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List species and varieties by family |
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| Oncaeidae Giesbrecht, 1893 ["1892"] ( Cyclopoida ) | |
| | | (1) Archioncaea Böttger-Schnack & Huys, 1997 | |
| | Rem.: | Total: 1 sp.
For Böttger-Schnack & Schnack (2013, p.18) the genus has a unique leg armature and combines primitive and advanced characters, which separates this taxon from all other known oncaeids. Most remarkable is the trisetose P5 exopod. On all four swimming legs, the middle and distal segments of the endopods are completely fused and the leg armature (Table 5, type 4) is characterized by the absence of an outer subdistal spine on the fused endopodal segments of P3 and P4 and showing only 2 outer spines on exopodite 3 of P2. |
| | [1] Archioncaea arabica Böttger-Schnack & Huys, 1997 (F) [Figs] | |
| | | (2) Conaea Giesbrecht, 1891 | |
| | Rem.: | type: Conäa rapax Giesbrecht,1891. 4 spp.
For Böttger-Schnack & Schnack (2013, p.19) within Conaea s.L. two subgroups can be distinguished (Table 3): the type species of the genus, C. rapax, (defined as subgroup Conaea s.str.) differs from all other Conaea species (difined as expressa-subgroup) by a different endopodal spine count on P2 (Table 5, type 3), and is also separated by details of the lateral elements on enddopodite 2 of A2, especially the lack of the typical Conaea-like spinulose ornamentation of antennary elements and a very small seta IV. Further differences are found in the proportional lengths of the urosomites. |
| | [1] Conaea expressa (Gordeeva, 1973) (F,M) [Figs] | |
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| | Conaea gracilis Dana, 1849 (F,M) | | Syn.: | Antaria gracilis Dana, 1849; 1852; ? non Conäa rapax Giesbrecht, 1891; 1892 (p.605); ? non Oncaea gracilis T. Scott, 1894 b (p.116, figs.F); Malt, 1982 (p.193); 1983 a (p.7); non Conaea gracilis : A. Scott, 1909 (p.244); Wilson, 1942 a (p.178); 1950 (p.191, figs.F,M); Owre & Foyo, 1964 a (p.369); 1967 (p.112); Boxshall, 1977 a (p.145); 1977 b (p.549) | | Ref.: | De Decker & Mombeck, 1964 (p.12, 18); Owre & Foyo, 1967 (p.112, figs.F,M); De Decker, 1968 (p.45); Binet & Dessier, 1968 (p.7); Furuhashi, 1966 (p.313); Deevey, 1971 (p.224); Heron, 1977 (p.86, 90, Rem.); Deevey & Brooks, 1977 (tab.2); Frontier, 1977 a (p.17); Vives, 1982 (p.296); Rudyakov, 1982 (p.209) | | Loc: | Afr. S (E & W), off Mauritanie, off Bermuda, Madagascar (Nosy Bé), Indien SW, Japon, Kuroshio zone | | Lg.: | (139) F: 1,05-0,88; M: 0,86-0,76; {F: 0,88-1,05; M: 0,76-0,86} | | Rem.: | espèce mal décrite. Voir remarques à Conaea rapax. ? Cf. Conaea rapax ou Oncaea gracilis |
| | [2] Conaea hispida Heron, 1977 (F,M) [Figs] | |
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| | [3] Conaea rapax Giesbrecht, 1891 (F,M) [Figs] | |
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| | [4] Conaea succurva Heron, 1977 (F) [Figs] | |
| | | (3) Epicalymma Heron, 1977 | |
| | Rem.: | Malt (1982, p.185) regards this genus as a synonym of Oncaea, contrary to Heron & al. (1984) and Heron & Bradford-Grieve (1995).
Type: Epicalymma schmitti. 7 spp.:
For Böttger-Schnack & Schnack, (2013, p.19) within the genus, one species, E. umbonata, may deserve a separate group status, based on morphological grounds (labral structure, antennary and maxillipedal elements) as discussed by Böttger-Schnack (2009), but not shown in detail in the present study. hence, the species has been subsumed under Epicalymma S.l. (in Table 3, p.12) to denote the potential existence of two lineages within this genus. |
| | [1] Epicalymma ancora (Gordeeva, 1973) (F) [Figs] | |
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| | [2] Epicalymma brittoni Heron, English & Damkaer, 1984 (F) [Figs] | |
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| | [3] Epicalymma bulbosa Böttger-Schnack, 2009 (F,M) [Figs] | |
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| | [4] Epicalymma exigua (Farran, 1908) (F,M) [Figs] | |
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| | [5] Epicalymma schmitti Heron, 1977 (F,M) [Figs] | |
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| | [6] Epicalymma umbonata Heron, 1977 (F,M) [Figs] | |
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| | [7] Epicalymma vervoorti Heron, English & Damkaer, 1984 (F,M) [Figs] | |
| | | (4) Monothula Böttger-Schnack & Huys, 2001 | |
| | Rem.: | type: Oncaea subtilis Giesbrecht,1892. Total: 1 sp. |
| | [1] Monothula subtilis (Giesbrecht, 1892) (F,M) [Figs] | |
| | | Myspictosum Kazatchenko & Andreev, 1977 | |
| | Rem.: | Cf. Oncaea | | | | (5) Oncaea Philippi, 1843 | |
| | Rem.: | Malt (1982, p.191, 192) regards the 2 genera Conaea and Epicalymma as synonyms of Oncaea, whereas Heron & al. (1984) and Heron & Bradford-Grieve (1995) believe that they should be maintained.
