem.: | Cf. Pseudochirella dentata |
| | Euchirella dubia A. Scott, 1909 (F) | | Ref.: | A. Scott, 1909 (p.60, figs.F) | | Rem.: | Cf. Pseudochirella obtusa |
| | Euchirella elongata Wolfenden, 1905 (F) | | Ref.: | Wolfenden, 1905 a (p.19, Descr.F, figs.F) | | Rem.: | Cf. Pseudochirella spectabilis |
| | [5] Euchirella formosa Vervoort, 1949 (F,M) [Figs] | |
|
| | [6] Euchirella galeatea Giesbrecht, 1888 (F,M) [Figs] | |
|
| | Euchirella gracilis Wolfenden, 1911 (F) | | Ref.: | Wolfenden, 1911 (p.237, Descr.F, figs.F); Vervoort, 1952 f (n°47, p.4, Rem.); Vaupel Klein,1984 a (p.47) | | Rem.: | Cf. Euchirella truncata |
| | [7] Euchirella grandicornis Wilson, 1950 (F) [Figs] | |
|
| | Euchirella granulata A. Scott, 1909 (F) | | Ref.: | A. Scott, 1909 (p.58, Descr.F, figs.F) | | Rem.: | Cf. Pseudochirella granulata |
| | Euchirella hessei Brady, 1883 | | Syn.: | Euchaeta hessei Brady, 1883 (p.63, figs.F,M); non E. hessei : A. Scott, 1909 (p.54, Rem.) | | Ref.: | Giesbrecht & Schmeil, 1898 (p.36); Sewell, 1929 (p.109: Rem.); 1947 (p.71, Rem.) | | Loc: | Atlant. N | | Rem.: | ? Cf. Euchirella rostrata (part.) |
| | Euchirella hirsuta Wolfenden, 1905 (F) | | Ref.: | Wolfenden, 1905 a (p.17, Descr.F, figs.F) | | Loc: | Antarct. | | Rem.: | Cf. Pseudochirella hirsuta |
| | Euchirella indica Vervoort, 1949 (F,M) | | Syn.: | Vervoort, 1949 (p.23, figs.F,M) | | Ref.: | Vaupel Klein, 1984 a (p.35) | | Rem.: | Cf. Euchirella messinensis indica |
| | Euchirella intermedia With, 1915 (F,M) | | Ref.: | With, 1915 (p.124, figs.F,M); Bradford & Jillett,1980 (p.33); Björnberg & al., 1981(p.632, figs.F); Vaupel Klein, 1984 a (p.47); Greze & al., 1985 (p.7) | | Rem.: | Cf. Euchirella truncata |
| | [8] Euchirella latirostris Farran, 1929 (F,M) [Figs] | |
|
| | [9] Euchirella lisettae Vaupel Klein, 1989 (F) [Figs] | |
|
| | [10] Euchirella maxima Wolfenden, 1905 (F,M) [Figs] | |
|
| | [11] Euchirella messinensis (Claus, 1863) (F,M) [Figs] | |
|
| | Euchirella messinensis indica Vervoort, 1949 (F,M) | | Syn.: | E. messinensis : A. Scott, 1909 (p.56); Tanaka, 1957 b (p.180, figs.F,M); E. indica Vervoort, 1949 (p.23, 28, figs.M); 1963 b (p.134); Grice & Hulsemann, 1968 (tab.2); Bradford & Jillett, 1980 (p.37, figs.F,M); Heinrich, 1990 (p.17); Shih & Young, 1995 (p.67); Hsiao & al., 2004 (p.325, tab.1) | | Ref.: | Tanaka & Omori, 1969 (p.48, figs.F,M, Rem.); 1969 a (p.157); Vaupel Klein, 1972 (p.505, figs.F, Rem.F); 1984 a (p.35, Rem.); Markhaseva, 1996 (p.158, figs.F,M); Chihara & Murano, 1997 (p.685, Pl.37,39: F,M); Mulyadi, 2001 (p.818, figs.F, Rem.) | | Loc: | Afr. S (S), Indien, Mer de Célèbes, mers de Chine, Taiwan E, Japon (Izu), Pacif. (tropical), Nouvelle-Zélande, Pacif. SE (tropic.), off Is. Juan Fernandez | | Lg.: | (37) F: 5,3-3,85; M: 4,2-4; (41) F: 5,05-4,25; M: 3,4; (56) F: 5,26; M: 4,62; (110) F: 4,9-4,4; M: 4,8-3,91; (855) F: 4,5; (866) F: 4,25-5,3; M: 3,4-4,8; { F: 3,85-5,30; M: 3,40-4,80} |
| | Euchirella messinensis messinensis Claus, 1863 (F,M) | | Syn.: | Undina messinensis Claus,1863 (p.187, figs.F,M); Euchirella messinensis : Giesbrecht, 1892 (p.232, 239, 244, figs.F,M); Esterly, 1905 (p.151, figs.F,M); Tanaka & Omori, 1969 (p.51, fig.F,M: a-e); Bradford & Jillett, 1980 (p.39, figs.F,M) | | Ref.: | Vaupel Klein, 1972 (p.504, figs.F); 1982 b (p.6 & suiv.); 1984 a (p.34, 35, Rem.); Markhaseva, 1996 (p.160, figs.F,M); Lapernat, 1999 (p.8); Bradford-Grieve & al., 1999 (p.879, 921, figs.F,M) | | Loc: | Médit., Atlant., Californie | | Lg.: | (37) F: 6,2-4,4; M: 5,46-2,8; (340) F: 4,9; M: 3,9 | | Rem.: | épi-bathypélagique. Cf. Euchirella messinensis |
| | [12] Euchirella orientalis Sewell, 1929 (F,M) [Figs] | |
|
| | [13] Euchirella paulinae Vaupel Klein, 1980 (F) [Figs] | |
|
| | Euchirella plumosa Brady, 1918 (F) | | Ref.: | Brady, 1918 (p.20, Descr.F, fig.F); Sewell, 1948 (p.575); Vaupel Klein, 1984 a (p.40, 45, 46) | | Rem.: | ? Cf. Euchirella latirostris ou E. rostromagna |
| | Euchirella propria Esterly, 1911 (M) | | Ref.: | Esterly, 1911 (p.321, figs.F); Vervoort, 1963 b (p.134, Rem.); Vaupel Klein, 1984 a (p.47) | | Rem.: | Cf. Euchirella truncata |
| | [14] Euchirella pseudopulchra Park, 1976 (F) [Figs] | |
|
| | [15] Euchirella pseudotruncata Park, 1975 (F,M) [Figs] | |
|
| | [16] Euchirella pulchra (Lubbock, 1856) (F,M) [Figs] | |
|
| | Euchirella pulchra pacifica Vaupel Klein, 1984 | | Ref.: | Vaupel Klein, 1984 a (p.44, Rem.) | | Loc: | Indo-Pacif. | | Rem.: | forme intermédiaire entre E. pulchra typica et l’espèce E. pseudopulchra. |
| | Euchirella pulchra typica Vaupel Klein, 1984 | | Ref.: | Vaupel Klein, 1984 a (p.44, Rem.) | | Loc: | Atlant. |
| | [17] Euchirella rostrata (Claus, 1866) (F,M) [Figs] | |
|
| | Euchirella rostrata magna Wolfenden, 1905 (F) | | Ref.: | Wolfenden, 1905 a (p.18); Vaupel Klein, 1984 a (p.44) | | Rem.: | Cf. Euchirella rostromagna |
| | [18] Euchirella rostromagna Wolfenden, 1911 (F,M) [Figs] | |
|
| | [19] Euchirella similis Wolfenden, 1911 (F) [Figs] | |
|
| | Euchirella simplex Esterly, 1911 (F) | | Ref.: | Esterly, 1911 (p.320, Descr.F, figs.F) | | Rem.: | Cf. Euchirella maxima |
| | [20] Euchirella speciosa Grice & Hülsemann, 1968 (F) [Figs] | |
|
| | Euchirella spinosa Wolfenden, 1905 (F) | | Ref.: | Wolfenden, 1905 a (p.119, Descr.F, figs.F) | | Rem.: | Cf. Pseudochirella spinosa |
| | [21] Euchirella splendens Vervoort, 1963 (F,M) [Figs] | |
|
| | Euchirella tanseii Omori, 1965 (F) | | Ref.: | Omori, 1965 a (p.60, Descr.F, figs.F); Tanaka & Omori, 1969 a (p.159, Rem.); Bradford & Jillett, 1980 (p.33, 43: Rem.); Vaupel Klein, 1984 a (p.48, Rem.); Markhaseva, 1996 (p.175, Rem.) | | Loc: | Japon (Sagami) | | Lg.: | (245) F: 4,3 | | Rem.: | ? Cf. Euchirella venusta (forme anormale) |
