Scolecitrichidae Giesbrecht, 1892 ( Clausocalanoidea ) | |
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Amallophora T. Scott, 1894 | |
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Rem.: | Bradford,1973 (p.138,139) originally considers this subgenus, then “genus”, as a synonym of Xanthocalanus. A certain number of forms, formerly placed in the genus Amallophora are divided since, in addition, among Xanthocalanus (Phaennidae), in the other genera of the Scolecitrichidae (see in Bradford et al., 1983, p. 71 & foll.). This way, the genus is emptied of its substance although certain species are difficult to transfer. |
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Amallophora affinis Sars, 1905 (F) |
Ref.: | Sars, 1905 b (p.21) |
Rem.: | Cf. Scaphocalanus affinis |
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Amallophora altera Farran, 1929 (F,M) |
Ref.: | Farran, 1929 (p.252) |
Rem.: | Cf. Mixtocalanus alter |
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Amallophora claviger T. Scott, 1909 (M) |
Ref.: | T. Scott, 1909 (p.124) |
Rem.: | Cf. Xanthocalanus claviger |
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Amallophora cornifer Tanaka, 1960 (M) |
Ref.: | Tanaka, 1960 a (p.107) |
Rem.: | Cf. Xanthocalanus cornifer |
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Amallophora crassirostris Tanaka, 1960 (M) |
Ref.: | Tanaka, 1960 a (p.109) |
Rem.: | Cf. Xanthocalanus crassirostris |
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Amallophora cristata Sciacchitano, 1930 (F,M) |
Ref.: | Sciacchitano, 1930 (p.22, figs.F,M); 1930 a (p.150, Descr.F,M); Sewell, 1948 (p.540: lapsus calami) |
Loc: | G. d'Aden |
N: | 1 |
Lg.: | (451) F: 5-4; M: 6-2 |
Rem.: | espèce douteuse. Cf . spp. inc. sedis. |
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Amallophora dubia T. Scott, 1894 (M) |
Syn.: | Scolecithrix (Amallophora) dubia T. Scott, 1894 b (p.55) |
Rem.: | Cf. Scopalatum dubia |
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Amallophora dubia similis T. Scott, 1894 (M) |
Syn.: | Scolecithrix (Amallophora) dubia similis T. Scott, 1894 b (p.56) |
Rem.: | Cf. Scaphocalanus major |
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Amallophora elegans Wolfenden, 1911 (F) |
Syn.: | Scaphocalanus elegans : Park, 1982 (p.77) |
Ref.: | Wolfenden, 1911 (p.266, Descr.F, figs.F); Sewell, 1948 (p.523); Bradford, 1973 (p.144); Bradford & al., 1983 (p.93) |
Loc: | Atlant. équatorial (off I. St. Paul NE) |
N: | 1 |
Lg.: | (10) F: 2,6 |
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Amallophora gracilis Wolfenden, 1911 (F) |
Ref.: | Wolfenden, 1911 (p.265, Descr.F, figs.F) |
Rem.: | Cf. Scaphocalanus : ? Scaphocalanus affinis ou Scaphocalanus magnus |
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Amallophora impar Wolfenden, 1911 (F) |
Ref.: | Wolfenden, 1911 (p.263, figs.F) |
Loc: | Antarct. (Indien) |
Lg.: | (10) F: 2,4 |
Rem.: | Le transfert à Scaphocalanus et la synonymie de cette forme ne sont pas définitivement établis. Cf. ? Scaphocalanus farrani |
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Amallophora irritans Tanaka, 1960 (M) |
Ref.: | Tanaka, 1960 a (p.113) |
Rem.: | Cf. Xanthocalanus irritans |
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Amallophora macilenta Grice & Hulsemann, 1970 (M) |
Ref.: | Grice & Hulsemann, 1970 (p.192) |
Rem.: | Cf. Xanthocalanus macilenta |
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Amallophora magna T. Scott, 1894 (F) |
Syn.: | Scolecithrix (Amallophora) magna T. Scott, 1894 b (p.55) |
Rem.: | Cf. Scaphocalanus magnus |
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Amallophora media Sars, 1907 (F) |
Ref.: | Sars, 1907 a (p.16) |
Rem.: | Cf. Scaphocalanus major |
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Amallophora obtusifrons Sars, 1905 (F) |
Syn.: | Amallothrix obtusifrons : Sars, 1925 (p.179, figs.F, [non M |
Rem.: | Cf. Pseudoamallothrix obtusifrons |
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Amallophora oculata Tanaka, 1960 (M) |
Ref.: | Tanaka, 1960 a (p.111) |
Rem.: | Cf. Xanthocalanus oculata |
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Amallophora robusta T. Scott, 1894 (F) |
Syn.: | Scolecithrix (Amallophora) robusta T. Scott, 1894 b (p.56) |
Rem.: | Transfert probable: Cf. Amallothrix robusta |
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Amallophora rotunda Wolfenden, 1906 (F) |
Ref.: | Wolfenden, 1906 (p.44, Descr.F, figs.F) |
Loc: | Atlant. NE |
N: | 1 |
Lg.: | (239) F: 2,15 |
Rem.: | Le statut de cette espèce reste à préciser. Cf. spp inc. sedis. |
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Amallophora rotunda Grice & Hulsemann, 1970 (M) |
Ref.: | Grice & Hulsemann, 1970 (p.192) |
Rem.: | Cf. Xanthocalanus rotunda |
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Amallophora serrata Tanaka, 1960 (M) |
Ref.: | Tanaka, 1960 b (p.104) |
Rem.: | Cf. Xanthocalanus serrata |
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Amallophora smithae Grice, 1962 (F) |
Ref.: | Grice, 1962 (p.205) |
Rem.: | Cf. Scopalatum smithae |
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Amallophora subbrevicornis Wolfenden, 1911 (F) |
Ref.: | Wolfenden, 1911 (p.262) |
Rem.: | Cf. Scaphocalanus subbrevicornis |
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Amallophora typica T. Scott, 1894 (F,M) |
Syn.: | Scolecithrix (Amallophora) typica T. Scott, 1894 b (p.54) |
Ref.: | Sewell, 1929 (p.175) |
Rem.: | Cf. Xanthocalanus typicus |
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(1) Amallothrix Sars, 1925 | |
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Rem.: | Bradford (1973, p.142) redefines this genus (type: Scolecithricella gracilis Sars,1905) what leads to various synonymies. Park, 1980 (p.29) contests the validity of the position of Bradford, preferring following that of Vervoort (1951). The count of the species is very difficult. In Bradford & al. (1983, p.78) one enumerates 10 "sure" (*), 20 probable, 3 that do not totally harmonize with the definition (**) and 2 that have been transferred in new genera. Provisional total of 16 spp + 1 unidentified. Type species: Scolecithricella gracilis Sars, 1905.
Definition from Bradford & al. (1983, p.77) : - Pedigerous segments 4 and 5 fused in female, with fusion line sometimes dorsally ; may be separate in male. - Rostrum of 2 filaments. - A1 22-23 segments in female ; 19-31 in male. - Mx1 inner lobe 1 (arthrite) with 2 posterior surface setae ; inner lobe 3 with 4 setae ; endopod segment 1 separated from segments 2 and 3. - P1 exopodal segment 1 usually with an external spine. - Male mouthparts slightly reduced. - Female P5 uniramous, 3-segmented, although 2 orall segments maybe fused ; terminal segment with 2 to 4 spines (inner one of which is longest) ; surface of leg often ornamented with small spinules. - Male left P5 endopod usually shorter than exopod, not extending past distal part of exopodal segment 2 ; right leg endopod short, at most reaching slighrly further than distal part of basis.
Diagnosis after Vyshkvartzeva (2000, p.234) : - Pediger somites 4 and 5 in female separte or partially or completely fused. - Posterolateral corners of last prosomal somite slighly produced, broadly rounded, usually with an incurvation near dorsal side. - In male, pegiger somites 4 and 5 separte, not produced, broadly rounded. - Rostrum bifurcated, both long proximal processes strong, sausage-shaped, tapering distally into sensory filament as long as proximal part or about 1/3 its length. - A1 female 23-24-segmented, 8th and 9th, sometimes also 24th and 25th segments fused. A1 male slightly asymmetrical : in left 8th-12th segments, in right also 20th and 21th segments fused. - A2 endopod about 2/3-4/5 ,length of exopod ; exopodal segments 2-6 with 1 seta each in female and male ; exopodal segment 7 usually with 1 medial and 3 distal setae. - Md basipod usually with 3 inner setae ; 2 setae frequently rudimentary ; endopod reachibg about 2/3 length of exopod. - Mx1 : inner lobe 1 with 2 (in A. aspinosa, with 4) posterior setae ; inner lobe 3 with 4 (in A. obscura, with 3) setae ; exopod with 8-9 setae. - Mx2 : inner lobes 1-4 with 3 sclerotized setae ; inner lobe 5 with 3 sclerotized setae (one seta being stronger, hook-like, denticulated) and 1 worm-like sensory filament. 3 endopodal segments with 3 long worm-like and 5 brush-like setae being shorter than 3 others. - P1 : exopodal segments 1-3 usually with 1 external spine about ½-4/5 length of exopodal segment 2 (in A. lobophora and A. obscura, exopodal segment 1 without external seta ; in A. aspinosa, exopodal segments 1 and 2 without external seta each). - P2 : distal outer corner of endopodal segment 1 produced into obtuse or acute spine-like process ; external spine of exopodal segment 1 usually long, more than half as long as exopodal segment 2. - Posterior sueface of P2-P4 exopod and endopod with spines and spinules frequently arranged in arcs ; on P4 ornamentation scarcer than on P2-P3. - P5 female usually 2-segmented ; common basal segment long ; distal segment elongate, curved inward, usually with 3 spines (internal spine the strongest, as long as segment or slightly longer or shorter, with external edge serrated ; apical spine about 1/4 -1/2 length of internal ; external spine small, usually situated opposite to internal spine ; in some species, P5 segments with spines posteriorly.