Type: Oncaea venusta Philippi,1843. Total: 71 spp. + 1 unidentified.:
After Ohtsuka & Nishida (2017, p.582), Oncaea abd related genera are associated with discarded appendicularian houses, feeding on plankters retained on the appendicularian feeding nets (Ohtsuka & Kubo, 1991; Ohtsuka & &l., 1993; 1996a). |
| | [1] Oncaea africana Shmeleva, 1979 (F) [Figs] | |
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| | [2] Oncaea alboranica Shmeleva, 1979 (F) [Figs] | |
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| | Oncaea ancora Gordeeva, 1973 (F) | | Rem.: | Cf. Epicalymma ancora |
| | [3] Oncaea anglica Brady, 1905 (F) | |
| | Rem.: | species doubtful. |
| | Oncaea antarctica Heron, 1977 | | Ref.: | Heron, 1977 (p.42, figs.F,M) | | Rem.: | Cf. Triconia antarctica |
| | [4] Oncaea atlantica Shmeleva, 1967 (F,M) [Figs] | |
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| | [5] Oncaea bathyalis Shmeleva, 1968 (F) [Figs] | |
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| | [6] Oncaea bispinosa Böttger-Schnack, 2002 (F,M) [Figs] | |
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| | Oncaea borealis Sars, 1918 (F,M) | | Syn.: | Oncaea conifera : Sars, 1900 (p.113, figs.F) | | Ref.: | Sars, 1918 (p.191, figs.F,M) | | Rem.: | Cf. Triconia borealis |
| | [7] Oncaea bowmani Heron, 1977 (F) [Figs] | |
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| | [8] Oncaea brocha Heron, 1977 (F,M) [Figs] | |
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| | [9] Oncaea brodskii Shmeleva, 1968 (F,M) [Figs] | |
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| | Oncaea canadensis Heron & Frost, 2000 (F,M) | | Ref.: | Heron & Frost, 2000 (p.1019, figs.F,M, tab.2, 3) | | Ref. compl.: | Nishibe & Ikeda, 2004 (Rem. p.933) | | Rem.: | Cf. Triconia canadensis |
| | [10] Oncaea clevei Früchtl, 1923 (F,M) [Figs] | |
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| | [11] Oncaea compacta Heron, 1977 (F,M) [Figs] | |
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| | Oncaea conifera Giesbrecht, 1891 (F,M) | | Syn.: | Oncäa conifera Giesbrecht, 1891; 1892 (p.591, 603, 774, figs.F,M); non O. conifera : Sars, 1900 (p.113); non Oncaea conifera var. III Giesbrecht, 1902 (p.41, Pl.13, figs.7-11); non O. conifera : Tanaka, 1960 (p.66, figs.F,M); B.D. Lee, 1966; non Oncaea conifera : Farran, 1936 a (p.127, figs.F: formes b & c); non O. conifera : Moulton, 1973 (p.147 & suiv.:'minus'); | | Rem.: | Cf. Triconia conifera |
| | Oncaea conifera furcula Farran, 1936 (F,M) | | Ref.: | Farran, 1936 a (p.127, figs.F); Sewell, 1947 (p.260, Rem.); Gardner & Szabo, 1982 (p.116, figs.F,M); Malt, 1983 (p.457) | | Rem.: | Cf. Triconia furcula |
| | [12] Oncaea convexa Heron, 1977 (F) [Figs] | |
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| | [13] Oncaea cristata Böttger-Schnack, 2005 (F,M) [Figs] | |
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| | [14] Oncaea crypta Böttger-Schnack, 2005 (F,M) [Figs] | |
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| | [15] Oncaea curta Sars, 1916 (F,M) [Figs] | |
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| | [16] Oncaea curvata Giesbrecht, 1902 (F,M) [Figs] | |
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| | [17] Oncaea damkaeri Heron, 1977 (F,M) [Figs] | |
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| | [18] Oncaea delicata Heron, English & Damkaer, 1984 (F,M) [Figs] | |
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| | Oncaea dentipes Giesbrecht, 1891 (F,M) | | Syn.: | Oncäa dentipes Giesbrecht, 1891; 1892 (p.591, 603, 774, Descr.F, figs.F) | | Rem.: | Cf. Triconia dentipes |
| | Oncaea derivata Heron & Bradford-Grieve, 1995 (F,M) | | Syn.: | Oncaea conifera : Moulton, 1973 (p.142 & suiv.: 'bumped') | | Ref.: | Heron & Bradford-Grieve, 1995 (p.25, Descr.F, figs.F); Heron & Frost, 2000 (p.1028, Descr.M, figs.F,M, tab.2, 3) | | Rem.: | Cf. Triconia derivata |
| | [19] Oncaea englishi Heron, 1977 (F,M) [Figs] | |
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| | Oncaea exigua Farran, 1908 (F,M) | | Ref.: | Farran, 1908 b (p.93); Malt, 1983 a (p.7, figs.F,M, Rem.); Greze & al., 1985 (p.8) | | Rem.: | Cf. Epicalymma exigua |