| | Euchirella trigrada Tanaka, 1957 (F) | | Ref.: | Tanaka, 1957 b (p.184, Descr.F, figs.F); Bradford & Jillett, 1980 (p.32); Vaupel Klein, 1984 a (p.38) | | Loc: | Japon | | Lg.: | (56) F: 5,23 | | Rem.: | Cf. Euchirella formosa |
| | [22] Euchirella truncata Esterly, 1911 (F,M) [Figs] | |
|
| | Euchirella tumida Brady, 1918 (F,M) | | Ref.: | Brady, 1918 (p.20, figs.F,M); Vervoort, 1963 b (p.135: Rem.); 1965 (p.86) | | Rem.: | Cf. (part.) Scolecithrix danae |
| | [23] Euchirella unispina Park, 1968 (F,M) [Figs] | |
|
| | [24] Euchirella venusta Giesbrecht, 1888 (F,M) [Figs] | |
|
| | Euchirella wolfendeni Farran, 1908 (F) | | Ref.: | Farran, 1908 b (p.38, Descr.F, figs.F) | | Rem.: | Cf. Pseudochirella pustulifera |
| | Euchirella sp. Tanaka, 1960 (juv. F) | | Ref.: | Tanaka, 1960 (p.38, figs. juv.F) | | Loc: | Antarct. (Indien) |
| | Euchirella sp. Séret, 1979 (F) | | Ref.: | Séret, 1979 (p.82, figs.F) | | Loc: | Antarct. (Kerguelen) | | Lg.: | (246) F: 3,12 |
| | Euchirella sp. C Vaupel Klein, 1984 | | Syn.: | E. bella (F) : C. B. Wilson, 1950 (p.218, figs.F) | | Ref.: | Vaupel Klein, 1984 a (p.49); 1989 (p.146) | | Rem.: | Cf. Euchirella lisettae. | | | (14) Gaetanus Giesbrecht, 1888 ( Aetideinae ) | |
| | Rem.: | Type: Gaëtanus miles Giesbrecht, 1888 (p.335). Park, 1975 a (p.11-12) joins together the two genera Gaetanus and Gaidius, but does not maintain this fusion in 1978 (p.127-136): Bradford & Jillett (1980) give a definition of the two genera. Markhaseva, 1996 retrieves the synonymy of the two genera, including 2 new species and correcting some synonymies. Provisional total: 25 spp. + 4 unidentified. |
| | Gaetanus affinis Sars, 1905 (F,M) | | Syn.: | Gaidius affinis Sars, 1905 b (p.9, Rem.F); Farran, 1908 b (p.32); Sars, 1925 (p.47, figs.F,M); Jespersen, 1934 (p.60); Sewell, 1948 (p.499, 545, 550, 566, 568); Vervoort, 1957 (p.56, figs.F, Rem.); Fagetti, 1962 (p.18); Vervoort, 1963 b (p.126); Park, 1978 (p.135: Rem.); Bradford & Jillett, 1980 (p.58, figs.F,M); Markhaseva, 1996 (p.187, 189: Rem.) | | Loc: | Sub-Antarct., G. de Guinée, off Açores E, Mer des Sargasses, Détr. de Davis S, Islande S, Irlande W, G. de Gascogne, Gibraltar W, Médit., Philippines, Colombie Britannique, Nouvelle-Zélande, off Galapagos, off Pérou, Chili | | Rem.: | Cf. Gaetanus brevispinus |
| | [1] Gaetanus antarcticus Wolfenden, 1905 (F,M) [Figs] | |
|
| | Gaetanus antarcticus Brady, 1918 (F) | | Ref.: | Brady, 1918 (p.19, figs.F); Sewell, 1948 (p.575); Park, 1975 a (p.10, Rem.); Razouls, 1982 (p.194, Rem.); Razouls, 1994 (p.71, figs.F) | | Loc: | Antarct., Tasmanie | | Lg.: | (247) F: 5 | | Rem.: | non synonyme de l'espèce décrite par Wolfenden en 1905. Si la forme est valide elle devient : Gaetanus bradyi . Pour Park (1975 a) elle serait synonyme de Gaetanus kruppi |
| | [2] Gaetanus armiger Giesbrecht, 1888 (F,M) [Figs] | |
|
| | Gaetanus ascendens Esterly, 1913 (F) | | Ref.: | Esterly, 1913 (p.182, Descr.F, figs.F); Sewell, 1947 (p.56); Park, 1975 a (p.10); Bradford & Jillett, 1980 (p.48) | | Rem.: | Cf. Gaetanus brevicornis |
| | Gaetanus atlanticus Wolfenden, 1904 (F) | | Ref.: | Wolfenden, 1904 (p.132, Rem.); Jespersen, 1934 (p.61); Sewell, 1947 (p.56); Davis, 1949 (p.29: Rem.); Vervoort, 1963 b (p.129); Park, 1975 a (p.10) | | Loc: | off Irlande W, Détroit de Davis S | | Lg.: | (49) F: 4,7-4,5; (229) F: 4,4 | | Rem.: | ? Cf. Gaetanus armiger |
| | Gaetanus boreale Wolfenden, 1902 (F) | | Syn.: | Gaidius borealis : Wolfenden, 1904 (p.131) | | Ref.: | Wolfenden, 1902 (p.365, Rem.F); Vervoort, 1957 (p.58); Park, 1975 a (p.11); Bradford & Jillett, 1980 (p.58); Markhaseva, 1996 (p.221) | | Rem.: | Cf. Gaetanus tenuispinus |
| | [3] Gaetanus brachyurus Sars, 1907 (F,M) [Figs] | |
|
| | Gaetanus bradyi Razouls, 1982 (F) | | Syn.: | G. antarcticus Brady, 1918 (p.19, Descr.F, figs.F), non G. antarcticus Wolfenden, 1905 | | Loc: | Antarct. | | Rem.: | Pour Park (1975 a, p.10): ? Cf. Gaetanus kruppi |
| | [4] Gaetanus brevicaudatus (Sars, 1907) (F) [Figs] | |
|
| | Gaetanus brevicaudatus Wolfenden, 1911 (F) | | Ref.: | Wolfenden, 1911 (p.227, Descr.F, figs.F); Sewell, 1947 (p.58); 1948 (p.523); Park, 1975 a (p.10); Bradford & Jillett, 1980 (p.48); Markhaseva, 1996, p.178, Rem.) | | Loc: | Atlant. (tropical S) | | Rem.: | Cf. Gaetanus antarcticus |
| | [5] Gaetanus brevicornis Esterly, 1906 (F,M) [Figs] | |
|
| | [6] Gaetanus brevirostris (Brodsky, 1950) (F,M) [Figs] | |
|
| | [7] Gaetanus brevispinus (Sars, 1900) (F,M) [Figs] | |
|
| | Gaetanus campbellae Park, 1975 (F,M) | | Ref.: | Park, 1975 a (p.12, Rem.: nom.nov.); Gardner & Szabo, 1982 (p.226, figs.F,M); Markhaseva, 1996 (p.196, Rem.) | | Rem.: | voir à Gaetanus intermedius Campbell,1930. Cf. Gaetanus simplex |
| | Gaetanus caudani Canu, 1896 (juv. M) | | Ref.: | Canu, 1896 (p.422, Descr. juv.M, figs. juv.M); Giesbrecht & Schmeil, 1898 (p.33, Rem.: juv. M); A. Scott, 1909 (p.46, Rem.); Vervoort, 1963 b (p.132, Rem.); Park, 1975 a (p.10, Rem.); Bradford & Jillett,1980 (p.48); Markhaseva, 1996 (p.211) | | Loc: | G. de Gascogne, Atlant.N, Féroé, Indonésie-Malaisie | | Rem.: | Cf. Gaetanus pileatus |
| | Gaetanus clarus Esterly, 1906 (M) | | Ref.: | Esterly, 1906 a (p.57, Descr.M, figs.M); Sewell, 1947 (p.62); Brodsky, 1950 (1967) (p.166, figs.M); Park, 1975 a (p.10, Rem.); Bradford & Jillett, 1980 (p.48); Markhaseva, 1996 (p.211) | | Rem.: | Cf. Gaetanus pileatus |
| | [8] Gaetanus curvicornis Sars, 1905 (F) [Figs] | |
|
| | Gaetanus curvispinus Wilson, 1950 (F,M) | | Ref.: | C.B. Wilson, 1950 (p.229, figs.F,M); Park, 1975 a (p.10) Bradford & Jillett, 1980 (p.48, Rem.); Markhaseva, 1996 (p.187, Rem.) | | Rem.: | sp. décrite très sommairement. Cf. Gaetanus brevicornis |
| | Gaetanus denticulatus Aurivillius, 1898 (F) | | Ref.: | van Breemen, 1908 a (p.38, 220, figs.juv.F) | | Rem.: | sp.non identifiable |