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Amallothrix aculeata Esterly, 1913 (F) |
Syn.: | Scolecithrix aculeata Esterly, 1913 (p.183) |
Rem.: | Probable dans ce G. |
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[1] Amallothrix arcuata (Sars, 1920) (F,M) (*) [Figs] | |
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Amallothrix auropecten Giesbrecht, 1892 (F,M) |
Syn.: | Scolecithrix auropecten Giesbrecht, 1892 (p.266) |
Rem.: | Cf. Archescolecithrix auropecten |
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Amallothrix curticauda A. Scott, 1909 (F) |
Syn.: | Scolecithricella curticauda A. Scott, 1909 (p.94) |
Rem.: | Probable dans ce G. |
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Amallothrix denticulata Tanaka, 1962 (M) |
Syn.: | Scolecithricella denticulata Tanaka, 1962 (p.85) |
Rem.: | Probable dans ce G. |
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[2] Amallothrix aspinosa (Roe, 1975) (F) (**) [Figs] | |
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[3] Amallothrix dentipes (Vervoort, 1951) (F,M) (*) [Figs] | |
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Amallothrix elephas Esterly, 1913 (F) |
Syn.: | Scolecithrix elephas Esterly, 1913 (p.185) |
Rem.: | Probable dans ce G. |
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Amallothrix emarginata Farran, 1905 (F,M) |
Syn.: | Scolecithrix emarginata Farran, 1905 (p.36, Descr.F, figs.F); ? Scolecithrix inornata Esterly, 1906 a (p.67, figs.F) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.228) |
Rem.: | Cf. Pseudoamallothrix emarginata. |
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[4] Amallothrix falcifer (Farran, 1926) (F,M) (*) [Figs] | |
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[5] Amallothrix farrani Rose, 1942 (F) [Figs] | |
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Rem.: | meso-bathypelagic. ? Cf. Scolecithricella farrani. The genus of this species remains to be determined. This species resemble Scolecithrix longipes Giesbrecht1,1892, and may be a synonym. |
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[6] Amallothrix gracilis (Sars, 1905) (F,M) (*) [Figs] | |
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Amallothrix hadrosoma Park, 1980 (F) (*) |
Syn.: | Scolecithricella hadrosoma Park, 1980 (p.66, Descr.F, figs.F) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.228) |
Lg.: | (523) F: 5,66-5 |
Rem.: | Cf. Pseudoamallothrix hadrosoma |
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Amallothrix incisa Farran, 1929 (F) |
Syn.: | Scolecithrix incisa Farran, 1929 (p.244) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.228) |
Rem.: | Cf.: Pseudoamallothrix incisa. |
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Amallothrix indica Sewell, 1929 (F,M) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.228) |
Rem.: | Cf.: Pseudoamallothrix indica |
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Amallothrix inornata Esterly, 1906 (F,M) |
Syn.: | Scolecithrix inornata Esterly, 1906 a (p.67) |
Rem.: | ? Cf. Amallothrix emarginata . La situation de cette espèce est à préciser. Voir à Scolecithrix inornata |
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[7] Amallothrix invenusta C.B. Wilson, 1950 (F) [Figs] | |
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Rem.: | G. not well confirmed. |
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Amallothrix laminata Farran, 1926 (F,M) |
Syn.: | Scolecithrix laminata Farran, 1926 (p.265) |
Rem.: | Cf. Pseudoamallothrix laminata . |
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Amallothrix lanceolata Tanaka, 1962 (M) |
Syn.: | Scolecithricella lanceolata Tanaka, 1962 (p.87) |
Rem.: | Probable dans ce G. |
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Amallothrix lobata Sars, 1920 (F) |
Syn.: | Scolecithricella lobata Sars, 1920 c (p.9) |
Rem.: | Cf.: Falsilandrumius lobatus |
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[8] Amallothrix lobophora (Park, 1970) (F, ?M) (*) [Figs] | |
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Amallothrix longispina Schulz, 1991 (F,M) |
Ref.: | Schulz, 1991 (p.206, figs.F,M) |
Loc: | Mer Arabe (Oman) |
Lg.: | (524) F: 1,56-1,42; M: 1,48-1,36 |
Rem.: | Cf. Pseudoamallothrix longispina |
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Amallothrix magnus Wolfenden, 1911 (F) |
Syn.: | Scolecithrix magnus Wolfenden, 1911 (p.258) |
Rem.: | Probable dans ce G. ? Cf. “ Scolecithrix ctenopus “ Group. Voir à Scolecithrix magnus |
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Amallothrix medius Wolfenden, 1911 (F) |
Syn.: | Scolecithrix medius Wolfenden, 1911 (p.256) |
Rem.: | Probable dans ce G. |
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Amallothrix modica Tanaka, 1962 (F) |
Syn.: | Scolecithricella modica Tanaka, 1962 (p.58, Descr.F, figs.F) |
Ref.: | Vyshkvartzeva, 2000 (p.235) |
Rem.: | Probable dans ce G. |
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Amallothrix mollis Esterly, 1913 (F) |
Syn.: | Scolecithrix mollis Esterly, 1913 (p.186) |
Ref.: | Vyshkvartzeva, 2000 (p.235) |
Rem.: | Probable dans ce G. |
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Amallothrix obscura Roe, 1975 (F) |
Syn.: | Scolecithricella obscura Roe, 1975 (p.318, Descr.F, figs.F); Bradford & al., 1983 (p.103) |
Ref.: | Vyshkvartzeva, 2000 (p.235-236: Rem.) |
Rem.: | Probable dans ce G. |
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Amallothrix obtusifrons Sars, 1905 (F) |
Syn.: | Amallophora obtusifrons Sars, 1905 b (p.22) |
Rem.: | Cf. Pseudoamallothrix obtusifrons. |
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[9] Amallothrix parafalcifer (Park, 1980) (F) (*) [Figs] | |
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[10] Amallothrix paravalida Brodsky, 1950 (F,M) [Figs] | |
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Rem.: | meso-bathypelagic-abyssal. Genus not well confirmed. After Tanaka (1962, p.75) the females from Sagami Bay (Japan) are identical with the species referred by Sars (1925) as Amallothrix valida, which differs from Scolecithrix valida Farran (1908) (= Amallothrix valida): 1- smaller sier; 2- the length of A1; 3- the length of the outer edge spine of the 1st exopodal segment of P2; 4- the structure of serration on the terminal spine of exopod of P2 and P3 ? See: Amallothrix valens. According to Park (1970) the synonymy is not doubtful, in contrary to Tanaka (1962) and Bradford & al. (1983). The geographic distribution of this species is uncertain. |
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Amallothrix profunda Brodsky, 1950 (M) |
Ref.: | Brodsky, 1950 (1967) (p.263, Descr.M, figs.M); Bradford, 1973 (p.144); Roe, 1975 (p.320, 321, Rem.); Park, 1982 (p.77); Bradford & al., 1983 (p.93, Rem.) |
Loc: | Pacif. NW |
Lg.: | (22) M: 2,6 |
Rem.: | ? Cf. Scolecithricella laminata . Cf. Pseudoamallothrix profunda |
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Amallothrix propinqua Sars, 1920 (F,M) |
Syn.: | Scolecithricella propinqua Sars, 1920 c (p.9) |
Ref.: | Vyshkvartzeva, 2000 (p.235) |
Rem.: | Probable dans ce G. |
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[11] Amallothrix pseudoarcuata (Park, 1970) (F) (*) [Figs] | |
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[12] Amallothrix pseudopropinqua (Park, 1980) (F,M) (*) [Figs] | |
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[13] Amallothrix robusta (T. Scott, 1894) (F) (*) [Figs] | |
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Amallothrix robustipes Grice & Hulsemann, 1965 (F) |
Ref.: | Grice & Hulsemann, 1965 (p.239); 1967 (p.16); Markhaseva & Ferrari, 2005 a (p.124) |
Rem.: | Cf. Brodskius robustipes. |
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[14] Amallothrix sarsi Rose, 1942 (F) [Figs] | |
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Rem.: | Probable transfer to Scolecithricella. |
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Amallothrix spinata Tanaka, 1962 (F) |
Syn.: | Scolecithricella spinata Tanaka, 1962 (p.81) |
Ref.: | Vyshkvartzeva, 2000 (p.235) |
Rem.: | Probable dans ce G. |
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Amallothrix timida Tanaka, 1962 (F) |
Syn.: | Scolecithricella timida Tanaka, 1962 (p.83) |
Ref.: | Vyshkvartzeva, 2000 (p.235) |
Rem.: | ? Cf. A. mollis. Probable dans ce G. |
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Amallothrix valens Farran, 1926 (F) |
Syn.: | Scolecithrix valens Farran, 1926 (p.263) |
Ref.: | Rose, 1933 (1934 a) (p.12); Vyshkvartzeva, 2000 (p.235) |
Rem.: | ? A. paravalida. Probable dans ce G. |
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[15] Amallothrix valida (Farran, 1908) (F,M) [Figs] | |
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Amallothrix sp. Marques, 1958 (F) |
Ref.: | Marques, 1958 (p.212) |
Rem.: | Cf. Scolecithricella marquesae |
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[16] Amallothrix sp. Roe, 1975 (M) [Figs] | |
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(2) Archescolecithrix Vyshkvartzeva, 1989 | |
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Rem.: | Type: Scolecithrix auropecten Giesbrecht,1892; Total: 1 sp. |
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[1] Archescolecithrix auropecten (Giesbrecht, 1892) (F,M) [Figs] | |
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(3) Bradfordiella Andronov, 2007 | |
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Rem.: | Type: Bradfordiella kurchatovi. 2 spp. The assignment of this genus to Scolecitrichidae is questionable; Andronov (2007) showed that the A1, with ancestral segments X-XV fused, the reduced distal part of the Mx1 and the huge Mx2 distinguish this genus from other ''Bradfordians'' genus. Markhaseva, Laakmann & Renz (2003, p.21) consider following characters that do not allow to specify its family placement: 1- Md basis lacking setae (versus 1 to 4 setae in other ''Bradfordians''; 2 - Mx1 setal formula 9, 0, 0, 3 (setae on fused distal basal endite and endopod), 2 and 5 setae; Bradfordiella shares a coxal endite without setae with Heteramalla and Pseudophaenna (versus coxal endite with 1-5 setae in other ''Bradfordians''), a proximal basal endite without setae with Kyphocalanus (Kyphocalanidae) (versus 2-4 setae in other ''Bradfordians''), a distal basal endite fused to the endopod with Rostrocalanidae, Brodskius and some species of Rythabis; 3 - Mx2 enditic lobe of endopod with 1 seta is shared with Kyphocalanus (versus usually 4, rarely 3 setae in other ''Bradfordians''; 4 - Mxp syncoxa in Bradfordiella without setae on praecoxal endites (versus setae present in other ''Bradfordians''); 5 - Mxp, endopodal segment 3 with 1 seta (versus 2-3 setae in other ''Bradfordians'' (Table 3-5 in Markhaseva & al., 2014, p.84) . Until a more detailed description of the genus is prepared, its familly placement remains uncertain. |
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[1] Bradfordiella fowleri (Farran, 1926) (F,M) [Figs] | |
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[2] Bradfordiella kurchatovi Andronov, 2007 (F) [Figs] | |
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(4) Brodskius Markhaseva & Ferrari, 2005 | |
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Rem.: | type: Brodskius benthopelagicus Markhaseva & Ferrari, 2005. Total: 6 spp. + 1 unidentified Markhaseva & Schulz (2007) include this genus in the Tharybidae family. Markhaseva, Laakmann & Renz (2014, p.79) maintain this genus in Tharybidae family.
Diagnosis from Markhaseva & Ferrari (2005, p.114) : Female: - Cephalosome and pediger 1 separate or fused, pedigers 4 and 5 separate. - Rostrum 2 delicate filaments. - Posterior corners of prosome laterally as an indented lobe. - A1 24-segmented. - A2 coxa and basis without setae. - Md gnathobase elongate, narrow medially with a knob on distal face, cutting edge narrow, with 2 distinct incisions separating groups of teeth. - Mx1 distal basal endite fused to unsegmented endopod, setae of distal basal endite inseparable from setae of endopod; 1 seta on proximal basal endite and 2 setae on distal basal endite + endopod long and thick with long setules. - Mx2 proximal praecoxal endite with 4 sclerotized setaze; both coxal endiotes and proximal basal endite with 1 thick seta, seta on basal endite thickest and claw-like; 5 worm-like setae with well-developeds setules and 2 or 3 short brush-like setae on distal basal endite + ramus. - Mxp syncoxa without seta on proximal praecoxal endite, 2 setae on middle praecoxal endite, 3 setae on distal praecoxal endite; all praecoxal setae sclerotized; coxal endite with 3 setae. - P1-P4 clausocalanoidean segmentation and setation. - P5 3-segmented; distal segment, the exopod, with 1 medial, 1 lateral and 1 subterminal seta, and terminal unarticulated extension. Male: - Adult male similar to female except: posterior corners of prosome not indented. - left A1 24-segmented, right 23-segmented; more and larger aesthetascs. Md gnathobase poorly-developed. - Mx1 reduced in size and setation. - Mxp praecoxal endites of syncoxa with 0, 2, 3 sclerotized setae smaller. - Von Vaupel klein 's organ of P1 without basal seta and anterior knob. - P5 biramous , right exopod 2-segmented, left exopod 3-segmented; both endopod 1-segmented, right small and left one longer than exopod.
For Markhaseva & Ferrari (2005, p.115) the synapomorphies of Brodskius are: 1- Md gnathobase narrow, with 2 distinct incisions separating groups of teeth; 2- Mx1 with 1 long, thick and heavily setulated seta on proximal basal endite and 2 such setae on distal basal endite + endopod; setae of distal basal endite inseparable from those of endopod. |
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[1] Brodskius abyssalis Markhaseva & Schulz, 2007 (F) [Figs] | |
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[2] Brodskius arcticus Andronov & Kosobokova, 2011 (F) [Figs] | |
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[3] Brodskius benthopelagicus Markhaseva & Ferrari, 2005 (F) [Figs] | |
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[4] Brodskius confusus Markhaseva & Ferrari, 2005 (F) [Figs] | |
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[5] Brodskius paululus (Park, 1970) (F, ?M) [Figs] | |
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[6] Brodskius robustipes (Grice & Hulsemann, 1965) (F) [Figs] | |
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[7] Brodskius sp. Markhaseva & Ferrari, 2005 (M) [Figs] | |
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Byrathis Markhaseva & Ferrari, 2005 | |
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Rem.: | Cf. Diaixidae |
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(5*) Cenognatha Bradford-Grieve, 2001 | |
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Rem.: | In the broad sens in this family. type: Neoscolecithrix antarctica Hulsemann, 1985. Total: 1 sp. (provisionally).
Diagnosis from Bradford-Grieve (2001a, p.792) : - Posterior prosome in lateral view extends into 2 posteriorly-directed spines. - Rostrum short and rounded with 2 filaments. - Mx2 endite 1 with 5 well-developed setae, endite 5 with 1 strong spine-like seta and 3 setae (2 of these possibly worm-like sensory setae in C. antarctica), endopod usually with 3 long worm-like setae and 5 brush-like sensory setae. - Mxp coxal endite 3 with 1 or 3 setae with bulbous base. P5 female with distal segment and its basis subequal in length. - Male P5 of similar lengths on both sides, styliform o,n right, endopod present on at least one side, 1-segmented and spine-like on right. - A2 exopod with full complement of ancestral setae as in Clausocalanus: 2 on segment 1, 3 on segment 2, 1 seta each on segments 3-5, segment 6 with 1 proximal seta and 3 terminal setae. - Edge of gnathobase of Md with dorsal spinulose seta unmodified. Endite 3 of Mx2 without modified seta. - Posterodistal border of basis of P1-P3 without spines. - Mxp praecoxa and coxa fused (based on C. farrani N. Vyshkvartzeva, pers. comm.), coxal endite 2 with 2 setae and 1 brush-like sensory seta. - Male mouthparts fully developed.
In this genus Bradford-Grieve, 2004 (p.285) includes the two species Neoscolecithrix caetanoi and farrani. According to Markhaseva & Schulz, 2010 (p.5) Procecenognatha (Diaixidae) and Cenognatha are closely related and share a combination of characters: short rostrum with 2 filaments; biramous right P5 in the male; a similar setation and segmentation pattern of the oral parts. For Markhaseva, Laakmann (2013, p.19) this genus must be transfered in Diaixidae family. |
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[1] Cenognatha antarctica (Hulsemann, 1985) (F, Juv.M) [Figs] | |
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(6) Diaiscolecithrix Markhaseva, Schulz & Renz, 2010 | |
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Rem.: | Type: Diaiscolecithrix andeep Markhaseva, Schulz & Renz, 2010. Total: 1 sp. + 1 undet.
Diagnosis from Markhaseva, Schulz & Renz (2010, p.114): Female: - Cephalosome and pediger 1 partly fused, pedigers 4 and 5 separate. - Posterior corners of prosome as short triangular lobes pointed distally. - Rostrum as a plate with filaments. - Upper and lower lips well developed and form circular oral cone-like structure). - Urosome 4-segmented; genital double-somite symmetrical. - A1 24-segmented, 1st segment with 3 setae. - A2 exopodal segment 1 witout setae, endopodal segment 1 with 1 seta; exopod nearly 1.5 times as long as endopod. - Md gnathobase elongate and slender with 2 acute spine-lke teeth distally; basis with 2 setae; endopoal segment 1 without setae; segment 2 with 9 setae; exopod 5-segmented with 1, 1, 1, 1, 2 setae. - Mx1 praecoxal arthrite with 7 setae; coxal endite, proximal basal endite and distal basal endite with 2 setae each; endopod segments fused with 4-5 setae including at least 1 sensory; exopod with 7 setae; coxal epipodite with 8 setae. - Mx2 praecoxa with outer hump,; proximal praecoxal endite with 3 setae; distal praecoxal endite with 1 seta; coxal endites with 2 setae each; proximal basal endite with 4 setae; distal basal endite okus endopod with 8 setae (3 long worm-like and 5 brush-like sensory setae). - Mxp syncoxa with 1, 2, and 1 setae on proximal praecoxal, middle and distal praecoxal endite respectively; coxal endite with 3 setae; basis with 3 medial setae; endopodal segments 2 to 5 with 4, 3, 3+1, 4 setae. - Segmentation and setation of P1-P4 typical for Clausocalanoidea. - P5 present, 2-segmented and lacking spines on both segments.