| | Oncaea expressa Gordeeva, 1973 (F,M) | | Ref.: | Gordeeva, 1973 (p.1573, figs.F,M); Malt, 1982 (p.192, 193) | | Loc: | off I. Ste-Hélène W | | Rem.: | Cf. Conaea expressa |
| | [20] Oncaea frosti Heron, 2002 (F,M) [Figs] | |
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| | Oncaea furcula Farran, 1936 (F,M) | | Syn.: | Oncaea conifera var. furcula Farran, 1936 a (p.127, figs.F: forme b); Bradford, 1972 (part., p.50); Oncaea conifera : Moulton, 1973 (p.142 & suiv.: 'Long' form); Oncaea furcula : Heron & Bradford-Grieve, 1995 (p.24, Redescr.F,M, figs.F,M) | | Rem.: | Cf. Triconia furcula |
| | [21] Oncaea furnestini Shmeleva, 1979 (F) [Figs] | |
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| | [22] Oncaea glabra Heron & Frost, 2000 (F,M) [Figs] | |
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| | Oncaea gracilis Dana, 1849 (F) | | Syn.: | Antaria gracilis Dana,1849; Giesbrecht, 1892 (p.591: sp. douteuse, 592) | | Ref.: | Malt, 1983 a (p.7, figs.F,M, Rem.); Böttger-Schnack, 1994 (p.277); Chihara & Murano, 1997 (p.980, Pl.221: F,M) | | Rem.: | La synonymie entre Conaea rapax Giesbrecht,1891 et cette forme insuffisamment décrite n'est pas admise par tous les auteurs. Giesbrecht (1892, p.592) estime qu'il s'agit d'une variété de Oncaea venusta . Voir remarques à Conaea gracilis et C. rapax |
| | [23] Oncaea grossa Heron & Frost, 2000 (F,M) [Figs] | |
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| | Oncaea hawii Böttger-Schnack & Boxshall, 1990 (F,M) | | Syn.: | Oncaea sp. Böttger-Schnack, 1988; Böttger-Schnack & Boxshall, 1990 (p.865, Descr.F, figs.F, tab.III) | | Rem.: | Cf. Triconia hawii |
| | [24] Oncaea heronae Malt, 1982 (F) [Figs] | |
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| | Oncaea hispida Malt, 1982 (F,M) | | Ref.: | Malt, 1982 (p.187, figs.F,M) | | Loc: | off NE Is. du Cap Vert | | Lg.: | (680) F: 0,56; M: 0,55 | | Rem.: | Heron & Bradford-Grieve (1995, p.46) ne considèrent pas la forme décrite par Malt (1982) comme synonyme de Conaea hispida Heron, 1977 |
| | [25] Oncaea illgi Heron, 1977 (F,M) [Figs] | |
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| | [26] Oncaea infantula Gordejeva, 1972 (F,M) [Figs] | |
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| | Oncaea inflexa Heron, 1977 (F,M) | | Ref.: | Heron, 1977 (p.45, figs.F,M); Malt,1983 (p.458, Rem.,); Heron & al., 1984 (p.465); Heron & Bradford-Grieve, 1995 (p.15, figs.F,M) | | Rem.: | Cf. Triconia inflexa |
| | [27] Oncaea insolita Heron & Frost, 2000 (F,M) [Figs] | |
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| | Oncaea ivlevi Shmeleva, 1966 (F,M) | | Ref.: | Shmeleva, 1966 (p.932, figs.F,M); Böttger Schnack, 2003 (p.192, Rem.) | | Rem.: | Cf. Spinoncaea ivlevi. |
| | Oncaea ivlevi 1 | | Ref.: | Böttger-Schnack, Hagen & Schnack-Shiel,2001 (p.1029, tab.1,2) | | Loc: | G. d'Aqaba, Mer Rouge (N-S) |
| | Oncaea ivlevi 2 | | Ref.: | Böttger-Schnack, Hagen & Schnack-Shiel,2001 (p.1029, tab.1,2) | | Loc: | Mer Rouge (N-S) |
| | [28] Oncaea lacinia Heron, English & Damkaer, 1984 (F,M) [Figs] | |
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| | [29] Oncaea latimana Gordejeva, 1975 (F,M) [Figs] | |
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| | [30] Oncaea longipes Shmeleva, 1968 (F,M) [Figs] | |
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| | [31] Oncaea longiseta Shmeleva, 1968 (F,M) [Figs] | |
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| | [32] Oncaea macilenta Heron, 1977 (F,M) [Figs] | |
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| | [33] Oncaea media Giesbrecht, 1891 (F,M) [Figs] | |
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| | Oncaea media major Sewell, 1947 (F,M) | | Ref.: | Sewell, 1947 (p.261, Rem.F); 1951 (p.375, Rem.); Malt, 1983 a (p.5, 7); Böttger-Schnack & al., 1989 (p.1089); 1995 (p.92) 1997 (p.409); Ohtsuka & al., 1996 a (p.91); Suarez-Morales & Gasca, 1998 a (p.112) | | Ref. compl.: | Lakkis & al., 2005 (p.152) | | Lg.: | (11) F: 0,817-0,733; (46) F: 0,82-0,55; (180) F: 0,76-0,7; (788) F: 0,77-0,65; {F: 0,55-0,82} |
