List species and varieties by family
Pseudocyclopiidae T. Scott,1894 ; emend. Jaume, Fosshagen & Iliffe, 1999 ( Clausocalanoidea )
(1) Frigocalanus Schulz, 1996 (? Pseudocyclopiidae )
Rem.: 1 sp.

[1] Frigocalanus rauscherti  Schulz, 1996   (F,M)    [Figs]
(2) Paracyclopia Fosshagen, 1985
Rem.: type: Paracyclopia naessi Fosshagen & Iliffe,1985. Total: 1 sp.

Diagnosis from Bradford-Grieve (1994, p.131) :
- As for the family definition.
- The posterior margins of urosomal segments finely striated, except anal segment.
- A1 reaches about half the length of prosome; 24-segmented in female; 23-segmented on the right side of the male.
- A2 exopod and endopod of equal length.Md blade with fine hyaline teeth.
- P1 endopod with 4 setae.
- Inner distal margin of basis of P2-P4 with a long strong spine, flanged distally.
- P5 3-segmented including a common basal segment in female, ending in a short segment with 2 distal points; simple, slender, asymmetrical with right leg longer in the male.

Paracyclopia gitana    Carola & Razouls, 1996   (M)
Ref.: Carola & Razouls, 1996 (p.448, Descr.M, figs.M)
Loc: Baléares (Minorque: grotte sous-marine)
Rem.: Cf. Stygocyclopia balearica

[1] Paracyclopia naessi  Fosshagen, 1985   (F,M)    [Figs]
(3) Pseudocyclopia T. Scott, 1892 emend
Rem.: Coastal forms. Type: Pseudocyclopia crassicornis T. Scott,1892. Total: 4 spp. + 1 inc. sed.

Diagnosis from Bradford-Grieve (1994, p.95) :
- As for the family definition.
- Body short and compact with the anterior head strongly vaulted dorsally and more or less compressed laterally.
- Rostral prominence with rostral filaments.
- Last pediger segment rounded off posteriorly.
- Urosome with anal segment well developed in both sexes.
- Caudal rami short with 4 terminal setae.
- A1 unusually short and tapering rapidly distally, 16- to 20-segmented, the 1st segment very large.
- A2 with 6-segmented exopod wihch is much shorter than endopod.
- Md blade very strong, masticatory part thickened with cutting edge irregularly curved and minutely denticulate; palp with both rami well-developed.
- Mx1 normal.
- Mx2 bearing terminal setae.
- Mxp slender with coxa remarkably produced distally; terminal part reflexed.
- Swimming legs with very strong outer-edge spines on the exopods; terminal spine on P2-P4 coarsely serrate.
- Coxa seta on P3 transformed into a strong spine.
- Female P5 3-segmented including a common basal segment, terminal segment largest and spiniferous.
- Male P5 of moderate size, uniramous on both sides; terminating in a styliform claw on the right; on the left with the basal part very swollen, remaining part 3-segmented.

Diagnosis after Jaume, Fosshagen & Iliffe (1999, p.403) :
- Pseudociclopiidae.
- Rostrum completely incorporated into cehalosome, pointed, with 2 tiny terminal filaments.
- Female A1 with compound segments I-IX and XII-XIII resulting in 18-segmented condition, although most species with segment X, or wegments X and XI, additionally incorporated into compound segment, resulting in 17 or 16-segmented condition.
- Male A1 symmetrical, with segmentation pattern identical to female except for additional compound segment XXII-XXIII, resulting in corresponding 17, 16 or 15-segmented conditions (see below : fig. 9A-C). Segment XV naked in both sexes. No duplication of aesthetascs on any segment of male A1.
- 1st A2 endopod segment fully incorporated into basis.
- Mx2 endopod unsegmented..
- P1 without outer basal spine ; endopod with 5 setae.
- 1st exopod segment of P3 frequently with accessory spine sub-marginally on posterior surface of segment, near to distal end of outer margin.
- P4 of both sexes differing only in presence of inner basal spine in female and absence in male.
- Female P5 3-segmented, uniramous, each ending y, 3 points. Male P5 asymmetrical, uniramous, one leg with basis swollen laterally, other longer, filiform.
- Female genital double-somite symmetrical.

[1] Pseudocyclopia caudata  T. Scott, 1894   (F)    [Figs]

[2] Pseudocyclopia crassicornis  T. Scott, 1892   (F,M)    [Figs]

[3] Pseudocyclopia giesbrechti  Wolfenden, 1902   (F,M)    [Figs]

[4] Pseudocyclopia insignis  Andronov, 1986   (F,M)    [Figs]

[5] Pseudocyclopia minor  T. Scott, 1892   (F,M)    [Figs]
Rem.: This form is considered as incertae sedis by Jaume & al. (1999, p.405). This form does not fit into the diagnosis of the genus. Even though it apparently exhibits a 17-segmented A1, with a segmentation pattern similar to that of Pseudocyclopia, its other characters are very peculiar. For example, one of the male P5 (Scott 1892, did not state precisely which one) is biramous, the proximal compound segment of A1 possesses a hook-like process, and the P1 display both an outer basal spine and only 4 setae on the endopod. The biramous condition of the male P5 is especially notable since all other Pseudocyclopiids have uniramous legs;
Likely, P5 male is incomplete in the Scott's drawing and only the left leg observe, nevertheless the size is very small in the genus. (C.R.)