Male unknown. |
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[1] Diaiscolecithrix andeep Markhaseva, Schulz & Renz, 2010 (F) [Figs] | |
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[2] Diaiscolecithrix sp. Markhaseva, Schulz & Renz, 2010 (F) [Figs] | |
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(7*) Falsilandrumius Vyshkvartzeva, 2001 | |
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Rem.: | In the broad sense in this family. Type: Scaphocalanus bogorovi. Total: 3 spp. For Markhaseva, Laakmann & Renz (2014, p.81) this genus must be transfered in Diaixidae family.
Diagnosis after Vyshkvartzeva (2001, p.79) : - Forehead with a low median crest or without crest. - Rostrum as a short plate with 2 long filaments. - Pedigers 4 and 5 separate. - Posterolateral corners of pediger somite 5 produced distally into triangular lobes with a rounded tip or terminating with a small tooth-like process. - Urosme of 4 somites, as long as ¼-1/5 of prosome. - Genital somite (laterally) with a conspicuous genital prominence. - Caudal rami as long as wide or slightly longer than wide. - A2 endopod as long as exopod or slightly longer. - Md basipod with endopod as long as exopod. - Mx1 with 2 posterior setae on inner lobe 1 ; 3-4 setae on inner lobe 2 and 4 setae on inner lobe 3 ; exopod with 10-11 setae. - Mx2 inner lobe 1 with 4 ( ?)-5 setae ; inner lobes 2-4 with 3 setae each ; inner lobe 5 with 4 sclerotized setae ; endopod with 3 worm-like and 5-6 brush-like sensory setae. - Mxp syncoxa in the middle of inner margin with 3 setae, 1 sometimes transformed into a brush-like sensory seta. - P1 basipod without inner distal seta ; endopod without setose outer lobe ; endopodal segments 1-3 with long, stout outer spine each, orendopodal segments 1 and 2 without outer spine. - In P2-P4, outer distal corner of all endopodal segments produced as a sharp spine-like process. - Outer sine of exopodal segment 1 of P2 shorter than outer spines of exopodal segments 2- 3. - Terminal spines of exopods of P2-P4 longer than exopodal segment 3. - Posterior surfaces of P2-P4 protopod and exopod segments usually covered with small denticles ; endopod segments with long spinules. - P5 3-segmented ; distal segment longer than wide and slightly flattened, usually with 4 spines (inner spine the longest, subequal in length to the segment ; outer spine situated before the middle of the outer margin). |
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[1] Falsilandrumius angulifrons (Sars, 1920) (F,M) [Figs] | |
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[2] Falsilandrumius bogorovi (Brodsky, 1955) (F, juv. C5 M) [Figs] | |
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[3] Falsilandrumius lobatus (Sars, 1920) (F) [Figs] | |
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(8*) Grievella Ferrari & Markhaseva, 2000 | |
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Rem.: | Type: Grievella shanki Ferrari & Markhaseva, 2000. Total: 1 sp. In the broad sense in this family. Total: 1 sp. For Markhaseva, Laakmann & Renz (2014, p.81) this genus must be transfered in Diaixidae family.
Genus' morphologic characters : - Cephalosome and pediger 1 fused, pedigers 4 and 5 fused. - Posterior corners of prosome rounded laterally, not reaching beyond the anterior margin of genital complex. - Genital complex symmetrical in ventral view, asymmetry in lateral view results from small integumental bumps. - 3 articulating abdominal somites posterior to genital complex. - Caudal rami with 4 large, terminal setae, 1 small medial-ventral seta, and 1 small lateral-dorsal seta - A1 24-segmented, reaching thoracic segment 3. Segment 22 with ear-like extension anteriorly with ear cavity facing proximally. - Mx2 proximal praecoxal endite with 5 setae and attenuation point, distal endite with 3 setae. Proximal coxal endite with 3 setae, distal coxal endite with 3 setae. Proximal basal endite with 1 long, thick seta, 2 long, thin seta and 1 poorly sclerotized setae. Distal basal lobe + exopod with 9 sensory setae, 3 worm-like setae distally and 6 brush-like setae, all about the same length and with short setules; 3 brush-like setae of the same thickness, 2 thinner, 1 very thin brush-like setae. - Mxp syncoxa with 1 long seta on proximal lobe; 2 long setae on middle lobe, both well-sclerotized; 3 short setae on distal lobe; coxal lobe with 3 setae and denticles on distal face. Basis with 3 setae on unattentuated proximal lobe and 2 setae on distal lobe; proximal denticles short and thick. Endopod 5-segmented from proximal to distal with 4, 4, 3, 4 (1 lateral),4 (1 lateral) setae respectively. . - P1 without seta but with medial denticlesBasis with medial seta sharply curved.Endopod 3-segmented; endopod 1-segmented complex with 3 medial and 2 terminal setae; quadrate protuberence of Von Vaupel-Klein's organ with anterior row of denticles proximally and posterior pore on lateral edge. - P2 exopod 3-segmented; endopod 2-segmented. - P3 exopod 3-segmented; endopod 3-segmented. - P4 exopod 3-segmented; endopod 3-segmented. - P5 coxa proximal; basis without seta fused to 1-segmented exopod with distally 1 medial, 1 terminal and 1 lateral setae.
Ferrari & Markhaseva (2000, p.1084) note that 4 derived characters (synapomorphies) states separate this species-type of the genus from other scolecitrichids: small integumental bumps on the genital complex; an ear-like extension on segment 22 of A1; 2 lateral setae on the distal endopodal segment of P2; and a denticle-like attenuation of the proximal praecoxal lobe of Mx2. The authors believe the first of these may be an autapomorphy. The type-species shares the absence of an outer soine on both the proximal and middle exopodal segments of P1 with a number of other scolecitrichids. This apomorphy that is shared with these other scolecitrichid species results from convergence; it is not evidence for monophyly.
The type-species shares 5 setae on the proximal praecoxal lobe of Mx2 with the scolecitrichids Xantharus renatehaassae, 1 of only 2 species in this genus, with Neoscolecithrix antarctica, 1 of 6 species in this genus, and with all 5 species in this genus, and with all 5 species of the genus Landrumius. The remaining species of Scolecitrichidae have 3 or 4 setae on the proximal praecoxal lobe of Mx2. Species of the Phaennidae have 5 setae on this proximal praecoxal lobe, except for a few species of Xanthocalanus with 4 setae. Species of Diaixidae, Parkiidae, and Tharybidae have 3 or 4 setae. Type -species shares 9 sensory setae on the distal basal lobe plus exopod of Mx2 with all 5 species of Landrumius. The remaining species of Scolecitrichidae have 8 sensory setae on the distal basal lobe plus exopod of Mx2, with the exception of Xantharus renatehaassae which has 8 sensory setae and 1 sclerotized seta. 8 sensory setae is the number most often reported for Phaennidae, Diaixidae, and Tharybidae, although 9 sensory setae have been reported for some phaennids and several tharybid-like copopods (unpubl. obs.). Species-type shares 1, 2 and 3 setae, from proximal to distal, on the 3 praecoxal lobes of Mxp with Xantharus renatehaassae, with all 5 species of Landrumius, with Neoscolecithrix antarctica and Neoscolecithrix magna; however, these latter 2 species differ quite significantly in other morphological features. Praecoxal lobes with 1, 2 and 3 setae also are known for some diaixids and tharybids. Setation for the remaining Scolecitrichidae usually is 1, 2, and 1 setae. For Phaennidae setal numbers 1, 2 and 2 seem to have been conserved. If it is assumed that the larger number of elements is the plesiomorphic state (see Monchenko & Von Vaupel-Klein, 1999) for these above 3 characters of the Mx2 and Mxp, then the states for the type-species (Grievella shanki) provide no direct information about its phylogenetic relationships with other copepods sharing the same character states. Among related families Phaennidae, Diaixidae, Tharybidae, and Parkiidae, synapomorphies have been proposed only for the latter family. The authors place the species Grievella shanki, (consequently the new genus) within the Scolecitrichidae tentativ/ The number and kind of sensory setae on the distal basal lobe plus exopod of Mx2 alone is not adequate to diagnose the Scolecitrichidae, or to separate all of its species from those of the Phaennidae and other families with these kinds of sensory setae. Careful redescriptions of the setarion of A2, Mx1, Mx2 and Mxp are required before the different synapomorphies of the 5 families with their included genera can be clarified. |
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[1] Grievella shanki Ferrari & Markhaseva, 2000 (F) [Figs] | |
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(9) Heteramalla Sars, 1907 | |
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Rem.: | type: Heteramalla dubia Sars, 1907. 1 sp. After A. Scott (1909, p.86) the main characteristics of Heteramalla are: 1 - A strong chitinised and slightly bifurcate lamelliform rostrum; 2 - An extraordinary development of two of the sensory appendages on the apex of Mx2; 3 - P5 very small and rudimentary in the female.
For Markhaseva, Laakmann & Renz (2003, p.22) this genus is removed from Scolecitrichidae, but the taxonomic position of this genus will probably clarified when the male is discovered. In this instance the genus has not been attributed to any of the currently known ''Bradfordians families''. This genus differs from Scolecitrichidae by the following characters: 1- Mx1 coxal endite without setae (versus usually 2 setae in scolecitrichids); 2 - Mx2 setal formula 5, 3, 3, 3, 4 (versus 3-4 setae at praecoxal endite in scolecitrichids); 3 - Mxp praecoxal endites setal formula 1, 1, and 2 (versus 1, 2, and 1 in scolecitrichids), with setae of praecoxal endites as: 1sc + 1w + (1sc + 1w) (versus single brush-like seta present in scolecitrichids) (Markhaseva, pers. com.). A Heteramalla male has not yet been found, therefore the taxonomic position of this genus will probably be clarified when the male is discovered. |
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[1] Heteramalla sarsi Roe, 1975 (F) [Figs] | |
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(10*) Landrumius Park, 1983 | |
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Rem.: | In the broad sense in this family. Total: 4 spp. For Markhaseva, Laakmann & Renz (2014, p.81) this genus must be transfered in Diaixidae family. |
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[1] Landrumius antarcticus Park, 1983 (F) [Figs] | |
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[2] Landrumius gigas (A. Scott, 1909) (F, juv.M) [Figs] | |
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[3] Landrumius insignis (Sars, 1920) (F) [Figs] | |
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[4] Landrumius sarsi (C.B. Wilson, 1950) (F) [Figs] | |
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(11) Lophothrix Giesbrecht, 1895 | |
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Rem.: | type : Lophothrix frontalis Giesbrecht,1895. Bradford & al. (1983) enumerates 2 "sure" species (*), 4 probable (4 having been transferred to the genus Landrumius) and 1 doubtful. Provisional total: 7 soo. (of which 1 doubtful).
Diagnosis from Bradford & al. (1983, p.88) : - Pedigers 4 and 5 fused or separate. - Rostrum may have short conical points or elongate filiform appendages. - Head with or without crest. - A1 24-segmented in female; about 22 segments in male. - Mx1 inner lobe 1 with 4 setae on posterior surface; inner lobe 3 with 4 setae. - P1 exopod segment 1 with or witout external spine; - Male mouthparts reduced. - Female P5 3-segmented, the last two of which may be fused; last segment with 3 or 4 spines. - Male P5 of Scaphocalanus type. |
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Lophothrix angulifrons Sars, 1920 (F) |
Syn.: | Scaphocalanus angulifrons Sars, 1920 c (p.8) |
Rem.: | Transfert dans ce G. à préciser (Bradford, 1973, p.144). Voir à Scaphocalanus angulifrons |
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[1] Lophothrix frontalis Giesbrecht, 1895 (F,M) (*) [Figs] | |
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Lophothrix frontalis major Sewell, 1929 (F,M) |
Ref.: | Sewell, 1929 (p.195, figs.F,M, Rem.); 1947 (p.149, fig.F); Tanaka, 1961 a (p.150, figs.F,M) |
Lg.: | (11) F: 6,718-6,250; M: 4,5; (29) F: 6,125-5,812; (108) F: 6,9; M: 5,39 |
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Lophothrix frontalis minor Sewell, 1929 (F) |
Ref.: | Sewell, 1929 (p.196, Descr.F, figs.F); 1947 (p.149, fig.F) |
Lg.: | (11) F: 5,633-4,75; (29) F: 5,375-4,875 |
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Lophothrix gigas A. Scott, 1909 (F,M) |
Syn.: | Brachycalanus gigas A. Scott, 1909 (p.81) |
Rem.: | Cf. Landrumius gigas |
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[2] Lophothrix humilifrons Sars, 1905 (F,M) [Figs] | |
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Rem.: | Bathypelagic. Sargasso Sea: 1500-2000 m (Deevey & Brooks, 1977, station "S"); 2000-1000 m (Harding, 1974). Probable in this Genus. Vervoort (1965, p.60) does not include Wilson's L. humilifrons into the synonymy of this species, because Wilson's specimens are exclusively male from the Albatros Expedition (1950), but the Vervoort's impression is that probably the species should be Scottocalanus thomasi A. Scott, of which species females are recorded from the same station (off Bataan, Philippines). For Park (1983, p.187) both the female and male are similar to L. frontalis may be easily distinguished. |
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Lophothrix insignis Sars, 1920 (F) |
Ref.: | Sars, 1920 c (p.8) |
Rem.: | Cf. Landrumius insignis |
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[3] Lophothrix latipes (T. Scott, 1894) (F,M) (*) [Figs] | |
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[4] Lophothrix quadrispinosa Wolfenden, 1911 (F) [Figs] | |
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Lophothrix sarsi Wilson, 1950 (F) |
Ref.: | C.B. Wilson, 1950 (p.253) |
Rem.: | Cf. Landrumius sarsi |
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Lophothrix securifrons Wolfenden, 1904 (F) |
Ref.: | Wolfenden, 1904 (p.120) |
Rem.: | Cf. Scottocalanus securifrons (F) (T. Scott,1894) |
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[5] Lophothrix similis Wolfenden, 1911 (F) [Figs] | |
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[6] Lophothrix simplex Wolfenden, 1911 (F) [Figs] | |
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Lophothrix thorsoni Björnberg, 1975 (F) |
Ref.: | Björnberg, 1975 (p.178) |
Rem.: | Cf. Landrumius gigas |
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[7] Lophothrix varicans Wolfenden, 1911 (F) [Figs] | |
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(12) Macandrewella A. Scott, 1909 | |
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Rem.: | Type: Macandrewella joanae A. Scott,1909. Total: 11 sp. (+ 1 ?).