| | Oncaea media minor Sewell, 1947 (F,M) | | Syn.: | ? Oncaea sp.1 Ferrari, 1975 (p.228, figs.) | | Ref.: | Sewell, 1947 (p.261, Rem.F); 1951 (p.375, Rem.); Malt,1983 a (p.5, 7); Böttger-Schnack & al., 1989 (p.1089); Huys & Boxshall, 1991 (p.292, 304, 311, figs.F,M); Böttger-Schnack, 1992 (p.304); Ohtsuka & al., 1996 a (p.91); Böttger-Schnack, 1997 (p.409); Boxshall & Halsey, 2004 (p.614: fig.M) | | Lg.: | (11) F: 0,65-0,58; M: 0,683-0,533; (180) F: 0,65-0,61 (non ! 1,65- 1,61); (788) F: 0,52-0,48; M: 0,44-0,41; {F: 0,48-0,65; M: 0,41-0,68} | | Rem.: | souvent confondu avec Oncaea scottodicarloi. |
| | [34] Oncaea mediterranea (Claus, 1863) (F,M) [Figs] | |
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| | [35] Oncaea memorata Gordejeva, 1973 (F,M) [Figs] | |
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| | [36] Oncaea minima Shmeleva, 1968 (F,M) [Figs] | |
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| | [37] Oncaea minor Shmeleva, 1979 (M) [Figs] | |
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| | Oncaea minuta Giesbrecht, 1892 | | Syn.: | Oncäa minuta Giesbrecht, 1892 (p.591, 603, 774, Descr.F, figs.F); | | Rem.: | Cf. Triconia minuta |
| | [38] Oncaea mollicula Gordejeva, 1975 (F,M) [Figs] | |
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| | [39] Oncaea neobscura Razouls, 1969 (F) [Figs] | |
| | Rem.: | Malt, 1982 a (p. 140) considers that it concerns a juvenile stage 5 of a male. |
| | [40] Oncaea notopus Giesbrecht, 1891 (F, ?M) [Figs] | |
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| | Oncaea obtusa Dana, 1852 (F,M) | | Ref.: | Brady, 1883 (p.120, figs.F,M); Grindley & Lane, 1979 (p.113); Kovalev & Shmeleva, 1982 (p.86) | | Lg.: | (135) F: 1,3 | | Rem.: | Cf. Oncaea venusta |
| | [41] Oncaea obscura Farran, 1908 (F,M) [Figs] | |
| | Rem.: | Malt (1982 a) believes that it concerns male juvenile forms (stage 5).
After Shiganova & al. (2012, p.99) this species new in the Black Sea (as other species) arise from a rise of the water surface layer temperature and/or increasing of shipping activity during the last decades. |
| | [42] Oncaea oceanica Gordeeva, 1972 (F,M) [Figs] | |
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| | [43] Oncaea olsoni Heron, 1977 (F,M) [Figs] | |
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| | [44] Oncaea ornata Giesbrecht, 1891 (F,M) [Figs] | |
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| | [45] Oncaea ovalis Shmeleva, 1966 (F,M) [Figs] | |
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| | [46] Oncaea parabathyalis Böttger-Schnack, 2005 (F,M) [Figs] | |
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| | [47] Oncaea paraclevei Böttger-Schnack, 2001 (F) [Figs] | |
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| | Oncaea paraobscura Shmeleva, 1979 | | Ref. compl.: | Razouls & Durand, 1991 (p.46) | | Loc: | Médit. | | N: | 1 | | Rem.: | espèce douteuse, non retenue |
| | [48] Oncaea parila Heron, 1977 (F,M) [Figs] | |
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| | Oncaea parobscura Shmeleva, 1979 (F) | | Ref.: | Shmeleva, 1979 (p.494, Descr.F, figs.F) | | Ref. compl.: | Kovalev & Shmeleva, 1982 (p.86); Greze & al., 1985 (p.8); Uysal & al., 2002 (p.18, tab.1) | | Loc: | Médit. (Mer d'Alboran, Bassin Levantin) | | N: | 3 | | Lg.: | (683) F: 0,36 | | Rem.: | Malt, 1982 a (p.140) estime que cette espèce est une forme juvénile mâle. |
| | [49] Oncaea petila Heron, 1977 (F) [Figs] | |
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| | [50] Oncaea philippinensis (Kazatchenko & Andreev, 1977) (M) | |
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| | [51] Oncaea platysetosa Boxshall & Böttger, 1987 (F) [Figs] | |
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| | Oncaea praeclara Humes, 1988 (F,M) | | Ref.: | Humes, 1988 (p.475, figs.F,M) | | Ref. compl.: | Suarez-Morales & Gasca, 1998 a (p.112) | | Loc: | Galapagos Rift, Pacif. Rise E, G. de Californie (Guaymas Basin) | | N: | 1 | | Lg.: | (699) F: 1,3-1,01; M: 0,98-0,91; {F: 1,01-1,30; M: 0,91-0,98} | | Rem.: | Cette espèce a été prélevée à très grande profondeur( 2000 à 2635 m) au moyen des submersibles 'Alvin' et 'Cyana' sur des sites d'hydro-thermalisme du Pacifique Est. Leurs dimensions sont parmi les plus grandes chez les Oncaea . Pour Böttger-Schnack, 2001(p.30, 52) cette espèce est synonyme de Oncaea venusta. |