| | [9] Gaetanus divergens Wolfenden, 1911 (F) [Figs] | |
|
| | Gaetanus ferox With, 1915 (M) |
| | Gaetanus hamatus A. Scott, 1909 (F) | | Ref.: | A. Scott, 1909 (p.50, Descr.F, figs.F); Sewell, 1947 (p.58); Park, 1975 a (p.10, 13); Bradford & Jillett, 1980 (p.48); Markhaseva, 1996 (p.187) | | Loc: | Indonésie-Malaisie (Mer de Banda), Pérou (in Björnberg, 1973) | | Rem.: | Cf.: Gaetanus brevicornis |
| | Gaetanus holti Farran, 1905 (F) | | Ref.: | Farran, 1905 (p.33, Descr.F, figs.F); Wolfenden, 1908 (p.31, Rem.): 1911 (p.232, figs.F); Sars, 1925 (p.57); Park, 1975 a (p.10); Bradford & Jillett, 1980 (p.48) | | Loc: | Atlant. | | Rem.: | Cf. Gaetanus latifrons |
| | [10] Gaetanus inermis Sars, 1905 (F) [Figs] | |
|
| | Gaetanus intermedius Campbell, 1930 (F,M) | | Syn.: | Gaetanus simplex Brodsky, 1950 (1967) (p.164 , figs.F,M); Vinogradov, 1968 (1970) (p.44); Vaupel-Klein, 1970 (p.4, figs.F); Shih & al., 1971 (p.143); Deevey & Brooks, 1977 (p.263); Gaetanus campbellae Park, 1975 a (p.12) | | Ref.: | Campbell, 1930 (p.178, Descr.F, figs.F); Sewell, 1948 (p.556); Bradford & Jillett, 1980 (p.48); Gardner & Szabo, 1982 (p.226, figs.F, M); Hattori, 1991 (tab.1, Appendix); Markhaseva, 1996 (p.196, Rem.) | | Loc: | ? Mer des Sargasses, Japon (off Sanriku), Mer d'Okhotsk, Mer de Béring, off Aléoutiennes S, Station 'P', Colombie britannique, Chili | | Rem.: | Hopkins (1985, p.167) et Hopkins & Torres (1988, p.82, tab.2) signalent cette espèce en Antarctique (Croker Passage, Mer de Weddell) sans doute à tort pour Gaidius brevispinus (et in Razouls & al., 2000, p.343, tab. 5, Appendix). Si les deux genres Gaetanus et Gaidius sont réunis, le nom est alors préoccupé (Wolfenden,1905 a) : Cf.: Gaetanus simplex |
| | [11] Gaetanus kruppii Giesbrecht, 1903 (F,M) [Figs] | |
|
| | [12] Gaetanus latifrons Sars, 1905 (F,M) [Figs] | |
|
| | Gaetanus longispinus Wolfenden, 1905 (F) | | Ref.: | Wolfenden, 1905 a (p.7, Descr.F, fig.F); Sars, 1925 (p.57); Park, 1975 a (p.10); Bradford & Jillett, 1980 (p.48) | | Rem.: | Cf. Gaetanus latifrons |
| | Gaetanus major Wolfenden, 1903 (F,M) | | Ref.: | Wolfenden, 1903 (in Fowler, 1903, p.125, Rem.); 1904 (p.114, figs.F); Wolfenden, 1911 (p.231, Rem.); Farran, 1926 (p.249); Sewell, 1947 (p.58); Vervoort, 1963 b (p.126, 127: Rem.); Park, 1975 a (p.10); Bradford & Jillett, 1980 (p.48) | | Lg.: | (250) F: 5,3; (229) M: 4,65 | | Rem.: | Proche de Gaetanus armiger. Cf. Gaetanus kruppi |
| | Gaetanus microcanthus Wilson, 1950 (F,M) | | Ref.: | C.B. Wilson, 1950 (p.231, figs.F,M); Park, 1975 a (p.10); Bradford & Jillett, 1980 (p.48); Markhaseva, 1996 (p.196) | | Loc: | off Pérou, off Galapagos | | Lg.: | (137) F: 3,85; M: 3,25 | | Rem.: | Cf.: Gaetanus kruppii |
| | [13] Gaetanus miles Giesbrecht, 1888 (F,M) [Figs] | |
|
| | [14] Gaetanus minispinus Tanaka, 1969 (F) [Figs] | |
|
| | [15] Gaetanus minor Farran, 1905 (F,M) [Figs] | |
|
| | [16] Gaetanus minutus (Sars, 1907) (F,M) [Figs] | |
|
| | [17] Gaetanus paracurvicornis Brodsky, 1950 (F, juv.M) [Figs] | |
|
| | [18] Gaetanus pileatus Farran, 1903 (F,M) [Figs] | |
|
| | [19] Gaetanus pseudolatifrons Markhaseva, 1996 (F) [Figs] | |
|
| | [20] Gaetanus pungens (Giesbrecht, 1895) (F,M) [Figs] | |
|
| | Gaetanus recticornis Wolfenden, 1911 (F) | | Ref.: | Wolfenden, 1911 (p.228, Descr.F, figs.F); Sewell, 1947 (p.57); C.B. Wilson, 1950 (p.233, figs.F), De Decker & Mombeck, 1964 (p.12); Björnberg, 1973 (p.324, 386); Park, 1975 a (p.10, Rem.: p. 22); 1978 (p.136); Bradford & Jillett, 1980 (p.48); Markhaseva, 1996 (p.211) | | Loc: | Afr. S (E), Atlant. (tropical), off Californie, off Galapagos, off Pérou, Chili | | Lg.: | (10) F: 5-4,6 | | Rem.: | Park (1975 a) estime que cette espèce est synonyme de G. pileatus |
| | Gaetanus rectus Wolfenden, 1911 (F) | | Ref.: | Wolfenden, 1911 (p.232, Descr.F, figs.F); Park, 1975 a (p.10); Markhaseva, 1996 (p.187) | | Rem.: | Cf. Gaetanus brevicornis |
| | [21] Gaetanus robustus Sars, 1905 (F,M) [Figs] | |
|
| | [22] Gaetanus rubellus Markhaseva, 1996 (F) [Figs] | |
|
| | [23] Gaetanus secundus Esterly, 1911 (F,M) [Figs] | |
|
| | [24] Gaetanus simplex Brodsky, 1950 (F,M) [Figs] | |
|
| | [25] Gaetanus tenuispinus (Sars, 1900) (F,M) [Figs] | |
|
| | Gaetanus unicornis Esterly, 1906 (F) | | Ref.: | Esterly, 1906 (p.157, Descr.F, figs.F); Park, 1975 a (p.10); Bradford & Jillett, 1980 (p.48); Markhaseva, 1996 (p.211) | | Rem.: | Cf. Gaetanus pileatus |
| | Gaetanus wolfendeni Park, 1975 (F) | | Syn.: | Gaetanus brevicaudatus Wolfenden, 1911 (p.227, figs.F) | | Rem.: | Park (1975 a, p.12), introduit ce changement de nom du fait de la mise en synonymie de Gaidius avec Gaetanus , ce qui n'est pas suivi par Bradford & Jillett (1980). Cf. Gaetanus antarcticus |
| | [26] Gaetanus sp. Paiva, 1963 (M) [Figs] | |
|
| | Gaetanus sp. Tanaka, 1969 (M) | | Syn.: | ? Gaetanus armiger : Tanaka, 1969 (p.257, Descr.M, figs.M) | | Loc: | off Perou | | Lg.: | (105) M: 2,55 | | Rem.: | Cf. Gaetanus sp.2 Tanaka, 1969 |
| | [27] Gaetanus sp. Tanaka, 1969 (F) [Figs] | |
|
| | [28] Gaetanus sp.2 Tanaka, 1969 (M) [Figs] | |
|
| | [29] Gaetanus sp. Tanaka & Omori, 1970 (M) [Figs] | |
| | | (15) Gaidiopsis A. Scott, 1909 ( Aetideinae ) | |
| | Rem.: | Type: Gaidiopsis crassirostris A. Scott, 1909. Total: 1 sp.
Diagnosis after A. Scott (1909, p.52):
Forehead without cephalic spine, wxith short one-pointed rostrum.
4th and 5th thoracic segments completely separated.
5th segment peoduced into strong lateral spines.
Abdomen 4-segmented.
A1 24-segmented.
A2, Md, Mx1 and Mxp, almost similar to those of Gaidius (= Gaetanus).
Exopodite of P1 distinctly 3-segmented.
P2, P3 and P4 similar to those of Gaidius (= Gaetanus).
Endopodite of P2 distinctly 2-segmented.