Pseudocyclopia muranoi    Ohtsuka, 1992   (F,M)
Ref.: Ohtsuka, 1992 (p.296, figs.F,M, Rem.); Chihara & Murano, 1997 (p.892, Pl.163: F,M)
Rem.: Transferred to : Thompsonia muranoi.

Pseudocyclopia stephoides    Thompson, 1895   (F,M)
Rem.: Cf. Thompsonia stephoides

Pseudocyclopia spp.    Jaume, Fosshagen & Iliffe, 1999   (M)
Ref.: Jaume & al., 1999 (p.403, figs.M)
Loc: Norvège (off Lyroddane)
(4) Stygocyclopia Jaume & Boxshall, 1995
Rem.: type: Stygocyclopia balearica. Total: 4 spp.
The species occur in marine caves or the mixed marine zone of anchialine caves.

Diagnosis after Jaume & al. (1999, p.405):
- Pseudocyclopiidae.
- Rostrum fused to cephalosome, with 2 apical filaments.
- A1 symmetrical in both sexes, 23-segmented in females, 22-segmented in males; compound segments I-IV and X-XI present in both sexes, XXI-XXIII in males; segtment XII naked. Male A1 with quadrithek on segments III, V, VII and IX.
- A2 endopod 1 free, not incotporated into basis.
- Mx2 endopod 2-segmented.
- Endopod of P1 with 5 setae.
- 1st exopod segment of P3 without accessory spine.
- P4 of both sexes similar, with or without inner coxal spine.
- Male P5 slender, uniramous, asymmetrical, each ending in two points; left longer than right, 5-segmented, not swollen proximally; right 3 or 4-segmented.
- Female P5 3-segmented, uniramous, each ending in three points.
- Female genital double-somite symmetrical.

[1] Stygocyclopia australis  Jaume, Boxshall & Humphreys, 2001   (F,M)    [Figs]

[2] Stygocyclopia balearica  Jaume & Boxshall, 1995   (F,M)    [Figs]

[3] Stygocyclopia philippensis  Jaume, Fosshagen & Iliffe, 1999   (F,M)    [Figs]
(5) Thompsonopia Jaume, Fosshagen & Iliffe, 1999
Rem.: Type: Thompsonia mediterranea. Total: 3 spp.

Diagnosis after Jaume, Fosshagen & Iliffe in Sarsia, 1999, 84. :
- Rostrum pointed, movable, not fused to cephalon, with 2 terminal filaments.
- A1 of both sexes 20-segmented, displaying characterisyic compound segment X-XIV.
- Female A1 with segment I completely incorporated into II-IV, and with segments XXII and XXIII separate.
- Male A1 with segment I separate ; articulation between segments XXII and XXIII partially expressed in right A1, not expressed in left ; quadrithek on segment VII.
- 1st A2 endopod segment fully incorporated into basis.
- Mx2 endopod unsegmented.
- P1 withouter basal spine ; endopod with 5 setae.
- 1st exopod segment of P3 lacking accessory spine on outer margin.
- P4 armature sexually dimorphic ; with male lacking inner coxal spine and with 2 proximal segments of endopod bearing ordinary setae ; those of female bearing inner coxal spine and with armature on 2 proximal endopod segments reansformed into robust spines.
- Female P5 3-segmented, uniramous, each ending in 3 points.
- Male P5 asymmetrical, uniramous, with basis of left leg swollen posteriorly ; right leg elongate, filiform.
- Female genital double-somite symmetrical.

Remarks: The sexual dimorphism involving the expression of the articulation between segment 1 and compound segment II-IV in the males of the new genus deserves a comment here. Whereas this state is manifested in the new species (T. mediterranea) and also in T. stephoides, both Thompson (1895) and Sras (1901-1903) described the male antennules of T. stephoides as being similar to the female, except they did not figure them. Examination of material of this species from other Norwegian localities has demonstrated that the articulation is fully expressed in the males. The same is also possibly true for the males of T. muranoi since Ohtsuka (1992) did not figure the male antennules.

[1] Thompsonopia mediterranea  Jaume, Fosshagen & Iliffe, 1999   (F,M)    [Figs]

[2] Thompsonopia muranoi  (Ohtsuka, 1992)   (F,M)    [Figs]

[3] Thompsonopia stephoides  (Thompson, 1895)   (F,M)    [Figs]

 Any use of this site for a publication will be mentioned with the following reference :

Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2024. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at [Accessed June 16, 2024]

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