Diagnosis from Bradford & al. (1983, p.91) : - Head with circular lens-like organs on frontal margin. - Head and pediger 1 fused, pedigers 4 and 5 separate. - Rostrum with slender filaments attached to common bifurcate base. - Posterior part of metasome and genital segment often asymmetrical. - P5 female absent or small, 1-segmented, attached to basal portion. - Male generally as female. - P5 male well developed; right leg endopod well developed, exopod with basal part produced internally and last segment usually forked apically; left leg endopod well developed and exopod with apical spine.
Ohtsuka & al. (2002, p.532, emended) : - Cephalosome fused to 1st pediger. - Rostrum bifurcate, bearing pair of long filaments at tip. - Single distinct cuticular lens present at base of rostrum. - Anteromedial crest-like plate absent on cephalosome. - 4th and 5th pedigers fused. - Last prosomal somite carrying 1 or 2 pairs of acute or lobate processes in female, and pair of short, poineted prominences in male. - Urosome 4-segmented, at most 1/3 as long as prosome in female; and 5-segmented in male, with anal somite almost telescoped into preceding somite. - Caudal setae symmetrical or asymmetrical in female. - A1 23-segmented in female - A1 male asymmetrical, 18-segmented on right side, 19-segmented on left side. - Mx2 endopod bearing 3 worm-like and 5 brush-like sensory setae. - Mouthparts of male similar to those of female, but with some elements on mandibular palp slightly more reduced. - Terminal elements of Mxp endopod more developed in male than in female. - Strong processes predent on posterior surface of endopods of P2 and P3 in both sexes. - Female right/left P5 present or absent. - Male P5 highly developed, both biramous ; right exopod 3-segmented, left 2-segmented; endopods uni-segmented. Second exopodal segment of right leg forming distinct chela with 3rd segment bearing inner thumb-like process. 3rd exopodal segment left leg armed with 2 membranous elements terminally.
Ohtsuka & al. (2002, p.534) point to the genus is clearly distinguishable from other scolecitrichid genera by the combination of the characters: presence of a single cephalic lens; the highly developed male P5 with the 2nd and 3rd exopodal segments of the right leg forming a chela. Later, Farran (1936) created a closely related genus Scolecocalanus with a single cephalic lens at the rostral base. Main differences between Macandrewella and Scolecocalanus are (see Campaner, 1989): an anterior crest-like cephalic expansion present in Scolecocalanus and absent in Macandrewella; the female P5 are 2-segmented with a common base or absent in mMcandrewella, whereas in Scolecocalanus, the left leg is uni-segmented with a stout terminal spine and the right leg is reduced. In addition, the rostral filaments are long and slender in Macandrewella but short and thick in Scolecocalanus. |
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[1] Macandrewella agassizi C.B. Wilson, 1950 (F,M) [Figs] | |
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[2] Macandrewella asymmetrica Farran, 1936 (F,M) [Figs] | |
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[3] Macandrewella chelipes (Giesbrecht, 1896) (F,M) [Figs] | |
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[4] Macandrewella cochinensis Gopalakrishnan, 1973 (F,M) [Figs] | |
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[5] Macandrewella joanae A. Scott, 1909 (F,M) [Figs] | |
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[6] Macandrewella mera Farran, 1936 (F) [Figs] | |
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[7] Macandrewella omorii Ohtsuka, Nishida & Nakaguchi, 2002 (F,M) [Figs] | |
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[8] Macandrewella scotti Sewell, 1929 (F,M) [Figs] | |
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[9] Macandrewella serratipes Ohtsuka, Nishida & Nakaguchi, 2002 (F,M) [Figs] | |
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[10] Macandrewella sewelli Farran, 1936 (F,M) [Figs] | |
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[11] Macandrewella stygiana Ohtsuka, Nishida & Nakaguchi, 2002 (F,M) [Figs] | |
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[12] Macandrewella tuberculata Chen, 1987 [Figs] | |
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(13) Mixtocalanus Brodsky, 1950 | |
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Rem.: | type: Amallophora altera Farran,1929. Total: 3 spp.
Diagnosis from Brodsky (1967, p.237) : - Head and 1st thoracic segment separate or fused; 4th and 5th thoracic segments separate. - Rostrum bifid. - Exopod of P1 with 3 outer spines. - Legs with posterior armature both on exopodites and on endopodites. - P5 female 3-segmented, without spines, except the distal segment, which bears 1 apical spine. - P5 male uniramous, right leg strongly reduced, left leg 5-segmented, several times longer than the right.
Nota: the male is considered as doubtful as Phaennidae. Mx2 male bears only 2 setae with very large sperical tips
For Brodsky (1967, p.237) the new genus, after male and female found, is closely related to Phaennidae and Scolecitrichidae and occupies ann intermediate position between them. Mixtocalanus closely resembles Xanthocalanus, but differs from it and from the remaining genera in the structure of the distal setae of Mx2; in the female there are 3 types of setae: 3 are band-shaped, 3 are apically plumose and 2 have spherical tips. Tips of all the types of setae are small, much smaller than in Heteramalla, which also differs in the structure of the rostrum.
Diagnosis from Bradford & al. (1983, p.91) : - head and pediger 1 fused or separate, pedigers 4 and 5 separate. - Rostrum bifurcate with 2 filaments in female. - Female Mx2 endopod with 3 worm-like and 5 brush-like sensoty filaments, 2 of which have spherical tips; all filaments smaller than in Heteramalla. - Male Mx2 with only 2 brush-like sensory filaments. - P1 exopod segment 1 with outer edge spine. - Female P5 3-segmented, with 1 terminal spine. - Male P5 uniramous, right leg strongly reduced, left leg several times longer than right. |
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[1] Mixtocalanus alter (Farran, 1929) (F,M) [Figs] | |
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[2] Mixtocalanus robustus Brodsky, 1950 (F) [Figs] | |
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[3] Mixtocalanus vervoorti (Park, 1980) (F,M) [Figs] | |
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(14*) Neoscolecithrix Canu, 1896 (part.) | |
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Rem.: | type: Neoscolecithrix koehleri Canu,1896. 8 spp. + 1 unnamed A comparison of the characters in the (F) leads Hulsemann (1985, p.60, 61) to define 2 groups: N. koehleri , N. magna , N. sp. Bradford 1973 et N. farrani , N. watersae , N. antarctica . For Vyshkvartzeva these three latter species belong to the Scolecitrichidae.
Diagnosis from Bradford-Grieve (2001a, p.791) : - Posterior prosome, in lateral view, extends into 2 posteriorly-directed spines. - Rostrum wide and long with 2 points each with filament. - Edge of Md gnathobase with very wide dorsal spinulose seta. - Mx2 endite 1 usually with 4 well-developed setae (sometimes 1 additional small spine present), endite 3 with 1 seta modified as a large brush-like sensory seta directed distally, endite 5 with 1 strong spine-like seta, 2 smaller setae and 1 worm-like sensory seta, and endopod with 5 long worm-like and 3 brush-like sensory setae. - Mxp coxal endite 3 with 1 of 3 setae with bulbous base. - Posterodistal border of basis of P1-P3 and often coxa of P2 and P3 with posterior surface spine (s). - Female P5 with distal segment at least twice as long as basis. - Male P5 asymmetrical, very short on one side, if endopod present it is rudimentary and fused to basis. - Male mouthparts fully developed, similar to those of female. - A2 exopod with full complement of ancestral setae as in Clausocalanus: 2 on segment 1, 3 on segment 2, 1 seta each on segments 3-5, segment 6 with 1 proximal seta and 3 terminal setae. - Mxp praecoxa and coxa separate, coxal endite 2 with 2 setae and 1 brush-like sensory seta. For Bradford-Grieve (2001a, p.791) the family relationship of the genus Neoscolecithrix s.l. is problematical. These species have been variously placed in the Phaennidae, (e.g. Bradford-Grieve, 1999) or Tharybidae (Bradford & al., 1983 and Vyshkvartzeva (2000) considers that species that are in Group II fit well within the Scolecitrichidae on the basis of the number of 3 worm-like and 5 brush-like sensory setae on the Mx2 endopod. nevertheless Group II species differ from the majority of Scolecitrichidae in important respects. Ohtsuka, Boxshall & Fosshagen (2003, p.53: History of the genus). For Markhaseva, Laakmann & Renz (2013, p.19) this genus must be tranfered in Diaixidae family. |
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Neoscolecithrix antarctica Hulsemann, 1985 (F, juv.M) |
Ref.: | Hulsemann, 1985 (p.55, Descr., figs. C5, F & M) |
Rem.: | Cf. Cenognatha antarctica |
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Neoscolecithrix bidentata Farran, 1905 (F) |
Syn.: | Oothrix bidentata Farran, 1905 (p.42, Descr.F, figs.F); Sewell, 1948 (p.501); ? Chen & Zhang, 1965 (p.58, figs.F); Shih & Young, 1995 (p.71) |
Ref.: | Bradford & al., 1983 (p.123); Hulsemann, 1985 (p.55, 60, Rem.) |
Rem.: | Cf. Neoscolecithrix koehleri. La forme rapportée par Chen & Zhang (1965) comme Oothrix bidentata (F) pourrait être un copépodite 5 d'une espèce encore non décrite de Neoscolecithrix ; Pour Ohtsuka & al., 2003 (p.60) cette forme pourrait être synonyme de Neoscolecithrix japonica , la localisation (Est de la mer de Chine) comme les conditions thermiques (eaux chaudes estivales) ainsi que des caractères morphologiques rendent douteuses la synonymie avec N. koehleri . |
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[1] Neoscolecithrix caetanoi Alvarez, 1985 (F,M) [Figs] | |
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Rem.: | hyperbenthic. ? Cf. Cenognatha. According to Alvarez (1985, p.200) the majority ofthe authors (Rose, 1933; Brodsky, 1967; Fosshagen, 1972 among others) included the genus in the Phaennidae. Bradford (1973) redefining the family, did not accept the above opinion, because the Neoscolecithrix species lack an important feature of the Phaennidae: the presence of spines on the endopods of P2 to P4. But she did not suggest to which family the genus should belong. Canu (1896) had already called attention to the similarity of the genera Scolecithrix, Xanthocalanus, and Neoscolecithrix, proposing that the last one be intermediate between the two. Fosshagen (1972) remarked that N. farrani is an association of characteristics of several families, and concluded that a revision of the systematic position of this genus and of the families related to it was necessary. The different modes of life have influence on the morphology of the animals, and this may mask the phylogenetic relationships existing between the several copepod genera. |
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[2] Neoscolecithrix farrani Smirnov, 1935 (F,M) [Figs] | |
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Rem.: | epibenthic-hyperbenthic. For Vyshkvartzeva (2000, p.217) this species belongs in the Scolecitrichidae. According to Bradford-Grieve in the genus Cenognatha. |
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[3] Neoscolecithrix japonica Ohtsuka, Boxshall & Fosshagen, 2003 (F,M) [Figs] | |
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[4] Neoscolecithrix koehleri Canu, 1896 (F,M) [Figs] | |
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[5] Neoscolecithrix magna (Grice, 1972) (F) [Figs] | |
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[6] Neoscolecithrix ornata Bradford-Grieve, 2001 (F) [Figs] | |
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[7] Neoscolecithrix watersae (Grice, 1972) (F,M) [Figs] | |
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Rem.: | bathypelagic (1465-1500 m), hyperbenthic. This species resembles Neoscolecithrix koehleri Canu, 1896. According to Campaner (1978 a, p.968) Xanthocalanus watersae (Grice, 1972) , from the structure of the P5, the species must be removed to the genus Neoscolecithrix. |
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[8] Neoscolecithrix sp. Bradford, 1973 (F, Juv.5 F) [Figs] | |
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Neoscolecithrix sp. Chahsavar-Archad & Razouls, 1982 (Juv.F) |
Ref.: | Chahsavar-Archad & Razouls, 1982 (p.27, fig.Fjuv.); Bradford-Grieve, 2001 a |
Loc: | off Is. du Cap Vert. S |
N: | 1 |
Rem.: | Cf. ? |
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(15) Omorius Markhaseva & Ferrari, 2005 | |
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Rem.: | type: Omorius atypicus. Markhaseva & Ferrari,2005. Total: Total: 2 spp.
Diagnosis adult female from Markhaseva & Ferrari (2005, p.143) : - Cephalosome and pediger 1 fused dorsally, separate laterally; pedigers 4 and 5 fused dorsally and separate laterally. - Posterior corners of prosome as rounded lobes in lateral view. - Rostrum without filaments. - A1 24-segmented. - A2 coxa with 1 seta; basis with 2 setae; exopod with 1, 3, 1, 1, 1, 1, 1, 3 setae. - Md gnathobase with a tooth-like knob on posterior face, cutting edge wide proximal to distal. - Mx1 praecoxal endite with 4 posterior setae and 9 terminal setae, 3rd and 4th (2nd and 3rd from proximal seta) thin and slightly curved; coxal epipodite with 9 setae; proximal basal endite with 2 setae; distal basal endite with 3 setae; endopod articulating with basis with groups of 2 and 5 setae; exopod with 4 setae. - Mx2 praecoxal endite with 4 setae; proximal and distal coxal endites with 3 setae, 1 thicker; and proximal basal endite with 4 setae, 1 thicker, 1 worm-like; distal basal endite + ramus with 5 long, worm-like setae and 3 short, brush-like setae. - Mxp praecoxal endites of syncoxa with 1, 2, 1 sclerotized setae; coxal endite with 3 setae; endopod of 5-segmented with 4, 4, 3, 3+1 and 4 setae. - P1-P4 clausocalanoidean segmentation and setation. - P5 3-segmented, distal segment with 3 setae and terminal attenuation.