Remarks: This new genus resembles Aetideopsis and also Gaidius (= Gaetanus). |
| | [1] Gaidiopsis crassirostris A. Scott, 1909 (F) [Figs] | |
| | | Gaidius Giesbrecht, 1895 ( Aetideinae ) | |
| | Rem.: | type: Gaidius pungens Giesbrecht,1895. Markhaseva (1996, p.176) considers this genus as a junior synonym of Gaetanus. | | | | Gaidius affinis Sars, 1905 (F,M) | | Rem.: | Cf. Gaidius brevispinus | | | | Gaidius boreale Wolfenden, 1902 (F) | | Rem.: | Cf. Gaidius tenuispinus | | | | Gaidius brevicaudatus Sars, 1907 (F) | | Rem.: | Cf. Gaetanus brevicaudatus Sars,1907 non Wolfenden,1911 | | | | Gaidius brevirostris Brodsky, 1950 (F,M) | | Rem.: | Cf. Gaetanus brevirostris | | | | Gaidius brevispinus Sars, 1900 (F,M) | | Rem.: | Cf. Gaetanus brevispinus | | | | Gaidius columbiae Park, 1967 (F,M) | | Rem.: | Cf. Gaidius minutus | | | | Gaidius cryptospinus Sars, 1905 (juv. F) | | Rem.: | Cf. Pseudochirella cryptospina | | | | Gaidius divaricatus Sars, 1905 (F) | | Rem.: | Cf. Pseudochirella divaricata | | | | Gaidius glacialis Brady, 1918 (juv. M) | | Ref.: | Brady, 1919 (p.19, Vervoort, 1951 (p.81, Rem.: juv.M); 1957 (p.58), Park, 1975 a (p.11); Markhaseva, 1996 (p.221) | | Lg.: | (247) Juv.: 2,85 | | Rem.: | Cf. Gaidius tenuispinus | | | | Gaidius inermis Sars, 1905 (F) | | Rem.: | Cf. Gaetanus inermis | | | | Gaidius intermedius Wolfenden, 1905 (F,M) | | Rem.: | Cf. Gaetanus brevispinus | | | | Gaidius major Wolfenden, 1904 (F,M) | | Ref.: | Wolfenden, 1904 (p.114, Rem.F,M); Mazza, 1967 (p.128: Rem.); Park, 1975 a (p.11); Markhaseva, 1996 (p.187) | | Lg.: | (229) F: = 4,65; M: ± 3,1 | | Rem.: | Cf. Gaetanus brevispinus | | | | Gaidius maximus Wolfenden, 1906 (F) | | Rem.: | Cf. Gaetanus robustus | | | | Gaidius minutus Sars, 1907 (F,M) | | Syn.: | Chiridius tenuispinus : Campbell, 1929 (p.311); Gaidius minutus Sars, 1907 a (p.10, Rem.F); 1925 (p.49, figs.F comme G. pusillus ) | | Rem.: | Park (1975 a) et Bradford & al. (1980) ne sont pas en accord sur la synonymie entre cette espËce et G. columbiae. Cf. Gaetanus minutus. | | | | Gaidius moderatus Tanaka, 1957 (F,M) | | Rem.: | Cf. Gaetanus minutus | | | | Gaidius notacanthus Sars, 1905 (F,M) | | Rem.: | Cf. Pseudochirella notacantha | | | | Gaidius parvispinus Farran, 1908 (F) | | Rem.: | Cf. Pseudochirella cryptospina | | | | Gaidius pungens Giesbrecht, 1895 (F,M) | | Rem.: | Cf. Gaetanus pungens. | | | | Gaidius robustus Sars, 1905 (F,M) | | Rem.: | Cf. Gaetanus robustus. | | | | Gaidius robustus Vervoort, 1949 (F) | | Ref.: | Vervoort, 1949 (p.12, Descr.F, figs.F); Grice & Hulsemann, 1967 (p.24, Rem.); Park, 1975 a (p.11); 1978 (p.127); Bradford & Jillett, 1980 (p.58) | | Loc: | Mer de Flores | | Rem.: | Cf. Gaidius brevispinus | | | | Gaidius similis T. Scott, 1894 (M) | | Syn.: | Euchaeta hessei similis T. Scott, 1894 b (p.58, Descr.M, figs.M), non G. similis : A. Scott, 1909 (p.51) | | Ref.: | With, 1915 (p.94, Rem.); Vervoort, 1957 (p.58); Park, 1975 a (p.11, Rem.) | | Loc: | G. de Guinée | | Lg.: | (57) M: 2,2 | | Rem.: | sp. douteuse. | | | | Gaidius tenuispinus Sars, 1900 (F,M) | | Rem.: | Cf. Gaetanus tenuispinus. | | | | Gaidius validus Farran, 1908 (juv. F) | | Rem.: | Cf. Gaetanus robustus | | | | Gaidius variabilis Brodsky, 1950 (F,M) | | Rem.: | Cf. Gaetanus minutus | | | | Gaidius sp. Tanaka, 1969 (F) | | Rem.: | femelle endommagÈe. Cf. Gaetanus sp. | | | | Gaidius sp. Tanaka & Omori, 1970 (M) | | Rem.: | Cf. Gaetanus sp. | | | | Gaidius sp.1 Jaschnov, 1927 | | Loc: | Mer de Kara | | | | Gaidius sp.2 Jaschnov, 1927 | | Loc: | Mer de Kara | | | | (16) Jaschnovia Markhaseva, 1980 | |
| | Rem.: | 2 spp. |
| | [1] Jaschnovia brevis (Farran, 1936) (F,M) [Figs] | |
|
| | Jaschnovia johnsoni Markhaseva, 1980 (F,M) | | Ref.: | Markhaseva, 1980 a (p.68, figs.F,M); 1996 (p.226) | | Rem.: | Cf. Jaschnovia brevis |
| | [2] Jaschnovia tolli (Linko, 1913) (F,M) [Figs] | |
| | | (17) Lutamator Bradford, 1969 ( Aetideinae ) | |
| | Rem.: | Type: Lutamator hurleyi Bradford, 1969. 3 spp.).
Diagnosis from Bradford (1969, p.491) [provisionally until further species are discovered] :
- head and 1st thoracic segment 1fused, 4th and 5th thoracic segments fused.
- Posterior thoracic borders extended into short points .
- Rostrum bluntly rounded.
- Posterior borders of abdominal segments edged with a weakly chitinised striated fringe.
- A1 24-segmented with long antennulate setae.
- A2 endopod and exopod equal; segment 1 of endopod wider proximally than distally; exopod tapers distally, segments 1 and 2 with 1 and 2 setae.
-- Md palp large.
- Mx1 inner lobe 1 (arthrite) with 11 spines and setae.
- Mxp basipod 1 twice as long as wide.
- P1 endopod 1-segmented.
- P2 endopod 2-segmented, all other rami 3-segmented; external spines on exopods large. Teeth on terminal spine of exopods connected by a lamella.
After Markhaseva & Schulz (2008, p.33) female specimens of Prolutamator share with Lutamator Bradford (1969) the following features: 1- rostrum as a poorly developed blunt plate; 2- A2 exopod nearly as long as endopod; 3- Md endopodal segment 2 with fewer than 7 setae; 4- coxal endite of Mxp syncoxa without sensory element; 5- P5 absent.
Except for the above characters (1-4) the genus Lutamator was defined by: 1- A2 endopodal segment 1 wider proximally than distally; 2- A2 exopodal segment 1 with 1 seta; 3- number of setae on the praecoxal arthrite of Mx1 reduced (11 setae). Its definition was considered tentative ''until further species are discovered'' (Bradford, 1969).
The genus was redefined by Alvarez (1984) and two species were included (L. hurleyi, L. elegans). The redefinition in genertal follows the earlier definition, but unfortunately has a misleading sentence that lacks the correct name of the described limb. |
| | [1] Lutamator elegans Alvarez, 1984 (F) [Figs] | |
|
| | [2] Lutamator hurleyi Bradford, 1969 (F) [Figs] | |
|
| | [3] Lutamator paradiseus Ohtsuka, Boxshall & Shimomura, 2005 (F,M) [Figs] | |
| | | (18) Mesocomantenna Alvarez, 1986 | |
| | Rem.: | 1 sp.: |
| | [1] Mesocomantenna spinosa Alvarez, 1986 (F) [Figs] | |
| | | Mesogaidius Wolfenden, 1911 | |
| | Rem.: | Cf. Gaetanus | | | | Mesogaidius intermedius Wolfenden, 1905 (F,M) | | Syn.: | Gaidius intermedius Wolfenden, 1905 a (p.6, figs.F); Markhaseva, 1996 (p.187) | | Rem.: | Cf. Gaetanus brevispinus | | | | Mesogaidius maximus Wolfenden, 1906 (F) | | Syn.: | Gaidius maximus Wolfenden, 1906 (p.35, Descr.F, figs.F) | | Ref.: | Wolfenden, 1911 (p.224, figs.F); Markhaseva, 1996 (p.217) | | Rem.: | Cf. Gaetanus robustus (Sars,1905) |
| | Mesundeuchaeta Wolfenden, 1911 | |
| | Rem.: | Cf. Undeuchaeta | | | | Mesundeuchaeta asymmetrica Wolfenden, 1911 (F) | | Ref.: | Wolfenden, 1911 (p.244, figs.F); Markhaseva, 1996 (p.302) | | Rem.: | Cf. Undeuchaeta incisa | | | | (19) Paivella Vervoort, 1965 ( Aetideinae ) | |
| | Rem.: | type: Paivella inaciae Vervoort,1965. 2 spp.: |
| | [1] Paivella inaciae Vervoort, 1965 (F,M) [Figs] | |
|
| | [2] Paivella naporai Wheeler, 1970 (F) [Figs] | |
| | | (20) Parabradyidius Schulz & Markhaseva, 2000 | |
| | Rem.: | type: Parabradyidius angelikae Schulz & Markhaseva, 2000. Total: 2 spp. (which one juvenil).
Diagnosis from Schulz & Markhaseva (2000, p.78) :
Female:
- Rostrum strongly bifurcate lacking filaments.
- 5th pedigerous somite tapering into long pointed process posteriorly.
- A2 exopod and endopod subequal in length; 1st exopodal segment with 1 seta, 2nd with 3 setae, 7th segment lacking medial seta.