| | [52] Oncaea prendeli Shmeleva, 1966 (F,M) [Figs] | |
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| | [53] Oncaea prolata Heron, 1977 (F,M) [Figs] | |
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| | [54] Oncaea pumilis Heron, 1977 (F,M) [Figs] | |
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| | Oncaea quadrata Heron & Bradford-Grieve, 1995 (F,M) | | Ref.: | Heron & Bradford-Grieve, 1995 (p.21, figs.F,M) | | Rem.: | Cf. Triconia quadrata |
| | Oncaea redacta Heron & Bradford-Grieve, 1995 (F,M) | | Syn.: | Oncaea conifera : Farran, 1936 a (p.127, figs.F: forme c); Bradford, 1972 (part., p.50); O. conifera : Moulton, 1973 (p.147 & suiv.:'minus'); | | Ref.: | Heron & Bradford-Grieve, 1995 (p.29, figs.F, non M); Heron & Frost, 2000 (p.1032, Redescr.M, figs.M, tab.2, 3) | | Rem.: | Cf. Triconia redacta |
| | [55] Oncaea rimula Heron & Frost, 2000 (F,M) [Figs] | |
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| | [56] Oncaea rotata Heron & Frost, 2000 (F,M) [Figs] | |
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| | [57] Oncaea rotunda Heron, 1977 (F,M) [Figs] | |
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| | [58] Oncaea rotundata Boxshall, 1977 (F,M) [Figs] | |
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| | Oncaea rufa Boxshall & Böttger, 1987 (F) | | Ref.: | Boxshall & Böttger, 1987 (p.553, Descr.F, figs.F) | | Rem.: | Cf. Triconia rufa |
| | Oncaea schmitti Heron, 1977 (F,M) | | Syn.: | Epicalymma schmitti Heron, 1977 (p.82) | | Ref.: | Malt, 1982 (p.190, figs.F, Rem.) | | Rem.: | Cf. Epicalymma schmitti |
| | [59] Oncaea scottodicarloi Heron & Bradford-Grieve, 1995 (F,M) [Figs] | |
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| | [60] Oncaea serrulata Böttger-Schnack, 2011 (F,M) [Figs] | |
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| | [61] Oncaea setosa Heron, 1977 (F,M) [Figs] | |
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| | [62] Oncaea shmelevi Gordejeva, 1972 (F,M) [Figs] | |
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| | Oncaea similis Sars, 1918 (F,M) | | Ref.: | Sars, 1918 (p.193, Descr.F, figs.F); Rose, 1933 a (p.303, figs.F) | | Rem.: | Cf. Triconia similis |
| | Oncaea subtilis Giesbrecht, 1892 (F,M) | | Syn.: | Oncäa subtilis Giesbrecht, 1892 (p.591, 603, Descr.F, figs.F) | | Rem.: | Cf. Monothula subtilis |
| | Oncaea succurva Heron, 1977 (F) | | Syn.: | Conaea succurva Heron, 1977 (p.90) | | Ref.: | Malt, 1982 (p.192, 193, Rem.) | | Rem.: | Cf. Conaea succurva |
| | [63] Oncaea tenella Sars, 1916 (F) [Figs] | |
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| | Oncaea tenella var. pacifica Olson, 1949 (F,M) | | Ref.: | Böttger-Schnack, 2002 (p.1120, Rem.: p.1125) | | Rem.: | Cf. Oncaea zernovi |
| | [64] Oncaea tenuimana Giesbrecht, 1891 (F,M) [Figs] | |
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| | Oncaea thoresoni Heron & Frost, 2000 (F,M) | | Ref.: | Heron & Frost, 2000 (p.1026, figs.F,M, tab.2, 3) | | Rem.: | Cf. Triconia thoresoni |
| | [65] Oncaea tregoubovi Shmeleva, 1968 (F,M) [Figs] | |
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| | Oncaea umbonata Heron, 1977 (F) | | Syn.: | Epicalymma umbonata Heron, 1977 (p.84) | | Ref.: | Malt, 1982 (p.191, 193) | | Rem.: | Cf. Epicalymma umbonata |
| | Oncaea umerus Böttger-Schnack & Boxshall, 1990 (F) | | Syn.: | Oncaea sp. Böttger-Schnack, 1988, 1990 | | Ref.: | Böttger-Schnack & Boxshall, 1990 (p.861, Descr.F, figs.F, tab.III); | | Rem.: | Cf. Triconia umerus |
| | [66] Oncaea venella Farran, 1929 (F,M) [Figs] | |
| | Rem.: | Farran (1929) characterises this variety by its dimensions. Tanaka (1960, p.72) observed in his material a small form (0/90-1.00 mm) corresponding to the venella form. The proportional lengths of the anterior and posterior regions of the body female are as 60 to 40. The anterior region about 1.6 times as long as broad (61 : 39), this is more slender than in forma typica (= O. venusta. The posterior region has the segments in the proportional lengths 10 : 45 : 7 : 7 : 9 : 22 = 100. The caudal rami 4 times as long as broad. No structural differences were observed between the two forms venusta and venella.