For Markhaseva & Ferrari (2005, p.143), at present3rd and 4th thin and slightly curved setae on the praecoxal arthrite of Mx1 is the only proposed synapomorphy for the monotypic Omorius. A sclerotized seta on distal praecoxal endite of Mxp syncoxa is shared with Archeoscolecithrix although the latter genus has 5 brush-like and 3 worm-like setae on the distal basal endite + ramus of Mx2, a situation reversed for Omorius. On the remaining 17 scolecitrichid-like genera, a brush-like seta replaces the sclerotized seta on the distal praecoxal endite of Mxp syncoxa. On the remaining 'bradfordian' genera, 2 or 3 setae are found on the endite. |
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[1] Omorius atypicus Markhaseva & Ferrari, 2005 (F) [Figs] | |
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[2] Omorius curvispinus Markhaseva & Schulz, 2007 (F) [Figs] | |
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(16) Parascaphocalanus Brodsky, 1955 (? Scolecitrichidae ) | |
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Rem.: | Type: Parascaphocalanus zenkevitchi Brodsky, 1955. Total: 1 sp.
Diagnosis from Vyshkvartzeva (2001, p.93) : - Forehead with a low median crest or without crest. - Rostrum as a short plate with 2 long filaments. - Pedigers 4 and 5 separate. - Posterolateral corners of pediger somite 5 produced distally into triangular lobes with a rounded tip or terminating with a small tooth-like process. - Urosme of 4 somites, as long as ¼-1/5 of prosome. - Genital somite (laterally) with a conspicuous genital prominence. - Caudal rami as long as wide or slightly longer than wide. - A2 endopod as long as exopod or slightly longer. - Md basipod with endopod as long as exopod. - Mx1 with 2 posterior setae on inner lobe 1 ; 3-4 setae on inner lobe 2 and 4 setae on inner lobe 3 ; exopod with 10-11 setae. - Mx2 inner lobe 1 with 4 ( ?)-5 setae ; inner lobes 2-4 with 3 setae each ; inner lobe 5 with 4 sclerotized setae ; endopod with 3 worm-like and 5-6 brush-like sensory setae. - Mxp syncoxa in the middle of inner margin with 3 setae, 1 sometimes transformed into a brush-like sensory seta. - P1 basipod without inner distal seta ; endopod without setose outer lobe ; endopodal segments 1-3 with long, stout outer spine each, orendopodal segments 1 and 2 without outer spine. - In P2-P4, outer distal corner of all endopodal segments produced as a sharp spine-like process. - Outer sine of exopodal segment 1 of P2 shorter than outer spines of exopodal segments 2- 3. - Terminal spines of exopods of P2-P4 longer than exopodal segment 3. - Posterior surfaces of P2-P4 protopod and exopod segments usually covered with small denticles ; endopod segments with long spinules. - P5 3-segmented ; distal segment longer than wide and slightly flattened, usually with 4 spines (inner spine the longest, subequal in length to the segment ; outer spine situated before the middle of the outer margin).
After Bradford & al. (1983, p.91) it is not clear that Parascaphocalanus is a scolecithrid. Brodsky (1955) figures the terminal part of Mx2 with 4 worm-like and 2 brush-like sendsory filaments, endopod segment 1 of P2 is small and narrow, and the male P5 very like that of some tharybids. On the other hand Parascaphocalanus appears to differ from most tharybids in that the A2 rami are of similar length and exopod segment 1 of P1 is without an outer edge spine. |
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[1] Parascaphocalanus zenkevitchi Brodsky, 1955 (F,M) [Figs] | |
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(17) Paraxantharus Schulz, 2006 | |
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Rem.: | type-species: Paraxantharus brittae Schulz,2006. Total: 2 spp. According to Markhaseva (2010, p.209) this genus is tentatively placed in the Diaixidae lineage (sensu Markhaseva & Ferrari, 2005); For Markhaseva, Laakmann & Renz (2014, p.81) this genus must be transfered in Diaixidae family.
Diagnosis male from Schulz (2006, p.48) : - cephalosome and pediger 1 fused, pedigers 4 and 5 incompletely coalescent. - Rostrum a bifurcate plate with 2 tiny filaments distinctly separate. - Urosome of 5 somites. - Anal somite very short (almost ''invisible''). - A1 extending to urosome, 23-segmented on left, 22-segmented on right side (due to fusion of ancestral segments XXII-XXIII); left segment XXII smaller than each of remaining segments. - A2 with exopod nearly 2 times length of endopod, ancestral segment I-IX armed with 1 seta each. - Mouthparts not reduced. - Md with gnathobase armed with 8 sharply pointed teeth. Mx2 with proximal praecoxal endite bearing 5 setae; endopod with 5 brush-like and 3 long worm-like plus 1 sclerotized setae. - Mxp syncoxa with 1, 2, 3, and 3 setae, endite 3 with 2sclerotized and 1 sensory setae bearing small terminal brush. - Legs of clausocalanoidean segmentation, 2nd and 3rd exopodal segments with strong outer spines, terminal exopodal spine large with comparatively few widely spaced teeth. - P1 endopod wih large outer lobe. P5 biramous with 1-segmented endopods, exopods 3-segmented; right exopod slender, with long terminal spine-like segment; left exopod shorter, compact. |
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[1] Paraxantharus brittae Schulz, 2006 (M) [Figs] | |
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[2] Paraxantharus victorbergeri Markhaseva, 2010 (F) [Figs] | |
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(18) Parkius Ferrari & Markhaseva, 1996 | |
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Rem.: | Boxshall & Halsey, 2004 (p.188; 190: F) deem that the Parkiidae family should be included in the Scolecitrichidae in the absence of a phylogenetic analysis. Total: 1 sp. + 1 undetermined For Markhaseva, Laakmann & Renz (2014, p.75, 77) this family is not included in the Scolecitrichidae, but in the Parkiidae family. |
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[1] Parkius karenwishnerae Ferrari & Markhaseva, 1996 (F, Juv.M) [Figs] | |
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[2] Parkius sp. (Grice & Hulsemann, 1967) (M) [Figs] | |
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(19) Plesioscolecithrix Markhaseva & Dahms, 2004 | |
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Rem.: | Type: Plesioscolecithrix juhlae Markhaseva & Dams, 2004. Total: 1 sp. Not included in the generic key in Boxshall & Halsey (2004, p.188). Earlier assigned to the Scolecitrichidae, later analysis of ''Bradforddians'' relationships showed that this genus does not belong to the monophyletic scolecitrichid lineage (Markhaseva & Ferrari, 2005). The taxonomic position of this genus therefore remains unresolved.
After Markhaseva, Laakmann & Renz (2013, p.22) this genus should be placed outside the Scolecitrichidae because of the following characters: 1 - the proximal part of right A1 in males has only ancestral segments X-XI fused (more segments fusions in scolecitrichids); 2 - A2 endopodal segment 1 with 1 seta (versus usually 2 setae in scolecitrichids); 3 - Mx1 praecoxal arthrite has no posterior setae (versus usually 1-4 posterior setae in scolecitrichids); 4 - Mx2 praecoxal artrite has 5 setae (versus 3-4) in scolecitrichids); 5 - Mxp praecoxa setal formula: 1, 2 and 2 setae (versus 1, 2 and 1 setae in scolecitrichids) (see Table 1-5), with setae of praecoxa as: 1sc + (1w+1sc) + (1sc+1br); 6 - male P5 right basipod not swollen (versus swollen in scolecitrichids); legs of silmple structure (versus complex in scolecitrichids); right and left coxa and basis nearly the same length (versus right basis significantly shorter than left bin scolecitrichids). |
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[1] Plesioscolecithrix juhlae Markhaseva & Dahms, 2004 (F,M) [Figs] | |
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(20) Pseudoamallothrix Vyshkvartzeva, 2000 | |
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Rem.: | type: Amallothrix profunda Brodsky,1950. Total: 15 spp. (of which 1 doubtful).
Diagnosis from Vyshkvartzeva (2000, p.227): - Pedigerous somites 4 and 5 usually fused in female ( articulation suture sometimes visible dorsally), usually separate in male and some females of large species. - Posterolateral corners of pedigerous somite 5 not produced, broadly rounded, in the middle part with a shallow incurvation (incision), or slightly produced, narrowly rounded with indentation near dorsolateral part. - Rostrum as a short plare with 2 strong but short rami continuing in 2 thick aesthetasc-like filaments; the latter longer than the strong proximal part and frequently notched at apex. - A1 female with 23-24 segments; 8th and 9th segments fused, 24th and 25th segme,ts sometimes fused. A1 of male slightly asymmetrical: left A1 20-segmented (8th-12th segments and 24-25th segments fused; right A1 with 19-segmented (also 20th-21th segments fused). - A2 endopod 1/2-2/, the length of exopod; exopodal segment 1 without seta, usually with a rounded swelling on internal margin (without swelling in P. cenotelis, P. longispina, P. ovata); exopodal segments 2-6 frequently with 1 seta each in female and male; seta of exopodal segment 2 usually shorter in female, than in male. - Mx1: inner lobe 1 with 2 or 4 posterior setae; inner lobe 3 usually with 4, sometimes with 3 setae (P. ovata, P. cenotelis); exopod usually with 8 setae (5 setae in P. ovata, P. cenotelis). - Mx2: inner lobes 1-4 with 3 setae each; inner lobe 5 with 2-3 sclerotized setae and 1 or sometimes 2 worm-like sensory setae; on inner lobe 3 , sometimes one of the 3 sclerotized setae transformed into sensory seta; 3-segmented endopod with 3 long worm-like and 5 brush-like sensory setae; distal brush of brush-like setae always small, but 3 of 5 setae have ''stem'' slightly or significantly longer and sometimes worm-like. - Male mouthparts similar to those of female in meristic details, but some setae are shorter , slightly reduced. - Each of the three segments of P1 exopod with external spine of about 1/2-4/5 the length of the segment, in P. longispina longer than segment (exopodal segment 1 without seta in P. birshteini; exopodal segments 1-2 without setae in P. canariensis. - External distal corner of endopodal segment 1 of P2 produced into long, spine-like process; external spine on endopodal segment 1 of P2 straight, usually short. - P2-P3 coxopod with a distinct indentation about in the middle of external margin ; internal margin usually with well marked projection distally ( without projection in P. longispina) ; the latter often with a notch ; dorsal surface of exopod and endopod segments with spines arranged in arcs and with numerous spinules. - P4 coxopod with external margin smooth, internal margin bearing a rounded lamelliform lobe ( absent in P. longispina) ; dorsal surface of segments with few spinules. - P5 female uniramous with 1 or sometimes 2 free subcylindrical segments and 2 (apical and ubapical) spines ; common basal segment long. - P5 male biramous, asymmetrical. In right leg, 1st and 2nd exopodal segments almost completely fused, 2nd segment with or without short distal projection, 3rd exopodal segment curved, bearing lamella-like spine on distal end ; right 1-segmented endopod usually longer than 3-segmented exopod or sometimes as long as exopod. Right P5 sometimes significantly shorter than left (P. indica). Sometimes (in P. ovata, ?P. cenotelis) P5 uniramous, right leg much shorter than left. The author in 2000 transfered 13 species in this genus. |
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[1] Pseudoamallothrix birshteini (Brodsky, 1955) (F) | |
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[2] Pseudoamallothrix canariensis (Roe, 1975) (F) [Figs] | |
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[3] Pseudoamallothrix cenotelis (Park, 1980) (F) [Figs] | |
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[4] Pseudoamallothrix emarginata (Farran, 1905) (F,M) [Figs] | |
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[5] Pseudoamallothrix hadrosoma (Park, 1980) (F) [Figs] | |
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[6] Pseudoamallothrix incisa (Farran, 1929) (F) [Figs] | |
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[7] Pseudoamallothrix indica (Sewell, 1929) (F,M) [Figs] | |
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[8] Pseudoamallothrix inornata (Esterly, 1906) (F,M) [Figs] | |
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[9] Pseudoamallothrix laminata (Farran, 1926) (F,M) [Figs] | |
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[10] Pseudoamallothrix longispina (Schulz, 1991) (F,M) [Figs] | |
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[11] Pseudoamallothrix obtusifrons (Sars, 1905) (F) [Figs] | |
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[12] Pseudoamallothrix ovata (Farran, 1905) (F,M) [Figs] | |
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[13] Pseudoamallothrix paralaminata Vyshkvartzeva, 2003 (F,M) | |
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[14] Pseudoamallothrix profunda (Brodsky, 1950) (F,M) [Figs] | |
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[15] Pseudoamallothrix soaresmoreirai (Bjornberg, 1975) (M) [Figs] | |
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(21) Puchinia Vyshkvartzeva, 1989 | |
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Rem.: | type: Puchinia obtusa. Total: 1 sp. The sensitive setae of the Mx2 indicate that this genus belongs to the Scolecithricidae and presents the most evolved characters in this family. Exop. 1 & 2 of the A2 with numerous small setae and the Ri of the Mx1 with 3 segments reveal archaic characters. 5 setae on the 2nd internal lobe of Mx; 2 sensory filaments and 1 usual chaeta in the middle protopodite of the Mxp; 1 sensory filament present on the 2nd-4th endites of Mx2. For Vyshkvartzeva, analysis of the evolutionary importance of the morphological features of Puchinia has revealed that it is characterized by a combination of a number of highly specialized features of its structure with the archaic ones, typical for the ancestor of the whole superfamily, by some parallelisms not only with the genera of its own family but with other Pseudocalanoidea. The bad condition of the type specimen makes it impossible to clarify the taxonomic position of the genus (Markhaseva pers. obs.). For Markhaseva, Laakmann & Renz (2014, p.23) the attribution of this genus to scolecitrichids is doubtful as Mx1 coxal endite bears 5 setae (versus 2 or rarely 4 setae in scolecitrichids) and Mxp praecoxa setal formula is 1, 2 and 2 (versus 1, 2 and 1 setae in scolecitrichids). |
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[1] Puchinia obtusa Vyshkvartzeva, 1989 (F) [Figs] | |
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(22) Racovitzanus Giesbrecht, 1902 | |
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Rem.: | type: Racovitzanus antarcticus Giesbrecht,1902. Total provisionally: 2 spp. For Tanaka (1961 a , p.185) the genus is closely allied to Scaphocalanus in the structure of the male P5 and intermediate between Scaphocalanus and Scolecithrix in having a rather long outer edge spine on the 2nd exopodal segment of P1.