- Md basis bearing 2 setae.
- Mx1 exopod with 11 setae.
- Exopod of P1 3-segmented; outer spine of 1st segment present and more slender than that of 2nd segment.
- Coxa of P1, P2 and P3 with outer patch of spinules.
- Posterior surfaces of P2 to P4 partly covered by spinules.
- P5 present, uniramous, indistinctly 3-segmented; apical segment small bearing spines or spine-like projections.
Male:
- Rostrum similar to that of female bur less diverging distally.
- Posterior corner of 5th pedigerous somite with small pointed process.
- Cephalosomal limbs more or less reduced.
- P5 biramous on both sides, simple, left leg longer than right.
The new genus differs from Aetideopsis, Bradyidius, Pseudotharybis in relative length of A2 endo- and exopod: while in these three genera the exopod is distinctly longer than the endopod (ca. 1.2 - 1.5 times), in Parabradyidius these rami are subequal or the exopod may be even slightly shoryter. In addition, Parabradyidius lacks the medial seta of the A2 7th exopodal segment present in these three taxa. |
| | [1] Parabradyidius angelikae Schulz & Markhaseva, 2000 (F,M) [Figs] | |
|
| | [2] Parabradyidius spinibasis (Bradford, 1969) (juv. F, juv. M) [Figs] | |
| | | (21) Paracomantenna Campaner, 1978 ( Aetideinae ) | |
| | Rem.: | type: Bryaxis minor Farran,1905. Total: 6 spp.: |
| | [1] Paracomantenna goi Ohtsuka, Boxshall & Shimomura, 2005 (F) [Figs] | |
|
| | [2] Paracomantenna gracilis Alvarez, 1986 (F) [Figs] | |
|
| | [3] Paracomantenna magalyae Campaner, 1978 (F) [Figs] | |
|
| | [4] Paracomantenna minor (Farran, 1905) (F,M) [Figs] | |
|
| | [5] Paracomantenna profunda Markhaseva & Renz, 2019 (F) [Figs] | |
|
| | [6] Paracomantenna wishnerae Markhaseva, 1995 (F) [Figs] | |
| | | (22) Prolutamator Markhaseva & Schulz, 2008 | |
| | Rem.: | Type: Prolutamator hadalis Markhaseva & Schulz, 2008. Total: 2 spp.. Total: 3 spp.
Abyssal above the sea bed.
Diagnosis from Markhaseva & Schulz (2008, p.32) :
Female:
- Cephalosome and pediger 1 more or less separate, 4th and 5th separate.
- Posterior corners of prosome short and rounded.
- Rostrum a poorly blunt plate.
- Urosome 4-segmented.
- Genital double-somite symmetrical.
- A1 24-segmented, 1st segment with 1 seta.
- A2 endopod nearly as long as exopod; endopodal segment 1 moderately wide in proximal part bearing 2 setae; basdis and exopodal segment 1 with 2 setae each.
- Md basis with 2-3 setae; endopoda segment 2 with 5-6 setae.
- Mx1 praecoxal endite with 9 terminal, 4 posterior and 1 anterior setae; coxal epipodite with 5-6 setae; distal basal endite with 5 setae; endopod with 14-15 setae.
- Mx2 endopod with 6 long and 2 short setae.
- Mxp proximal praecoxal endite with 1 seta, middle and distal endites with 2 and 3 setae, respecyively; coxal endite without sensory element.
- P1 exopodal segments 1-3 with lateral spine each.
- Segmentation of P1-P4 as typical of Aetideidae.
- P5 absent.
Male unknown. |
| | [1] Prolutamator hadalis Markhaseva & Schulz, 2008 (F) [Figs] | |
|
| | [2] Prolutamator minor Markhaseva & Schulz, 2008 (F) [Figs] | |
|
| | [3] Prolutamator pseudohadalis Markhaseva & Renz, 2019 (F) [Figs] | |
| | | Pseudaetideus Wolfenden, 1904 ( Aetideinae ) | |
| | Rem.: | type: Euchaeta armata (Boeck,1872). For Vervoort (1952) this species is close to Chiridius obtusifrons . Cf. Aetideopsis | | | | (23) Pseudeuchaeta Sars, 1905 ( Aetideinae ) | |
| | Rem.: | type: Autanepsius minor Wolfenden, 1906. 8 spp.
Definition after Bradford & Jillett (1980, p.63) :
- Robust copepods with blunt, rounded rostrum.
- Head and pediger segment 1 partially fused, 4th and 5th pediger segments partially fused.
- Posteolateral corners of last thoracic segment extended into points
- Posterior borders of urosomal segments edged with weakly scleritized fringe.
- Female A1 24-segmented, extending at most length of prosome, bears few sensory filaments and numerous annulate setae, long distally , endopod setae with as in Bradyidius
- A2 exopod and endopod equal with 2 setae on exopodal segments 1 and 2.
- Endopod of Md small.
- Mx1 inner lobe 1 (arthrite) with 11 or 13 spines and setae, posterior surface having 1 or 3 setae.
- Mxp large with sensory organ on basipodal segment 1 (as in Comantenna), endopod setae with crescent-shaped appendages along their length.
- Exopodal segment 1 of P1 without outer edge spine; external exopod spines large.
- Posterior surface of endopods of P3 and P4 with fine spinules.
- Terminal spines of leg exopods with long widely-spaced teeth joined by a lamella.
P5 absent.
Male:
- A1 23-segmented.
- Mouthparts slightly reduced.
- Basipodal segment 1 of Mxp with distal sensory appendage.
- Left P5 with exopod 3-segmented, small lobe on segment 3 lined with spinules, endopod 1-segmented, equals length of exopodal segment 1; right leg exopod 3-segmented, segments 1 and 3 with outer edge spine, segment 3 with long terminal spine extending as sword-like expansion , endopod 1-segmented, club-shaped. |
| | [1] Pseudeuchaeta acuticornis Markhaseva & Schulz, 2006 (F) [Figs] | |
|
| | [2] Pseudeuchaeta arctica Markhaseva, 1986 (F,M) [Figs] | |
|
| | [3] Pseudeuchaeta brevicauda Sars, 1905 (F,M) [Figs] | |
|
| | [4] Pseudeuchaeta flexuosa Bradford, 1969 (F) [Figs] | |
|
| | [5] Pseudeuchaeta magna Bradford, 1969 (F) [Figs] | |
|
| | [6] Pseudeuchaeta major (Wolfenden, 1906) (F) [Figs] | |
|
| | [7] Pseudeuchaeta spinata Markhaseva, 1986 (F) [Figs] | |
|
| | [8] Pseudeuchaeta vulgaris Markhaseva, Mohrbeck & Renz, 2017 (F) [Figs] | |
| | | (24) Pseudochirella Sars, 1920 ( Euchirellinae ) | |
| | Rem.: | type not designated, Pseudochirella obesa proposed by von Vaupel Klein (1984 a, p.52).
Total: 30 spp. + 1 unidentified.
Definition after Bradford & Jillett (1980, p.66) :
Female
- Body resembling Euchirella and Chirundina.
- Head and pediger segment 1 fused, segments 4 and 5 separate or fused.
- Forehead rounded in lateral view, without crest.
- Rostrum distinct, one-pointed.
- Posterolateral corners of last thoracic segment rounded or produced into spines.
- Genital segment asymmetrical in many forms.
- A1 as long as body.
- A2 endopod strongly developed, usually one half length of exopod or slightly longer; exopod segments 1 and 2 separate.
- Mx1 with larger number of setae than in Euchirella: 5 setae on basipodal segment 2; 16 setae on endopod.
- Segments of legs as in Euchirella.
- Exopodal segment 1 of P1 with well developed outer edge spine.
- Coxa of P4 with 6-14 strong spines on posterior surface.
- Setae on outer edge of caudal rami thin and usually elongate.
Male:
- Slender body.
- A1 with reduced number of segments, some setae thickened.
- A2 endopod more than half length of exopod.