Heron (2002) considers this variety as a true species (after essentially its dimensions and some minor morphological details), and distinguishes it as a new species O. frosti (see remark for this species).
After Heron (2002, p.150) this is the first time that Oncaea venusta var. venella is elevated to the species level. The difference in size specifications has caused some confusion for investigators. The smaller size reported in Farran (1936) is considered the most authentic, since it was based on additional material. Because specimens have been found conforming to Farran’s size range from three different areas, the smaller size for O. venella appears to be confirmed.
Its presence in the eastern Pacific is confirmed; however, Böttger-Schnack & Huys (2004) contest the geographic localisations of this form which is not always validly identified, and even the existence of this species that corresponds only to a simple form.
See remarks in Oncaea venusta.
After Böttger-Schnack & Schnack ( 2013, p. 7, Table 2) the species from Heron (2002) is insufficiently described .
Observed in the ballast waters of ships in San Francisco. |
| | [67] Oncaea venusta Philippi, 1843 (F,M) [Figs] | |
| | Rem.: | Epi- to mesopelagic, also demersal and semi-parasitic.
Sampling depth (Antarct., sub-Antarct.) : 400 m. Overall depth Range in Sargasso Sea: 0-500 m (Deevey & Brooks, 1977, Station "S"). 0-1236 m at Station T-1 (E Tori Is, E middle Japan) from Furuhashi (1966 a).
This species presents 2 forms (sub-species?) that are characterised by their dimensions. Sewell (1947) suggests also that their spawning season is different. For this author the forma typica is characterized by: The proportional lengths of the anterior and posterior regions of the body as 59 : 41, and the length and breadth of the anterior region are as 59 to 29.. The proportional lengths of the segments of the posterior region (5th thoracic segment to caudal rami) as 13 : 46: 7: 7: 9 : 18. In the forma venellaThe proportional lengths of the anterior and posterior regions of the body as 60 to 40, and the length and breadth of the anterior region as 60 to 24. The proportional lengths of the segments of the posterior region (5th thoracic segment to caudal rami) as 13 : 43 : 7 : 8 : 9: 20 = 100. The proportions of the caudal rami are as 23 to 6.
For Tanaka (1960, p.71) the two forrms : large and small, are present in his material. Female: The proportional lengths of the anterior and posterior regions of the body are as 57 to 43. The anterior region is 1.7 times as long as wide (63 : 37). The proportional lengths of the segments of the posterior region (caudal rami included) as 13 : 48 : 5 : 5 : 9 : 20 = 100. Caudal rami 4 times as long as broad.
Heron (2002) considers the two varieties O. venusta typica and O. venusta venella as two species (see remarks in Oncaea venella and frosti), position objected to by Böttger-Schnack & Huys (2004).
After KazKazmi (2004, p.232), although free living and pelagic, the species has been found on fish gills (Kazatchenko & Adeev, 1977) or in a sponge (Ho, 1984) and now from the sand (Kazmi & Naushaba, 2000).
After Benedetti & al. (2018, p.1, Fig.2) this species belonging to the functional group 5 corresponding to small sac-spawning detritivorous.