Diagnosis from Bradford & al. (1983, p.91) ; - Rostrum cylindrical with 2 small filaments terminally. - Mx1 inner lobe 1 with 2 setae on posterior surface; inner lone 3 with 3 setae. - Mx2 endopod with worm-like and brush-like sensory filaments. - P1 exopod segment 1 without external spine. - Female P5 absent or 2-segmented, with 1 or 2 spines, neither of which are external. - Male P5 of the Scaphocalanus type.
After Bradford-Grieve (1983, p.91) the nature of the sensory filaments on Mx2 is difficult to observe and there is considerable variation in the number and type recorded by various authors. The male which Vervoort (1957) attributes erroneously to R. antarcticus appears to differ from other Racovitzanus in having an outer edge spine on exopod segment 1 of P1. |
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[1] Racovitzanus antarcticus Giesbrecht, 1902 (F,M) [Figs] | |
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Racovitzanus erraticus Vervoort, 1957 (M) |
Ref.: | Vervoort, 1957 (p.99) |
Rem.: | Cf. Racovitzanus antarcticus |
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[2] Racovitzanus levis Tanaka, 1961 (F,M) [Figs] | |
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Racovitzanus nanus Tanaka, 1953 |
Ref.: | Tanaka, 1953 (p.132, nom. nudum.) |
Rem.: | Cf. Racovitzanus porrectus |
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Racovitzanus pacificus Esterly, 1905 (F) |
Syn.: | Scolecithrix pacifica Esterly, 1905 (p.168) |
Ref.: | Bradford & al., 1983 (p.91); Park, 1983 (p.171, Rem.) |
Rem.: | Probable dans ce G. (maintenu provisoirement à Scolecithrix) |
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Racovitzanus porrectus Giesbrecht, 1888 (F) |
Syn.: | Scolecithrix porrecta Giesbrecht,1888; 1892 (p.266) |
Ref.: | Bradford & al., 1983 (p.91); Park, 1983 (p.171, Rem.) |
Lg.: | (143) F: 2,3 |
Rem.: | Probable dans ce G. (Maintenu provisoirement à Scolecithrix) |
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Racovitzanus sp. Grice & Hulsemann, 1967 (M) |
Ref.: | Grice & Hulsemann, 1967 (p.26, Descr.M, figs.M) |
Rem.: | Cf. Parkius sp. |
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(23*) Rythabis Schulz, 1995 | |
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Rem.: | Type: Rythabis atlantica Schulz & Beckmann, 1995. Total: 4 spp.
Diagnosis from Schulz, 1995 (1995, p.199) : - Body compact (tharybids). - Cephalosome and pediger 1 separate, pedigers 4 and 5 mostly separate. - Anal somite as long as preceding somite. - Rostrum a knob-like prominence lacking filaments. - A1 with fusion patterns of segments typical of superfamily Clausocalanoidea, and with ancestral segments XXVI and XXVII-XXVIII separated. - Exopod of A2 more than twice the length of endopod. - Md gnathobase bearing 3 acute teeth ventrally; basis with 2 long curved setae. - Mx1 well equipped with setal armature. - Mx2 comprising short praecoxa and relatively large endopod . - Mxp slender. - Legs of typical clausocalanoid segmentation. - Inner endopodal lobe of P1 reduced. - Outer exopodal spines of P1 and P2 (presumably P3 and P4 as well) exceptionally strong and ornamented with small marginal denticles. - Posterior face of P2-P5 spinose. - P5 uniramous, 3-segmented, terminal segment bearing 2 distal, 1 inner and 1 outer denticulated spines. - Male unknown (up to 2015).
For Schulz & Beckmann (1995, p.203) this genus shows closest affinities to the genus Tharybis, particularly on account of its compact ovate body form, the conical rostral knob lacking filaments, the separated segments XXVI and XXVII-XXVIII of A1, presence of identical numbers of large and massive spines on the praecoxal arthrite of Mx1, occurrence of 2 different types of maxillar aesthetascs and presence of 3 basal setae on Mxp, and non-broadened exopods of P1 to P4. However, the following are distinguishing characters taht separate members of Rythabis from Tharybis: 1- 4th and 5th pedigers mostly separated; 2- anal somite as long as preceding somite; 3- basis of Md equpped with only 2 setae; 4- maxillular epipodite, basis, endopod and exopod comparatively well developed and armed with higher numbers of setae; 5- brush-like aesthetascs of Mx2 and Mxp of different structure; 6- outer exopodal spines of P1 and P2 ornamented with denticles; 7- P5 having the exopod armed with 1 inner, 1 lateral and 2 distal articulated spines. In members of the genus Tharybis the terminal segment of P5 always carries a single inner distal spine in addition to usually 2 spine-like protrusions.
Markhaseva & Schulz (2007, p.742) consider the family placement as incompletely resolved. For Markhaseva, Laakmann & Renz (2014, p.85) the characters following prevent the placement of this genus in Tharybidae (placed in this family by Schulz & Beckmann, 1995, followed by Bradford-Grieve, 2004 and Vyshkvartzeva, 2005), as in the Scolecitrichidae (placed by Ohtsuka & al. 2003, and Boxshall & Halsey, 2004). The following characters prevent a placement of the genus Rythabis into the Scolecitrichidae: 1 - Mx2 endopod setal formula has usually 6 worm-like + 2 brush-like sensory setae (versus usually 3w + 5br sensory setae in scolecitrichids, however 1 species of Rythabis shares the 4w + 4br setation with the scolecitrichid genus Racovitzanus) and the Mxp praecoxal setal formula 1, 2 and 2 (versus 1, 2, and 1 setae in scolecitrichids), setae at praecoxa 1w + 2 sclerotised setae + (1sc +1br). Except for the differences in limbs setation, Rythabis is distinguised from scolecithrichids: 1 - The short A2 endopod (shared with Pseudophaenna, Diaixis and Tharybis: 2 - Mxp syncoxa with a row of long thin spinules proximal of coxal endite (absent in scolecitrichids); 3 - Md gnathobase with a row of spinules along cutting edge (absent in scolecitrichids); 4 - female P5 with 4 spines at the exopod (usually 3 spines in scolecitrichids). The following characters prevent the placement of Rythabis in Tharybidae: 1 - Mx1 coxal and distal basal endites with 4-5 setae (versus 2-3 setae in Tharybis), in some species the distal basal endite and endopod are fused (versus separate in Tharybis); 2 - Mx2 endopod setal formula 6w + 2br or 4w + 4br (versus 3w + 5br, 3W + 6br, or 3w + 5br + 1sc in Tharybis); 3 - Mxp praecoxal setal formula 1, 2, 2 (versus 1, 2 and 3 in Tharybis).
Laakmann, Markhaseva & Renz (2019, p.330, Table 1) consider this genus as incertae sedis genera in the ''Bradfodians families" .
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[1] Rythabis asymmetrica Markhaseva & Schulz, 2007 (F) [Figs] | |
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[2] Rythabis atlantica Schulz, 1995 (F) [Figs] | |
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[3] Rythabis heptneri Markhaseva & Ferrari, 2005 (F) [Figs] | |
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[4] Rythabis schulzi Markhaseva & Ferrari, 2005 (F) [Figs] | |
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(24) Scaphocalanus Sars, 1900 | |
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Rem.: | type: Amallophora magna T. Scott, 1894. For Bradford & al (1983) there are 14 'sure' species(*) and 14 'probable'(**). Provisional total 32 spp. (of which 1 doubtful) + 5 undet. After Tanaka (1961 a, p.157) the species of the genus resemble each other so closely in general appearance and in the structure of P5 that is very difficult to discriminate them. Moreover, P5 female exhibit a considerable degree of variability in the proportional lengths and the number of spines on the distal segment. The P5 os often furnished with abnormal endopod as illustrated by Vervoort (1957) in S. brevicornis Sars.
Diagnosis after Bradford & al. (1983, p.93) : - Pedigerous segments 4 and 5 fused in female, usually separate in male. - Head may have crest. - Rostrum of 2 filaments. - A1 22-segmented in female, with segments 4-20 flattened and expanded posteriorly; of 19-21 segments in male. - Mx1 inner lobe 1 usually with 3 setae on posterior surface; inner lobe 3 with 3 or 4 setae; endopod segment 1 separated from segments 2 and 3. - Mx2 endopod with 3 worm-like and 5 brush-like sensory filaments. - P1 exopod segment 1 without outer spine. P2 endopod segment 1 without pointed nouter distal extension. Male mouthparts reduced, especially Mx2 and inner lobes of Mx1. - Female P5 absent or, more usually, present and 3-segmented, often with last two segments fused; terminal segment with 2 to 4, usually strong spines. - Male left P5 endopod longer than exopod; right P5 endopod long, usually reaching segment 2, exopod segment 3 long and blade-like.
Diagnosis after Park (1982, p.78) : Female: - Head and 1st pedigeous segment fused; 4th and 5th thoracic segments fused, often with a line of segmentation visible dorsally. - Urosome 4-segmented, followed by moderately developed caudal rami. - Rostrum of 2 filaments, usually well-developed. A1 with segments 8 through 10 fused and with a transparent strip along posterior edges of segments 5 through 22. - Both endopod and exopod of A2 well developed; 1st exopodal segment without a seta. - Md with strong masticatory blade and well-developed basis followed by small endopod and large exopod. - Mx1 with coxa bearing 3 inner lobes and 1 outer lobe, and basis carrying endopod and exopod. - Mx2 with 5 well-developed lobes and a small endopod; endopod with 3 long vermiform and 5 short brush-like sensory setae. - Mxp with large coxa and basis followed by 5 small endopodal segments; basis with 3 middle and 2 terminal setae. - P1 with 1-segmented endopod and 3-segmented exopod; 1st exopodal segment without spines or setae. - P2 with 2-segmented endopod and 3-segmented exopod; coxa fringed medially with hair followed by a well-developed seta; basis devoid of setae and hair along inner margin. - P3 and P4, both endopod and exopod 3-segmented; basis of P3 similar to P2; in P4, coxa without inner marginal hair but with a moderately developed inner seta; - P5 absent or more usually 2-segmented; distal segment often divided partially into 2, with strong inner and terminal spines and ofren with 1 or 2 small outer spines.
Male: - Head and 1st pedigerous segment fused, 4th and 5th thoracic segments incompletely fused with line of segmentation clearly visible dorsally. - Urosome 5-segmented, followed by small caudal rami; - Rostrum of 2 filaments, usually well-developed. - A1 with segments 8 through 12 fused, without a transparent strip along posterior edge; segments 20 and 21 fused on right A1, but separate on left. - Md blade poorly developed, with small teeth on its distal edge; palp well-developed with a broad basis followed by strong endopod and exopod. - Mx1 with poorly developed inner lobes and well-developed outer lobe and exopod. - Mx2 poorly developed, with 5 lobes followed by an endopod bearing 3 vermiform and 5 brush-like sensory filaments. - Mxp relatively small; setae poorly developed, except those on last 2 endopodal segments; basis with 3 middle anfd 2 terminal setae. - P1 to P4 similar to those of female. - P5 asymmetrical, each leg with 2-segmented basipod followed by endopod and exopod; basis elongate in left leg but short in right. In left leg, exopod3-segmented; distal segment with hair and bristles; endopod 1-segmented with a small terminal spine and longer than exopod; In right leg, exopod 2-segmented with well-developed distal spine; 1st segment ofthen divided into two by a line; endopod 1-segmented with a terminal spine.