- P5 with coxa and basis swollen; endopod elongate, 1-segmented, truncate at apex; left leg exopod 3-segmented, segments short, last segment rounded distally; right leg exopod with 2 elongate segments. |
| | [1] Pseudochirella accepta Zvereva, 1976 (F) [Figs] | |
|
| | [2] Pseudochirella batillipa Park, 1978 (F,M) [Figs] | |
|
| | [3] Pseudochirella bilobata Vervoort, 1949 (F) [Figs] | |
|
| | [4] Pseudochirella bowmani Markhaseva, 1986 (F) [Figs] | |
|
| | [5] Pseudochirella calcarata Sars, 1920 (juv. F) [Figs] | |
|
| | [6] Pseudochirella cryptospina (Sars, 1905) (juv. F) [Figs] | |
|
| | [7] Pseudochirella dentata (A. Scott, 1909) (F) [Figs] | |
|
| | [8] Pseudochirella divaricata (Sars, 1905) (F) [Figs] | |
|
| | [9] Pseudochirella dubia (Sars, 1905) (F,M) [Figs] | |
|
| | Pseudochirella elongata Wolfenden, 1905 (F,M) | | Syn.: | Euchirella elongata Wolfenden, 1905 a (p.19, Descr.F, figs.F); Sewell, 1948 (p.574) | | Ref.: | Sewell, 1948 (p.574); Vervoort, 1957 (p.69); Park, 1978 (p.159, figs.F,M); Markhaseva, 1984 (p.391, figs.F,M); 1989 (p.39, 41); Markhaseva, 1996 (p.289) | | Loc: | Antarct., Arct. | | Rem.: | Cf. Pseudochirella spectabilis |
| | Pseudochirella fallax Sars, 1920 (F,M) | | Syn.: | Bradyetes major Sars, 1907 a (p.9, Rem.F) | | Ref.: | Sars, 1920 c (p.6, Rem.); 1925 (p.95, figs.F, Rem.M); Sewell, 1948 (p.500); Markhaseva, 1989 (p.35, 40, 41, Descr.M, figs. M); Vaupel Klein, 1995 (p.914, Rem.); Markhaseva, 1996 (p.264, figs.F,M) | | Loc: | Atlant. (tempÈrÈ), AÁores, CaraÔbes, Mer des Sargasses, Indien W | | Rem.: | Cf. Pseudochirella major |
| | [10] Pseudochirella formosa Markhaseva, 1989 (F) [Figs] | |
|
| | [11] Pseudochirella gibbera Vervoort, 1949 (F) [Figs] | |
|
| | Pseudochirella granulata A. Scott, 1909 (F) | | Syn.: | Euchirella granulata A. Scott, 1909 (p.58, Descr.F, figs.F) | | Ref.: | Vervoort, 1952 g (n_48, p.4, Rem.); Bradford & Jillett, 1980 (p.66); Markhaseva, 1989 (p.55); 1996 (p.262, Rem.) | | Loc: | Mer de Ceram (Manipa Strait) | | Rem.: | Cf. Pseudochirella dubia |
| | Pseudochirella gurjanovae Brodsky, 1955 (M) | | Ref.: | Brodsky, 1955 a (p.189, Descr.M, figs.M); Bradford & Jillett, 1980 (p.66); Markhaseva, 1986 a (p.841, Rem.) | | Rem.: | Cf. Batheuchaeta gurjanovae |
| | [12] Pseudochirella hirsuta (Wolfenden, 1905) (F,M) [Figs] | |
|
| | [13] Pseudochirella limata Grice & Hulsemann, 1968 (F) [Figs] | |
|
| | [14] Pseudochirella lobata (Sars, 1907) (F) [Figs] | |
|
| | Pseudochirella magna Wolfenden, 1911 (F,M) | | Syn.: | Chirundina magna Wolfenden, 1911 (p.241, Descr.F, figs.F) | | Ref.: | Vervoort, 1949 (p.48, figs.M); Bradford & Jillett,1980 (p.27, 66) | | Rem.: | Cf. Chirundinella magna |
| | [15] Pseudochirella major (Sars, 1907) (F,M) [Figs] | |
|
| | [16] Pseudochirella mariana Markhaseva, 1989 (F) [Figs] | |
|
| | [17] Pseudochirella mawsoni Vervoort, 1957 (F,M) [Figs] | |
|
| | [18] Pseudochirella notacantha (Sars, 1905) (F,M) [Figs] | |
|
| | [19] Pseudochirella obesa Sars, 1920 (F,M) [Figs] | |
|
| | [20] Pseudochirella obtusa (Sars, 1905) (F,M) [Figs] | |
|
| | [21] Pseudochirella pacifica Brodsky, 1950 (F,M) [Figs] | |
|
| | [22] Pseudochirella palliata (Sars, 1907) (F) [Figs] | |
|
| | Pseudochirella polyspina Brodsky, 1950 (F) | | Syn.: | Pseudeuchirella polyspina: Bjˆrnberg, 1973 (p.324, 388) | | Ref.: | Brodsky, 1950 (1967) (p.186, Descr.F, figs.F); Park, 1978 (p.169, figs.F,M); Bradford & Jillett, 1980 (p.66); Gardner & Szabo,1982 (p.252, figs.F); Markhaseva, 1989 (p.36); 1996 (p.278, 281: Rem.) | | Rem.: | Cf. Pseudochirella obtusa |
| | [23] Pseudochirella pustulifera (Sars, 1905) (F,M) [Figs] | |
|
| | [24] Pseudochirella scopularis (Sars, 1905) (F) [Figs] | |
|
| | [25] Pseudochirella semispina Vervoort, 1949 (F) [Figs] | |
|
| | [26] Pseudochirella spectabilis (Sars, 1900) (F,M) [Figs] | |
|
| | Pseudochirella spinifera Brodsky, 1950 (M) | | Ref.: | Brodsky, 1950 (1967) (p.188, figs.M); Park, 1978 (p.169, 171: Rem.); Bradford & Jillett, 1980 (p.66); Markhaseva, 1989 (p.36, Rem.); 1996 (p.278, 281: Rem.) | | Rem.: | Cf. Pseudochirella obtusa |
| | [27] Pseudochirella spinosa (Wolfenden, 1905) (F) [Figs] | |
|
| | Pseudochirella squalida Grice & Hulsemann, 1967 (F,M) | | Ref.: | Grice & Hulsemann, 1967 (p.24, figs.F); Ferrari, 1980 (p.537, figs.F,M, juv.); Bradford & Jillett, 1980 (p.67); Markhaseva, 1989 (p.37); 1996 (p.275) | | Rem.: | Cf. Pseudochirella notacantha |
| | Pseudochirella superba With, 1915 (F,M) | | Syn.: | Undeuchaeta superba With, 1915 (p.137) | | Ref.: | Vervoort, 1952 g (n_48, p.3, fig.M, Rem.F,M); Bradford & Jillett, 1980 (p.80, Rem.); Markhaseva, 1996 (p.302) | | Rem.: | Cf. Undeuchaeta incisa Esterly,1911 |
| | [28] Pseudochirella tanakai Markhaseva, 1989 (F, ?M) [Figs] | |
|
| | Pseudochirella tuberculata Tanaka, 1957 (F,M) | | Ref.: | Tanaka, 1957 b (p.195, Descr.F, figs.F); Roe, 1975 (p.307, figs.M); Bradford & Jillett, 1980 (p.66); Markhaseva, 1996 (p.276) | | Rem.: | Cf. Pseudochirella obesa |
| | Pseudochirella tuberosa Grice & Hulsemann, 1968 (F) | | Ref.: | Grice & H¸lsemann, 1968 (p.327, Descr.F, figs.F); Bradford & Jillett, 1980 (p.66); Markhaseva, 1996 (p.272) | | Loc: | Is. Juan Fernandez | | Rem.: | Cf. Pseudochirella mawsoni |
| | [29] Pseudochirella vervoorti Tanaka & Omori, 1969 (F) [Figs] | |
|
| | [30] Pseudochirella vulgaris Markhaseva, 1997 (F) [Figs] | |
|
| | [31] Pseudochirella sp. Bradford, 1972 (F) [Figs] | |
| | | Pseudogaetanus Brodsky, 1950 ( Aetideinae ) | |
| | Rem.: | type: Pseudogaetanus robustus (Sars,1905). Cf. Gaetanus | | | | Pseudogaetanus robustus Sars, 1905 (F,M) | | Syn.: | Gaetanus robustus Sars, 1905 b (p.11, Rem.F) | | Ref.: | Brodsky, 1950 (1967) (p.169, figs.F,M); Tanaka & Omori, 1967 (p.244) | | Rem.: | Cf. Gaidius robustus (Sars,1905) | | | | (25) Pseudotharybis T. Scott, 1909 ( Aetideinae ) | |
| | Rem.: | Genus excluded from the Tharybidae by Bradford (1976 a) and considered as intermediary between the Aetideidae and the Tharybidae by Markhaseva (1996). Type: Pseudotharybis zetlandicus. Total: 7 spp.
Definition after Bradford & Jillett (1980, p.71) :
- All characteristics as in Bradyidius except that females have P5 3-segmented, terminal segment with 2 terminal and 1 external spine-like extensions.
- Males also almost identical with Bradyidius in having both biramous and uniramous P5, but instead of posteriorly directed spine arising from very posterior last thoracic border; Pseudotharybis males have a small tooth placed more dorsally on last metasomal segment. |
| | [1] Pseudotharybis brevispinus (Bradford, 1969) (F) [Figs] | |
|
| | [2] Pseudotharybis dentatus (Bradford, 1969) (M) [Figs] | |
|
| | Pseudotharybis dubius T. Scott, 1909 (F) | | Ref.: | T. Scott, 1909 a (p.35, Descr.F, figs.F); Bradford, 1976 a (p.9, Rem.: Genre erronÈ) | | Loc: | Irlande N | | Rem.: | Cf.: spp. inc. sedis |
| | [3] Pseudotharybis magnus (Grice & Hulsemann, 1970) (M) [Figs] | |
|
| | [4] Pseudotharybis polaris Markhaseva & Schulz, 2008 (F) [Figs] | |
|
| | [5] Pseudotharybis robustus (Bradford, 1969) (F,M) [Figs] | |
|
| | Pseudotharybis spinibasis Bradford, 1969 (Juv. F & M) | | Syn.: | ? Aetideopsis magna (M) Grice & H¸lsemann, 1970 (p.188, figs.M); Bradyidius spinibasis: Bradford, 1969 b (p.481, figs. juv. F&M); Bradford & Jillett, 1980 (p.21, 74, figs.F,M) | | Loc: | Nouvelle-ZÈlande (N), ? off Woods Hole | | Lg.: | (201) juv.F: 3,35; juv.M: 3,9 | | Rem.: | Cf. Parabradyidius spinibasis |
| | [6] Pseudotharybis zetlandicus T. Scott, 1909 (F) [Figs] | |
| | | (26) Pterochirella Schulz, 1990 | |
| | Rem.: | Type: Pterochirella tuerkayi Schulz, 1990. Total: 1 sp.