See in DVP Conway & al., 2003 (version 1) |
| | Oncaea venusta typica Farran, 1929 (F,M) | | Syn.: | Oncaea venusta : Farran, 1929 (p.284) | | Ref.: | Farran, 1929 (p.284); Sewell, 1947 (p.263, Rem.F); 1951 (p.371, Rem.: parasites); Corral Estrada, 1970 (p.217); Boxshall, 1977 a (p.127, Rem.); 1977 b (p.548); Malt, 1983 (p.454, figs.F); 1983 a (figs.F,M, Rem., p.5, 7); Ho, 1984 (p.42, Rem.); Böttger-Schnack & al., 1989 (p.1089); Böttger-Schnack, 1994 (p.277); 1995 (p.92); Ohtsuka & al., 1996 a (p.91); Chihara & Murano, 1997 (p.981, Pl.223: F,M); Nakata & al., 2001 (p.335, tab.4, 5, 7); Böttger-Schnack, 2001 (p.31, 46, figs.F,M, Rem.); Rezai & al., 2004 (p.490, tab.2); Böttger-Schnack & Huys, 2004 (p.1, Rem.); Böttger-Schnack & al., 2004 (p.1130, tab.1, Rem.) | | Ref. compl.: | Böttger-Schnack & al., 2001 (p.1029, tab.1); Razai & al., 2004 (p.490, tab.2) | | Lg.: | (11) F: 1,25-1,185; (139) F: ± 1,13; (788) F: 1,23-1; M: 0,95-0,88; (819) F: 1-1,2; M: 0,88-0,95; (833) F: 1,23-1,09; M: 0,98-0,89; {F: 1,00-1,25; M: 0,88-0,95} | | Rem.: | Cette variété est caractérisée d’après Farran (1929) par ses dimensions: F = 1,08 -1,16 mm; M = 0,78-0,81 mm |
| | Oncaea venusta “robust form” | | Ref.: | Boxshall, 1977 a (p.127, fig.F, Rem.); 1977 b (p.546) | | Lg.: | (139) F: 1,04-0,88; {F: 0,88-1,04} |
| | Oncaea venusta venella Farran, 1929 | | Ref.: | Farran, 1929 (p.210, 284, fig.F); Boxshall, 1977 a (p.127: Rem.); Böttger-Schnack & al., 2004 (p.1130, tab.1, Rem.) | | Loc: | Atlant. (tropical), Pacif. SW, Grande Barrière) | | Lg.: | (35) F: 1,07-0,92 | | Rem.: | Cf. Oncaea venella. Voir remarques in Böttger-Schnack & Huys, 2004 (p. 2 & suiv.) |
| | [68] Oncaea vodjanitskii Shmeleva & Delalo, 1965 (F,M) [Figs] | |
|
| | [69] Oncaea waldemari Bersano & Boxshall, 1994 (F,M) [Figs] | |
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| | [70] Oncaea walleni Heron, 1977 (F) [Figs] | |
|
| | [71] Oncaea zernovi Shmeleva, 1966 (F,M) [Figs] | |
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| | [72] Oncaea sp.1 [Heron & Frost, 2000] (F,M) [Figs] | |
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| | Oncaea sp.1 Vilela, 1968 | | Ref.: | Vilela, 1968 (p.33, figs. Juv.F) | | Loc: | Portugal | | Lg.: | (327) juv.F: 0,99-0,81 |
| | Oncaea sp.2 Vilela, 1968 (M) | | Ref.: | Vilela, 1968 (p.33, figs.M) | | Loc: | Portugal | | Lg.: | (327) M: 0,87 |
| | Oncaea sp. A Razouls, 1974 (F) | | Ref.: | Razouls, 1972 (p.95, Annexe: p.118, figs.F); 1974 b (p.241, figs.F) | | Loc: | Médit. (G. du Lion: Banyuls) | | Rem.: | Cf. Oncaea minuta |
| | Oncaea sp.1 Ferrari, 1975 (F) | | Ref.: | Ferrari, 1975 (p.228, figs.F); Boxshall, 1977 a (p.143); Böttger-Schnack, 2001 (p.29, Rem.) | | Loc: | G. du Mexique | | Lg.: | (687) F: 0,71-0,68 | | Rem.: | Cf. ? Oncaea media ou curta |
| | Oncaea sp.2 Ferrari, 1975 (F) | | Ref.: | Ferrari, 1975 (p.228, figs.F); Boxshall, 1977 a (p.143); Böttger-Schnack, 2001 (p.29, Rem.) | | Loc: | G. du Mexique | | Lg.: | (687) F: 0,58-0,54 | | Rem.: | Cf. ? Oncaea curta ou venusta venella |
| | Oncaea sp. K Böttger-Schnack, Hagen & Schnack-Schiel, 2001 | | Ref.: | Böttger-Schnack & al., 2001 (p.1029, tab.1, 2) | | Loc: | G. d'Aqaba, Mer Rouge (N-S) |
| | Oncaea 'zernovi' Böttger-Schnack, Hagen & Schnack-Schiel, 2001 | | Ref.: | Böttger-Schnack & al., 2001 (p.1029, tab.1, 2) | | Loc: | G. d'Aqaba, Mer Rouge (N-S) | | | (6) Spinoncaea Böttger-Schnack, 2003 | |
| | Rem.: | type: Oncaea ivlevi. Total: 3 spp.