Bradford & al. (p.93) note that the number of setae on Mx1 inner lobes 1 and 3 appear not to be as constant (see Bradford, 1973). Scaphocalanus difficilis Roe, 1975 has 2 posterior surface setae on inner lobe 1 and S. difficilis, S. curtus, S. similis, S. invalidus all have 3 setae on inner lobe 3 (see Hure & Scotto di Carlo, 1968). |
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Scaphocalanus acrocephalus Sars, 1900 (F,M) |
Ref.: | Sars, 1900 (p.36); Vervoort, 1957 (p.94, Rem.); Park, 1982 (p.75, 76, Rem.); Kosobokova & Hirche, 2000 (p.2029, tab.2) |
Loc: | Arct. |
Rem.: | Cf. Scaphocalanus magnus (T. Scott,1894) |
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[1] Scaphocalanus acuminatus Park, 1970 (F) (*) [Figs] | |
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[2] Scaphocalanus acutocornis Vyshkvartseva, 1987 (F) [Figs] | |
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[3] Scaphocalanus affinis (Sars, 1905) (F,M) (*) [Figs] | |
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[4] Scaphocalanus amplius Park, 1970 (F,M) [Figs] | |
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Scaphocalanus angulifrons Sars, 1920 (F,M) (**) |
Ref.: | Sars, 1920 c (p.8, Rem.F); 1925 (p.170, Descr.F, figs.F) |
Rem.: | Cf. Falsilandrumius angulifrons. |
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[5] Scaphocalanus antarcticus Park, 1982 (F,M) [Figs] | |
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[6] Scaphocalanus australis Park, 1982 (F,M) [Figs] | |
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Scaphocalanus avia Tanaka, 1962 (M) (**) |
Syn.: | Scolecithricella avia Tanaka, 1962 (p.51) |
Rem.: | Transfert probable dans ce G. |
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Scaphocalanus bogorovi Brodsky, 1955 (F) (**) |
Ref.: | Brodsky, 1955 a (p.193, Descr.F, figs.F) |
Rem.: | Transfert probable à Scolecithricella ou à un nouveau genre. Cf. Falsilandrumius bogorovi |
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[7] Scaphocalanus brevicornis (Sars, 1900) (F,M) (*) [Figs] | |
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[8] Scaphocalanus brevirostris Park, 1970 (F,M) (*) [Figs] | |
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[9] Scaphocalanus californicus Davis, 1949 (F) (**) [Figs] | |
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[10] Scaphocalanus cristatus (Giesbrecht, 1895) (F,M) [Figs] | |
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[11] Scaphocalanus curtus (Farran, 1926) (F,M) (*) [Figs] | |
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[12] Scaphocalanus difficilis Roe, 1975 (F) (*) [Figs] | |
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[13] Scaphocalanus echinatus (Farran, 1905) (F,M) (*) [Figs] | |
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Scaphocalanus elegans Wolfenden, 1911 (F) (**) |
Syn.: | 1982 (p. 77); Bradford & al.,1983 (p.93)
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Loc: | Atlant. (off Is. St Pierre & St Paul NE) |
Rem.: | Probable dans ce G. |
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[14] Scaphocalanus elongatus A. Scott, 1909 (F,M) (**) [Figs] | |
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Rem.: | meso-bathypelagic. Sampling depth (Antarct., sub-Antarct.): 0-3000 m. Sargasso Sea: 500-2000 m (Deevey & Brooks, 1977, station "S"); 2000-1000 m (Harding, 1974). Distributional range (m) from Roe (1972) Day: 720-900, Night: 720-800; from Grice & Hulsemann (1965): 1500-4000 (in Kuriyama & Nishida, 2006). Genus not assured but probable. For Vervoort (1965, p.63), Tanaka (1961 a, p.166) has, probably correctly, referred Brodsky's male specimens of S. brevicornis to S. elongatus. S. elongatus from Hamahera Sea, appears uncrested, though a weak line on the head seems to have been present. Additional specimens were recorded by Sewell (1929) from the Bay of Bengal, but as these specimens differ in many respects from Scott's original specimen and from those recorded by Tanaka, their identification as S. elongatus is very doubtful. The specimen from the Gulf of Guinea found by Vervoort is the first from the Atlantic Ocean. First occurrence in Chilean waters by Hidalgo & al. (2010) |
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[15] Scaphocalanus emine Uysal & Shmeleva, 2002 (F) [Figs] | |
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[16] Scaphocalanus farrani Park, 1982 (F,M) [Figs] | |
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Scaphocalanus glacialis Tanaka, 1961 (M) |
Ref.: | Tanaka, 1953 (p.132 : nom. nud.); 1961a (p.177) |
Rem.: | Cf. Scaphocalanus curtus |
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Scaphocalanus gracilicauda Tanaka, 1937 (M) |
Syn.: | S. gracilicaudatus : Brodsky, 1950 (1967) (p.256, figs.M) |
Ref.: | Tanaka, 1937 (p.262) |
Rem.: | Cf. Scaphocalanus subbrevicornis |
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Scaphocalanus gracilis Wolfenden, 1911 (F) |
Syn.: | Amallophora gracilis Wolfenden, 1911 (p.265, Descr.F, figs.F); Sewell, 1948 (p.502); Bradford, 1973 (p.144); Bradford & al., 1983 (p.95, Rem.); Bradford & Wells, 1983 (p.2) |
Ref.: | Sewell, 1948 (p.502); Tanaka, 1961 a (p.,161, 163, Rem.); Park, 1982 (p.77, 79, 82: Rem.) |
Loc: | Atlant. S (off Tristan da Cunha) |
Lg.: | (10) F: 4,85 |
Rem.: | Cf. ?Scaphocalanus affinis ou magnus |
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[17] Scaphocalanus impar (Wolfenden, 1911) (F) (**) [Figs] | |
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Rem.: | Bathypelagic. Sampling depth (Antarct.) :1200 m. Genus poorly assured but probable. Geographical distribution uncertain. ? Cf. Scaphocalanus brevicornis and Scaphocalanus farrani. According to Tanaka (1961a, p.173) this species is closely allied to Scaphocalanus brevicornis Sars, but can be distinguished by the armature on P5 male. The present specimen has the P5 quite similar in structure to those figured by Wolfenden. |
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[18] Scaphocalanus insignis Brodsky, 1950 (F) (**) [Figs] | |
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[19] Scaphocalanus insolitus C.B. Wilson, 1950 (F) (**) [Figs] | |
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[20] Scaphocalanus invalidus Hure & Scotto di Carlo, 1968 (F) (*) [Figs] | |
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Scaphocalanus lobatus Sars, 1920 (F) (**) |
Syn.: | Scolecithricella lobata Sars, 1920 c (p.9) |
Ref.: | Bradford, 1973 (p.144: comme S. lobatus ) |
Rem.: | Possible dans ce G. |
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[21] Scaphocalanus longifurca (Giesbrecht, 1888) (F,M) (*) [Figs] | |
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Scaphocalanus macropes Tanaka, 1953 |
Ref.: | Tanaka, 1953 (p.132: nom. nud.); 1961 a (p.173) |
Rem.: | Cf. Scaphocalanus subbrevicornis |
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[22] Scaphocalanus magnus (T. Scott, 1894) (F,M) (*) [Figs] | |
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Scaphocalanus magnus major Sewell, 1947 (F) |
Ref.: | Sewell, 1947 (p.144, figs.F) |
Loc: | Mer Arabe S |
Lg.: | (11) F: 4,967 |
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Scaphocalanus magnus minor Sewell, 1947 (F) |
Ref.: | Sewell, 1947 (p.146, figs.F) |
Loc: | Mer Arabe, G. d’Oman |
Lg.: | (11) F: 3,55 |
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[23] Scaphocalanus major (T. Scott, 1894) (F,M) (*) [Figs] | |
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[24] Scaphocalanus medius (Sars, 1907) (F,M) [Figs] | |
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Scaphocalanus minutus Tanaka, 1937 (M) |
Syn.: | Scaphocalanus minuta Tanaka, 1937 (p.262) |
Rem.: | Cf. Scaphocalanus impar |
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Scaphocalanus obscurus Esterly, 1913 (F) (**) |
Syn.: | Scolecithrix obscura Esterly, 1913 (p.184) |
Rem.: | Transfert dans ce G. douteux, peut-être Lophothrix. |
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Scaphocalanus pacificus Mori, 1932 (F) |
Ref.: | Mori, 1932 a (p.169, figs.F); Mori, 1937 (1964) (p.49, figs.F) |
Loc: | Japon S |
Lg.: | (151) F: 2,6 |
Rem.: | Un doute subsiste sur la synonymie : Cf. Lophothrix latipes |
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[25] Scaphocalanus parantarcticus Park, 1982 (F,M) [Figs] | |
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[26] Scaphocalanus paraustralis Schulz, 1987 (F) [Figs] | |
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[27] Scaphocalanus polaris Brodsky, 1950 (F) (**) [Figs] | |
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Scaphocalanus profundus Brodsky, 1950 (M) (**) |
Syn.: | Amallothrix profunda Brodsky, 1950 (1967) (p.263) |
Rem.: | Possible dans le G. Scaphocalanus . ? Cf. Scolecithricella laminata. Voir à Amallothrix profunda |
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[28] Scaphocalanus pseudobrevirostris Schulz, 1987 (F) [Figs] | |
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Scaphocalanus similis Hure & Scotto di Carlo, 1968 (F) (*) |
Ref.: | Hure & Scotto di Carlo, 1968 a (p.157, Descr.F, figs.F); Bradford, 1973 (p.143); Park, 1982 (p.122, Rem.F, figs.F); Bradford & al., 1983 (p.93); Campaner, 1984 a (p.167, 169: tab.1); Scotto di Carlo & al., 1991 (p.271); Hure & Krsinic, 1998 (p.101) |
Loc: | Médit. (Mer d'Alboran, Adriatique S), Australie W (off Cap Leeuwin S) |
Lg.: | (533) F: 1,08-0,93 |
Rem.: | Cf. Scaphocalanus curtus |
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[29] Scaphocalanus somaliensis Vyshkvartzeva & Prusova, 1997 (F) [Figs] | |
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[30] Scaphocalanus subbrevicornis (Wolfenden, 1911) (F,M) (*) [Figs] | |
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Scaphocalanus subcurtus Park, 1970 (F) |
Ref.: | Park, 1970 (p.476, 499, Descr.F, figs.F) |
Ref. compl.: | Bradford, 1973 (p.143); Park, 1982 (p.77); Bradford & al., 1983 (p.93); Lozano Soldevilla & al., 1988 (p.59) |
Loc: | G. du Mexique, Caraïbes, Canaries, Médit. (in Kovalev & Shmeleva, 1982, p.84) |
Lg.: | (88) F: 1,06-0,96 |
Rem.: | Pour Bradford (1973), Bradford & al (1983) et Campaner, 1984 a (p.167, 169: tab.1) cette espèce serait synonyme de Scaphocalanus curtus (Farran, 1926) |
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[31] Scaphocalanus subelongatus Brodsky, 1950 (F) (**) [Figs] | |
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Scaphocalanus temporalis Tanaka, 1953 (M) |
Ref.: | Tanaka, 1953 (p.132 : nom. nud.); 1961 a (p.177, Rem.) |
Rem.: | Cf. Scaphocalanus longifurca |
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[32] Scaphocalanus vervoorti Park, 1982 (F,M) [Figs] | |
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[33] Scaphocalanus sp. Tanaka, 1961 [Figs] | |
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[34] Scaphocalanus sp. a Vidal, 1971 (F) [Figs] | |
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[35] Scaphocalanus sp. b Vidal, 1971 (F) [Figs] | |
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[36] Scaphocalanus sp. Roe, 1975 (M) [Figs] | |
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[37] Scaphocalanus sp. Park, 1982 (M) [Figs] | |
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(25) Scolecithricella Sars, 1902 | |
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Rem.: | Bradford & al. (1983) numbers 9 'safe' species (*) in this genus and 16 probable, 3 whose generic status would require to be clarified, and 2 congeneric species of a group : 'Scolecithrix ctenopus' near Scolecithricella. Type: Scolecithrix minor Brady, 1883. Provisional total 31 spp. + 6 undet.
Sano & al. (2013, p.23) note that the genus is considerd omnivore/detritivore in previous studies, and their chemo-sensory setae are considered to be involved in the detection of detritus (see in Nishida & Ohtsuka, 1997).
Diagnosis after Vyshkvartzeva (2000, p. 233): - Pediger somites 4 and 5 in female completely fused, more or less produced distally, narrowly or broadly rounded, sometimes with a incision near the dorsal side, in male separate, broadly rounded. - Rostrum bifurcated, both long proximal processes strong, sausage-shaped, tapering distally into sensory filaments, varying from short to 3/5 length of proximal part. - A1 female 20-23-segmented, 8-10th segments of the typical 25-segmented calanoid antennule fused ; 24th and 25th, 1st and 2nd, and 12th and 13th segments sometimes also fused, partly or completely. A1 male symmetrical , with 18-19-segmented (segments 8-12th, 20-21th completely fused, 24-25th sometimes partly fused. - A2 : endopod about 2/3 exopod. Exopodal segment 1 without seta or swelling ; exopdal segments 2-6 with 1 seta each in both sexes ; in female, seta of exopodal segment 2 sometimes shorter than in male. - Md endopod reaching the end of exopod ; basis with 1 inner seta reduced in male. - Mx1 : inner lobe 1 with 1 or 2 posterior setae ; inner lobe 3 with 3 setae ; exopod with 5, 6, 7 or 8 setae. - Mx2 : inner lobe 2 with 3 setae ; inner lobes 3-4 with 2-3 setae ; inner lobe 5 with 3 setae (one of them more stout) and usually with 1 worm-like sensory seta ; distal 3 segments of endopod with 3 worm-like and 5 brush-like sensory setae ; 2 of 5 brush-like setae shortyer, their brushes slightly larger. - P1 exopod with 2 outer spines (exopdal segment 1 without outer spine) ; spine of exopodal segment 2 not longer than 2/3 length of exopodal segment 3. - Outer distal corner of endopodal segment 1 of P2 produced distally into obtuse or acute spine-like process ; external spine of exopodal segment 1 of P2 usually long, more than half as long as exopodal segment 2. - P4 coxopod near outer distal margin with obtuse, knob-like process. Anterior and posterior surfaces of protopod, endopod and exopod with sparse small spines. P5 female uniramous, 1-segmented ; segment flat, attached tp common, very short coupler, armed with long inner spine (slightly shortyer, equal or slightly longer than segment), shorter apical spine and a minute outer spine, the latter frequently absent. P5 sometimes completely absent. - P5 male biramous, asymmetrical ; length of right and left legs subequal or right leg slightly shorter ; legs shorter than, about equal to or a little longer than urosome. Endopods of both legs short and rudimentary ; exopods usually 3-segmented, segments subcylindrical. In right leg, exopodal segments 1 and 2 partially or completely fused ; exopodal segment 2 with or more frequently without short medio-distal projection ; exopodal segment 3 usually with a minute seta distally, or tapering as a long spiniform process, or termibating with a characteristic grooved structure. Distal segment of left leg exopod usually the shortest, with spinules. Left endopod ginger-like, not longer than half of exopod, usually with a minute seta distally. For the author 11 species are included in this genus, and possibly 4 species belong to this genus but their descriptions are insufficient and usually males unknown : S. farrani (Rose, 1942), (= ? S. longipes) ; S. longipes Giesbredht, 1892 ; S. longispina Chen & Zhang, 1965 ; S. pearsoni Sewell, 1914 ; S. sarsi (Rose, 1942) ( = ? Scolecithricella globulosa) ; S. vespertina Tanaka, 1962.