This genus shows affinities with Bradyetes and Pseudeuchaeta.
Diagnosis from Schulz (1990, p.182) :
Male :
- Cephalosome and 1st pedigerous somite fused, 4th and 5th pedigerous somite fused.
- Dorsal body wall of prosome transformed into deep transverse cleft encompassed by number of paired lobe-like structures.
- 1st pedigerous somite with paired winglike extension.
- Urosome 5-segmented.
- Rostrum absent, 2 delicate filaments present.
- A2 with endopod equaling exopod in length.
- Mouthparts considerably reduced.
- Endopods of P1 1-segmented, of P2 2-segmented, all other rami 3-segmented.
- P5 biramous with left and right protopod of about equal length, not swollen; endopods 1-segmented; exopods 2- to 3-segmented; distal segment of right side particularly long, tapering distally.
For Schulz (1990, p.187) according to Bradford & Jillett (1980) several features of this genus point to the family Aetideidae. In the male sex: 1 - fusion of cephalosome with 1st pedigerous somite and of somites 4 and 5; 2 - reduction or total lack of a rostrum; 3 - reduced number of segments of A1, swollen proximal segments armed with many large club-shaped aesthetascs; 4 - considerably reduced mouthparts; 5 - segmentation of P1 to P4, and pronounced, acute teeth on terminal spines of P2 to P4; 6 - presence of both endopods on P5. Furthermore, the presence of rows of spinules on the prominent outer lobe of the endopod of P1, described by Vaupel-Klein (1972) for the genus Euchirella, seems to support this classification. Pterochirella differs from both genera in the incomplete fusion of segments 9 and 10, 12 and 13, and complete fusion of segments 24 and 25 of the A1. Segments 20 and 21 are fused on the right but separate on the left A1. Furthermore, Pterochirella lacks the extraordinarily strong spine on lobe 5 of the Maxilla typical of Bradyetes. P5 of Pterochirella is distinguished from those of Bradyetes and Pseudeuchaeta by the comparatively short left protopod and the relative length of the endopods of which the right one is longer than the left. Morever, the right exopod terminates in a more or less straight distal segment, not curved as it is found in the other genera..
Pterochirella is assignable to the subfamily Euchirellinae of Brodsky (1950 [1967]) characterized particularly by 1- the more complex structure of male P5 and 2- the shortened urosome in the female (absent still in 2015). |
| | [1] Pterochirella tuerkayi Schulz, 1990 (M) [Figs] | |
| | | (27) Senecella Juday, 1923 | |
| | Rem.: | freshwater (relict) and brackish. Total: 2 spp.: |
| | [1] Senecella calanoides Juday, 1923 (F,M) [Figs] | |
|
| | [2] Senecella siberica Vyshkvartzeva, 1994 (F,M) [Figs] | |
| | | Snelliaetideus Vervoort, 1949 ( Aetideinae ) | |
| | Rem.: | type: S. arcuatus Vervoort,1949. Cf. Aetideus | | | | Snelliaetideus arcuatus Vervoort, 1949 (F) | | Ref.: | Vervoort, 1949 (p.4, Descr.F, figs.F), Park, 1978 (p.108, figs.F) | | Loc: | IndonÈsie, Antarct., Chili, Mer des Antilles, G. du Mexique | | | | (28) Sursamucro Bradford, 1969 ( Aetideinae ) | |
| | Rem.: | type: Sursamucro spinatus Bradford,1969. Total: 1 sp.
Defnition after Bradford & Jillett (1980, p.77) :
Female:
- Head and 1st pediger segment fused, 4th and 5th pediger segments separate.
- Posterolateral corners of last thoracic segment with points turned dorsally.
- Rostrum bluntly rounded.
- Posterior urosomal borders fringed with well defined teeth.
- A1 24-segmented, bearing long annulate setae with some sensory filaments.
- A2 exopod and endopod equal; exopod with 2 setae on segments 1 and 2.
- Mandibular palp large.
- Mx1 inner lobe 1 (arthrite) with 14 spines and setae.
- Mxp basipodal segment 1 with length twice width.
- P1 exopod with 3 external spines.
- External spines on exopods large on all legs.
- Posterior surface of endopod of P2 to P4 bear fine spinules.
- Terminal spines of P2-P4 exopods with coarse teeth without connecting lamella.
- P5 usually absent.
- Male unknown. |
| | [1] Sursamucro spinatus Bradford, 1969 (F) [Figs] | |
| | | (29) Undeuchaeta Giesbrecht, 1888 ( Euchirellinae ) | |
| | Rem.: | type: Undeuchaeta major Giesbrecht,1888. Total: 5 spp. (of which 1 doubtful) + 1 sp.
Diagnosis after Tanaka (1957 b, p.199) :
- Head and 1st pediger segment fused, 4th and 5th pedigers fused.
- Head with or without crest.
- Posterolateral corners of last thoracic segment bluntly produced in female; rounded in male.
- Rostrum 1-pointed.
- Caudal rami short; appendicular seta short.
- A1 23-segmented.
- Endopod of A2 more than half the length of exopod.
- Mx1 exopod short, number of seta on endopod unreduced.
- Mx2 as in Aetideus.
- Mxp endopod very short.
- P2 endopod 1-segmented.
- P5 absent in female.
- P1 exopod 3-segmented.
- P5 male biramose.
Definition after Bradford & Jillett (1980, p.80):
Female:
- Head and pediger segment 1 indistinctly separate, pediger segments 4 and 5 fused.
- Forehead rounded or crested.
- Posterolateral corners of last thoracic segment more or less asymmetrical, rounded or triangular with rounded apex.
- Genital segment asymmetrical with ventral/or dorsal spine.
- Rostrum small, one-pointed.
- A1 23-segmented.
- A2 endopod more than half length of exopod.
- Mx1 endopod with 15 setae, basipodal segment 2 with 5 setae.
- P1 exopodal segments 1 and 2 partially separate, segment 1 without external spine.
- Endopod of P2 1-segmented.
- Coxa of P4 with or without internal edge spines; if present spines do not protrude posteriorly as in Pseudochirella;
Male:
- Body slender.
- P5 biramous on both sides; basal segments swollen, endopods 1-segmented; left leg exopod 3-segmented, apical segment with styliform distal portion with patch of hairs at midlength.
Diagnosis after Vives & Shmeleva (2007, p.603):
Female:
- Head and 1st pediger fused, sometimes with line of separation, 4th and 5th pedigers fused.
- Prosome longer than 3 times the urosome.
- A1 extenends to end of 3rd urosomal segment.
- Genital segment asymmetrical, sometimes with spicules on the margin posterior dorsal.
- P1 exopod with 2 or 3 segments and P2 to P4 with 3 segments.
- P1 endopod with 1 segment, P2 with 1 or 2 segments and P3-P4 with 3 segments.
-P5 absent.
Male:
- Geenerally without crest, but if exists, very small.
- Segmentation of prosome as in the female.
- Posteroangles of last thoracic segment slightly asymmetrical.
- Urosme 5-segmented.