For Böttger-Schnack & Schnack (2013, p.19) the genus displays intrageneric variation in the spine count on the exopod segment of P2 (Table 5, type 6). Similar variation has also been found in the longipes-group and the zernovi -group (Table 5: type 5 and type 7a, respectively). In allthese cases, however, all the other morphological characters of the species, including details of ornamentation, were similar (unlike the situation within Conaea s.l.), indicating that they are closelt related sibling/sister species within the same group. Hence, differences in exopodal spine counts on the distal segment of P2 alone are not regaeded as an indication for defining separate subgroups within any group defined in the present study. |
| | [1] Spinoncaea humesi Böttger-Schnack, 2003 (F,M) [Figs] | |
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| | [2] Spinoncaea ivlevi (Shmeleva, 1966) (F,M) [Figs] | |
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| | Spinoncaea ivlevi "robust form" | | Ref.: | Böttger-Schnack, 2003 (p.204, figs.F) |
| | Spinoncaea ivlevi "elongate form" | | Ref.: | Böttger-Schnack, 2003 (p.204, figs.F) |
| | [3] Spinoncaea tenuis Böttger-Schnack, 2003 (F,M) [Figs] | |
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| | Spinoncaea tenuis "typical form" | | Ref.: | Böttger-Schnack, 2003 (p.222, Rem.) |
| | Spinoncaea tenuis "elongate form" | | Ref.: | Böttger-Schnack, 2003 (p.222, Rem.) | | | (7) Triconia Böttger-Schnack, 1999 | |
| | Rem.: | type: Oncaea conifera Giesbrecht,1891. Total: 26 spp. + 1 indet.
Generic name derived from the Latin tres, tria, meaning three and conus, meaning cone, and refers to the presence of a conical projection on the distal endopodal segments of the last three pairs of swimming legs.
Böttger-Schnack (1999) established the genus Triconia which is composed of all oncaeids having a large conical process on the distal margin of the endopod of swimming legs 2-4.
This genus was subdivided into 3 subgroups: similis, conifera, dendipes.
After reexamination, the similis sub-group includes 2 groups, which can be distinguished by the proportional spine lengths on the endopods of swimming P2: 1- containingT. minuta, T. umerus, T. gonopleura, T. parasimilis, present a relatively short distal endopodal spine: 2- including T. similis, T. hawii, T. recta, T. denticulata, with a relatively long distal endopodal spine. |
| | [1] Triconia antarctica (Heron, 1977) (F,M) [Figs] | |
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| | [2] Triconia borealis (Sars, 1918) (F,M) [Figs] | |
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| | [3] Triconia canadensis (Heron & Frost, 2000) (F,M) [Figs] | |
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| | [4] Triconia conifera (Giesbrecht, 1891) (F,M) [Figs] | |
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| | [5] Triconia constricta Wi, Böttger-Schnack & Soh, 2012 (F,M) [Figs] | |
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| | [6] Triconia denticula Wi, Shih & Soh, 2011 (F) [Figs] | |
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| | [7] Triconia dentipes (Giesbrecht, 1891) (F,M) [Figs] | |
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| | [8] Triconia derivata (Heron & Bradford-Grieve, 1995) (F,M) [Figs] | |
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| | [9] Triconia elongata Böttger-Schnack, 1999 (F,M) [Figs] | |
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| | [10] Triconia furcula (Farran, 1936) (F,M) [Figs] | |
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| | [11] Triconia giesbrechti Böttger-Schnack, 1999 (F, M) [Figs] | |
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| | [12] Triconia gonopleura Böttger-Schnack, 1999 (F) [Figs] | |
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| | [13] Triconia hawii (Böttger-Schnack & Boxshall, 1990) (F,M) [Figs] | |
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| | [14] Triconia hirsuta Wi, Böttger-Schnack & Soh, 2010 (F) [Figs] | |
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| | [15] Triconia inflexa (Heron, 1977) (F,M) [Figs] | |
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| | [16] Triconia minuta Giesbrecht, 1892 (F, M) [Figs] | |
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| | [17] Triconia pacifica Cho, Kim, Böttger-Schnack, Lee, 2013 (F,M) [Figs] | |
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| | [18] Triconia pararedacta Wi, Böttger-Schnack & Soh, 2012 (F) [Figs] | |
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| | [19] Triconia parasimilis Böttger-Schnack, 1999 (F) [Figs] | |
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| | [20] Triconia quadrata (Heron & Bradford-Grieve, 1995) (F,M) [Figs] | |
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| | [21] Triconia recta Böttger-Schnack, 1999 (F,M) [Figs] | |
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| | [22] Triconia redacta (Heron & Bradford-Grieve, 1995) (F,M) [Figs] | |
|
| | [23] Triconia rufa (Boxshall & Böttger, 1987) (F,M) [Figs] | |
|
| | [24] Triconia similis (Sars, 1918) (F,M) [Figs] | |
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| | [25] Triconia thoresoni (Heron & Frost, 2000) (F,M) [Figs] | |
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| | [26] Triconia umerus (Böttger-Schnack & Boxshall, 1990) (F,M) [Figs] | |
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| | [27] Triconia sp. Böttger-Schnack, 1999 [Figs] | |
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 Any use of this site for a publication will be mentioned with the following reference :
Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2024. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed November 21, 2024] © copyright 2005-2024 Sorbonne University, CNRS
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