- P5 biramous, asymmetrical and longer than urosome. the left leg longer than the right; endopod 1-segmented, exopod 3-segmented; right leg exopod 2-segmented. |
| | Undeuchaeta bispinosa Esterly, 1911 (F,M) | | Syn.: | non U. bispinosa (M) : Sewell, 1929 (p.126, figs.M) | | Ref.: | Esterly, 1911 (p.318, Descr.F, figs.F); Sewell, 1929 (part., p.124, figs.F); Sewell, 1947 (p.103); 1948 (p.328, 530, 531, 534, 540, 541, 542, 552, 562); Vervoort, 1957 (p.73, Rem.); Gardner & Szabo, 1982 (p.248, figs.F); Bradford & Jillett, 1980 (p.80); Markhaseva, 1996 (p.301, 302, Rem.) | | Loc: | G. d’Aden, Mer Arabe, Is. Maldives, G. du Bengale, IndonÈsie-Malaisie, off Hokkaido SE, Californie | | Lg.: | (143) F: 4,5 | | Rem.: | Cf. Undeuchaeta intermedia |
| | Undeuchaeta dubia Sars, 1905 (F) | | Ref.: | Sars, 1905 b (p.15, Rem.F); Markhaseva, 1996 (p.262) | | Rem.: | Cf. Pseudochirella dubia |
| | [1] Undeuchaeta incisa Esterly, 1911 (F,M) [Figs] | |
|
| | [2] Undeuchaeta intermedia A. Scott, 1909 (F,M) [Figs] | |
|
| | Undeuchaeta japonica Chiba, 1956 (F) | | Ref.: | Chiba, 1956 (p.48,49, figs.F); Furuhashi, 1965 (p.22) | | Loc: | off Japon | | Lg.: | (536) F: 1,34 | | Rem.: | non Undeuchaeta. Cf.: ? (voir spp. inc. sedis) |
| | Undeuchaeta lobata Sars, 1907 (F) | | Ref.: | Sars, 1907 a (p.11, Rem.F) | | Rem.: | Pseudochirella lobata |
| | Undeuchaeta magna Tanaka, 1957 (F) | | Ref.: | Tanaka, 1957 b (p.203, Descr.F, figs.F); Paiva, 1963 (p.39, figs.F); 1969 (p.266, figs.F); Park, 1978 (p.182, 183, Rem.); Bradford & Jillett,1980 (p.80); Markhaseva, 1996 (p.302) | | Loc: | Japon (Izu), Indien E, Is. du Cap Vert | | Lg.: | (73) F: 6,29; (103) F: 6,03 | | Rem.: | ? Cf. Undeuchaeta incisa |
| | [3] Undeuchaeta major Giesbrecht, 1888 (F,M) [Figs] | |
|
| | Undeuchaeta minor Giesbrecht, 1888 (F,M) | | Syn.: | Undeuchaeta minor Giesbrecht, 1888; 1892 (p.228, 232, figs.F) | | Ref.: | Willey, 1918 (p.187, figs.M); Vervoort, 1957 (p.71); Park, 1978 (p.182); Bradford & Jillett, 1980 (p.80); Markhaseva, 1996 (p.310) | | Rem.: | Cf. Undeuchaeta plumosa |
| | Undeuchaeta obtusa Sars, 1905 (F) | | Ref.: | Sars, 1905 b (p.13, Rem.F) | | Loc: | Atlant., Médit. | | Rem.: | Cf. Pseudochirella obtusa |
| | Undeuchaeta palliata Sars, 1907 (F) | | Ref.: | Sars, 1907 a (p.11, Rem.F) | | Loc: | Açores | | Rem.: | Cf. Pseudochirella palliata |
| | [4] Undeuchaeta plumosa (Lubbock, 1856) (F,M) [Figs] | |
|
| | Undeuchaeta pustulifera Sars, 1905 (F) | | Ref.: | Sars, 1905 b (p.14, RemF) | | Rem.: | Cf. Pseudochirella pustulifera |
| | Undeuchaeta scopularis Sars, 1905 (F) | | Ref.: | Sars, 1905 b (p.14, Rem.F) | | Rem.: | Cf. Pseudochirella scopularis |
| | Undeuchaeta spectabilis Sars, 1900 (F,M) | | Ref.: | Sars, 1900 (p.59, figs.F,M) | | Rem.: | Cf. Pseudochirella spectabilis |
| | Undeuchaeta superba With, 1915 (F,M) | | Ref.: | With, 1915 (p.137); Sewell,1948 (p.564, Rem.); Bradford & Jillett, 1980 (p.80); Vaupel Klein, 1984 a (p.54); Markhaseva, 1996 (p.302) | | Rem.: | non Cf. Pseudochirella superba ; Cf. Undeuchaeta incisa |
| | [5] Undeuchaeta sp. Mulyadi, 2004 (F) [Figs] | |
| | | Undinopsis Sars, 1902 ( Aetideinae ) | |
| | Rem.: | type: Undinopsis bradyi Sars,1902 Cf. Bradyidius | | | | (30) Valdiviella Steuer, 1904 (? Aetideidae, Aetideinae ) | |
| | Rem.: | Type: Valdiviella oligarthra Steuer,1904. Total: 7 spp.
For Tanaka (1957 b, p.205) the genus Valdiviella was created by Steuer (1904) to accomodate a new species (V. oligarthra which is closely related to Euchaeta and also to some genera of Aetideidae.
According to Steuer, the genus has the mandible similar in structure to those of Gaetanus and Chiridius; Mx1 as in Undeuchaeta; Mx2 as in Aetideus, thus the genus Valdiviella is regarded as a member of the family Aetideidae, whereas Sars (1925, p.98) doubts of this justification. A. Scott states that ''the genus is closely related to Euchaeta, but easily recognised by its bifurcate rostrum; the incomplete segmentation of the exopodites of P1 and P2, and indistinctly 3-segmented endopodites of P3 and P4''.
- The genus has general appearance as in Euchaeta; head and 1st thoracic segment fused, 4th and 5th thoracic segments fused; posterior margin of last thoracic segment rounded; abdomen 4-segmented; appendicular seta short; endopod of A2 about as long as exopod; endopod of Md about as long as exopod;
- Mx1 differs from that of Euchaeta : setae on endopod and 2nd basal segment much reduced, endopod with only 3 setae, 2nd basal with 3 setae whereas that of Euchaeta with 3+3+3 setae on endopod, and 5 on 2nd basal segment;
- Mx2 resembles that of Aetideidae, setae on 5th lobe strong; Mxp resembles that of Undeuchaeta and Euchaeta, but row of spinules on anterior margin of 2nd basal segment are situated on the proximal half, wherea, it is on the distal half in Euchaeta, and thet are feeble.
- P1 as in Euchirella : 1st and 2nd exopodal segments fused; endopod of P2 1-segmented, that of P3 2-segmented; P4 exopod and endopod 3-segmented each; P5 absent in female; male P5 resemble those of Chirundina. The genus has a closer relationship to Aetideidae : rostrum resembles that of Aetideopsis or Undinopsis in its bifurcate shape;
Mx1 has the reduced number of setae on endopod and on 2nd basal segment.
- Mx2 as those of Aetideus in having 1 strong seta on 5h lobe (= endite).
- P1 similar to those of Euchirella.
- P2 as in Euchirella or Undeuchaeta.
- The genus has resemblance to Euchaeta: general appearance, structure of A1, Md and Mxp. Thus Valdiviella has the generic characters common both Aetideidae and Euchaetidae; it is intermediate form between Undeuchaeta and Euchaeta. For the author it seems reasonable, as Steuer stated, to accomodate it in the family Aetideidae.
For Vervoort (1957, p.87) the genus, in many respects, takes an intermediate position between the Aetideidae and the Euchaetidae. It approaches the Aetideidae in the structure of the rostrum, swimming legs, in the case of males, the structure of P5. The structure of Mxp shows unmistakable affinities with the Euchaetidae.
The belonging of this genus to this family is discussed by Heptner (1971) and by Zvereva (1975).
Definition after Bradford & Jillett (1980, p.83) Female:
- General appearance as in Euchaeta.
- Rostrum bifurcate.
- Head and pediger segment 1 fused, 4th and 5th pediger segments fused..
- Posterolateral corners of last thoracic segment rounded.
- A1 23-segmented (as in Euchaeta).
- A2 exopod and endopod equal in length.
- Mx1 with reduced setattion on inner lobe 1 (arthrite), endopod and basipodal segment 2 (11, 3, 3 setae respectively); posterior surface of inner lobe 1 with only 1 seta.
- Mx2 as in Aetideus, seta on lobe 5 strong.
- Mxp as in Undeuchaeta and Euchaeta, but row of spinules on anterior margin of basipodal segment 2 situated proximally not distally as in Euchaeta
- Swimming legs with some degree of fusion between proximal segments.
- P5 absent. |
| | [1] Valdiviella brevicornis Sars, 1905 (F,M) [Figs] | |
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| | [2] Valdiviella brodskyi Zvereva, 1975 (F) [Figs] | |
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| | Valdiviella gigas Brady, 1883 (M) | | Syn.: | Euchaeta gigas Brady, 1883 (p.65) | | Ref.: | Sewell, 1929 (p.142, figs. Juv., Rem.: p.146); Vervoort, 1957 (p.86); Bradford & Jillett, 1980 (p.83) | | Rem.: | Cf. Valdiviella oligarthra |
| | [3] Valdiviella ignota Sewell, 1929 (M) [Figs] | |
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| | [4] Valdiviella imperfecta Brodsky, 1950 (F) [Figs] | |
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| | [5] Valdiviella insignis Farran, 1908 (F,M) [Figs] | |
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| | [6] Valdiviella minor Wolfenden, 1911 (F,M) [Figs] | |
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| | [7] Valdiviella oligarthra Steuer, 1904 (F,M) [Figs] | |
| | | Wilsonidius Tanaka, 1969 ( Aetideinae ) | |
| | Rem.: | type:Wilsonidius alaskaensis Tanaka,1969. Cf.: Chirundina | | | | Wilsonidius alaskaensis Tanaka, 1969 (F) | | Syn.: | W. alaskensis Tanaka, 1969 (p.268, Descr.F, figs.F) | | Ref.: | Bradford & Jillett, 1980 (p.92) | | Loc: | G d'Alaska (cÙte) | | Rem.: | Cf. Chirundina alaskaensis | | |
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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2024. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed November 21, 2024] © copyright 2005-2024 Sorbonne University, CNRS
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