Definition from Bradford, Haakonssen & Jillett (1983, p.102) : - Pedigerous segments 4 and 5 fused. - Rostrum of 2 filaments. - A1 22 or 23-segmented in female ; 19 segments in male. - Mx1 inner lobe 1 (arthrite) with 2 posterior surface setae ; inner lobe 3 (basal endite) with 3 setae ; endopod segment 1 usually separate from segments 2 and 3. - Mx2 endopod with 3 worm-like and 5 brush-like elements. - P1 exopod segment 1 usually without an external spine. - Male mouthparts slightly reduced compared with female. - Female P5 uniramous, 1-segmented, flattened, plate-like, attached to common basal segment. - Male P5 biramous on both sides, both endopods very short. |
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[1] Scolecithricella abyssalis (Giesbrecht, 1888) (F, M) (*) [Figs] | |
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Scolecithricella altera Farran, 1929 (F) |
Syn.: | Amallophora altera Farran, 1929 (p.252) |
Ref.: | Park, 1980 (p.70, Rem., figs.F); Vyshkvartzeva, 1989 (p.7, 16, Rem.) |
Rem.: | Cf. Mixtocalanus alter |
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Scolecithricella arcuata Sars, 1920 (F,M) |
Ref.: | Sars, 1920 c (p.10) |
Rem.: | Cf.: Amallothrix arcuata |
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Scolecithricella aspinosa Roe, 1975 (F) |
Rem.: | Cf.: Amallothrix aspinosa |
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Scolecithricella auropecten Giesbrecht, 1892 (F,M) |
Syn.: | Scolecithrix auropecten Giesbrecht, 1892 (p.266) |
Ref.: | Vyshkvartzeva, 1989 (p.5, 6, Rem.) |
Rem.: | Cf. Archescolecithrix auropecten |
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[2] Scolecithricella avia Tanaka, 1962 (M) (**) [Figs] | |
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Scolecithricella beata Tanaka, 1962 (F) |
Ref.: | Tanaka, 1962 (p.50); Bradford, 1973 (p.142) |
Rem.: | Cf. Scolecithricella tropica |
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Scolecithricella canariensis Roe, 1975 (F) |
Syn.: | Scolecithricella sp. Roe, 1972 (p.304, 310) |
Ref.: | Roe, 1975 (p.323, Descr.F, figs.F); Bradford & al., 1983 (p.103) |
Loc: | Is. du Cap Vert, off Canaries |
Lg.: | (8) F: 3,42-3,12 |
Rem.: | Cf. Pseudoamallothrix canariensis |
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Scolecithricella cenotelis Park, 1980 (F) (*) |
Ref.: | Park, 1980 (p.59, Descr.F, figs.F); Bradford & al., 1983 (p.103, 109: Rem.) |
Loc: | Antarct. , sub-Antarct. |
Lg.: | (523) F: 2,34-1,96 |
Rem.: | Cf. Pseudoamallothrix cenotelis |
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Scolecithricella ctenopus Giesbrecht, 1888 (F,M) |
Syn.: | Scolecithrix ctenopus Giesbrecht, 1888 |
Rem.: | Bradford (1973, p.146) considère cette forme comme le type d'un groupe d'espèces dont le genre, qui reste à définir, est intermédiaire entre Scolecithrix , Scolecithricella et Amallothrix . Voir à Scolecithrix ctenopus. Vyshkvartzeva, 1999 (2000) (p.219, 221) en fait le type d'un nouveau genre: Scolecitrichopsis. |
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[3] Scolecithricella curticauda A. Scott, 1909 (F) (**) [Figs] | |
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[4] Scolecithricella dentata (Giesbrecht, 1892) (F,M) (*) [Figs] | |
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[5] Scolecithricella denticulata Tanaka, 1962 (M) (**) [Figs] | |
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Scolecithricella dentipes Vervoort, 1951 (F,M) |
Syn.: | Scolecithricella (Amallothrix) dentipes Vervoort, 1951 (p.103) |
Rem.: | Cf. Amallothrix dentipes |
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Scolecithricella dubia Giesbrecht, 1892 (M) |
Syn.: | Scolecithrix dubia Giesbrecht, 1892 (p.266, fig.M); non Scolecithricella dubia : Tanaka, 1937 (p.261, Rem.) |
Rem.: | Cf. Scolecithricella dentata |
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Scolecithricella emarginata Farran, 1905 (F,M) |
Syn.: | Scolecithrix emarginata Farran, 1905 (p.36) |
Rem.: | Cf. Pseudoamallothrix emarginata |
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Scolecithricella farrani Rose, 1942 (F) (**) |
Syn.: | Amallothrix farrani Rose, 1942 (p.130) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.234) |
Rem.: | ? Cf.: Scolecithrix longipes. Probable dans le G. Scolecithricella . |
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[6] Scolecithricella globulosa Brodsky, 1950 (F,M) (**) [Figs] | |
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Scolecithricella gracilis Sars, 1905 (F,M) |
Ref.: | Sars, 1905 b (p.21) |
Rem.: | Cf. Amallothrix gracilis |
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[7] Scolecithricella grata Grice & Hulsemann, 1967 (F) (**) [Figs] | |
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Scolecithricella hadrosoma Park, 1980 (F) |
Ref.: | Park, 1980 (p.66) |
Rem.: | Cf. Amallothrix hadrosoma |
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Scolecithricella ingolfi With, 1915 (F) |
Ref.: | With, 1915 (p.207, Descr.F, figs.F); Sewell, 1948 (p.502); Bradford & al., 1983 (p.104) |
Loc: | Detr. de Davis, Islande |
Lg.: | (7) F: ? |
Rem.: | sp. douteuse, forme juvénile ? |
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Scolecithricella lamellifer Tanaka, 1962 (F) |
Ref.: | Tanaka, 1962 (p.78) |
Rem.: | Cf. Pseudoamallothrix laminata |
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Scolecithricella laminata Farran, 1926 (F) |
Syn.: | Scolecithrix laminata Farran, 1926 (p.265, Descr.F, figs.F) |
Rem.: | Cf.: Pseudoamallothrix laminata |
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[8] Scolecithricella lanceolata Tanaka, 1962 (M) (**) [Figs] | |
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Scolecithricella lobata Sars, 1920 (F) |
Syn.: | Amallothrix lobata : Sars, 1925 (p.184, Descr.F, figs.F) |
Ref.: | Sars, 1920 c (p.9, Rem.F) |
Rem.: | Cf. Falsilandrumius lobatus |
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Scolecithricella lobophora Park, 1970 (F,?M) |
Syn.: | Amallothrix lobophora : Roe, 1975 (p.329, Descr. M, figs.M, Rem.); Vives, 1982 (p.292) |
Ref.: | Park, 1970 (p.511, Descr.F, figs.F); Vyshkvartzeva, 1999 (2000) (p.235) |
Rem.: | Cf. Amallothrix lobophora. |
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Scolecithricella longipes Giesbrecht, 1892 (F) (**) |
Syn.: | Scolecithrix longipes Giesbrecht, 1892 (p.266) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.234) |
Rem.: | Probable dans le G. Scolecithricella |
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[9] Scolecithricella longispinosa Chen & Zhang, 1965 (F) (**) [Figs] | |
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[10] Scolecithricella marginata (Giesbrecht, 1888) (F) [Figs] | |
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[11] Scolecithricella maritima Grice & Hulsemann, 1967 (F) (**) [Figs] | |
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Rem.: | Family and Genus to reconsider. For Grice & Hulsemann (1967) this species is similar to Scolecithrix ctenopus Giesbrecht (1888, 1892), but the two may be distinguished. |
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Scolecithricella marquesae Vervoort, 1965 (F) |
Syn.: | Scolecithricella (Amallothrix) marquesae Vervoort, 1965 (p.74, Descr.F, figs.F, Rem) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.221, Rem.) |
Rem.: | Probable dans le Groupe “ Scolecithrix ctenopus “. Cf. Scolecitrichopsis tenuipes |
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[12] Scolecithricella minor (Brady, 1883) (F,M) (*) [Figs] | |
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Scolecithricella minor occidentalis Brodsky, 1950 (F,M) |
Ref.: | Brodsky, 1950 (1967) (p.270, fig.178 b); Vervoort, 1965 (p.83); Park, 1980 (p.35: Rem.); Bradford & al., 1983 (p.109); Sirenko & al., 1996 (p.349); Kosobokova & al., 1998 (tab.2); Kosobokova & Hirche, 2000 (p.2029, tab.2) |
Loc: | Antarct., Atlant.N, Arct. (Sibérie, Mer de Laptev, Lomonosov Ridge, Mer de Chukchi), Nouvelle-Ecosse |
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Scolecithricella minor orientalis Brodsky, 1950 (F,M) |
Ref.: | Brodsky, 1950 (1967) (p.270, fig.178 a); Vervoort, 1965 (p.83); Vaupel-Klein, 1970 (p.20); Bradford & al., 1983 (p.109) |
Loc: | Japon, Pacif. NW, Mer de Béring, Station 'P' |
Lg.: | (205) F: 1,3; M: 1,3 |
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[13] Scolecithricella modica Tanaka, 1962 (F) (**) [Figs] | |
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Scolecithricella neptuni Cleve, 1904 (F,M) (**) |
Ref.: | Cleve, 1904 a (p.197, 206, figs.F,M); Bradford, 1973 (p.147, Rem); Bradford & al., 1983 (p.103,123) |
Loc: | Afr. S (W) |
Lg.: | (12) F: 1,4; M: 1,2 |
Rem.: | mésopélagique. Présente des affinités avec Scolecithricella, mais statut mal défini. Peut-être appartenance aux Tharybidae. Cf. Tharybis neptuni. |
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[14] Scolecithricella nicobarica (Sewell, 1929) (F,M) (**) [Figs] | |
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[15] Scolecithricella obscura Roe, 1975 (F) (**) [Figs] | |
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[16] Scolecithricella orientalis Mori, 1937 (F,M) (**) [Figs] | |
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Scolecithricella ovata Farran, 1905 (F,M) (*) |
Syn.: | Scolecithrix ovata Farran, 1905 (p.37, Descr.F, figs.F) |
Rem.: | épipélagique à bathypélagique. Cf. Pseudoamallothrix ovata |
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[17] Scolecithricella pacifica Chiba, 1956 (F) (**) | |
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Scolecithricella parafalcifer Park, 1980 (F) |
Ref.: | Park, 1980 (p.50) |
Rem.: | Cf. Amallothrix parafalcifer |
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[18] Scolecithricella paramarginata Schulz, 1991 (F,M) (*) [Figs] | |
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Scolecithricella paravalida Brodsky, 1950 (F) |
Syn.: | Amallothrix paravalida : Brodsky, 1950 (1967) (p.263, Descr.f, figs.F); Scolecithricella (Amallothrix) paravalida : Vervoort, 1965 (p.68, Rem.) |
Ref.: | Tanaka, 1962 (p.73, figs.F, Rem.); Bradford, 1973 (p.143); Bradford & al., 1983 (p.78, Rem.) |
Rem.: | Les auteurs ne sont pas en accord sur les synonymies; Pour Park (1970, p.511) cette forme est un synonyme junior de Scolecithricella valens (= Scolecithix valens Farran, 1926) contrairement à Tanaka (1962). Cf. Amallothrix paravalida |
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[19] Scolecithricella pearsoni Sewell, 1914 (F,M) (**) [Figs] | |
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[20] Scolecithricella profunda (Giesbrecht, 1892) (F,M) (*) [Figs] | |
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[21] Scolecithricella propinqua (Sars, 1920) (F,M) (**) [Figs] | |
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Scolecithricella pseudoarcuata Park, 1970 (F) |
Ref.: | Park, 1970 (p.511, figs.F) |
Rem.: | Cf. Amallothrix pseudoarcuata |
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Scolecithricella pseudoculata Campaner, 1979 (F) |
Ref.: | Campaner, 1979 (p.82, Descr.F, figs.F) |
Rem.: | Probable dans le G ' Scolecithrix ctenopus '. Cf. Scolecitrichopsis pseudoculata |
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Scolecithricella pseudopropinqua Park, 1980 (F,M) |
Ref.: | Park, 1980 (p.51) |
Rem.: | Cf. Amallothrix pseudopropinqua |
|
Scolecithricella sarsi Rose, 1942 (F) (**) |
Syn.: | Amallothrix sarsi Rose, 1942 (p.132) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.234, Rem.) |
Rem.: | Probable dans le G. Scolecithricella |
|
Scolecithricella schizosoma Park, 1980 (F,M) (*) |
Ref.: | Park, 1980 (p.43, figs.F,M); Bradford & al., 1983 (p.103, 111, Rem., figs.F,M); Vyshkvartzeva, 2000 (p.234, Rem.) |
Loc: | Antarct., sub-Antarct. |
Lg.: | (523) F: 2,16-1,66; M: 2,28-2,1 |
Rem.: | Cf. Scolecithricella globulosa |
|
Scolecithricella similis Wolfenden, 1904 (F) |
Syn.: | Scolecithrix similis Wolfenden,1904 (p.119, Descr.F, figs.F); Van Breemen, 1908 a (p.73, Rem.F, fig.F); Rose, 1942 (p.149: Rem.) |
Ref.: | Lysholm & al., 1945 (p.28) |
Loc: | Féroé, Atlant.N, Canaries |
Lg.: | (229) F: 1,5 |
Rem.: | Probable dans le genre Scolecithricella. Cf. ? |
|
[22] Scolecithricella spinacantha C.B. Wilson, 1942 (F) (**) [Figs] | |
|
|
[23] Scolecithricella spinata Tanaka, 1962 (F) (**) [Figs] | |
|
|
Scolecithricella spinipedata Mori, 1937 (F) |
Ref.: | Mori, 1937 (1964) (p.53) |
Rem.: | Cf. Scolecithrix ctenopus |
|
Scolecithricella subdentata Esterly, 1905 (F) |
Syn.: | Scolecithrix subdentata Esterly, 1905 (p.267) |
Rem.: | Probable dans le G. Scolecithricella . Cf. Scolecithricella ovata |
|
Scolecithricella subvittata Rose, 1942 (F) |
Ref.: | Rose, 1942 (p.142); Mazza, 1966 (p.70) |
Rem.: | Hure & Krsinic (1998, p.49, 101) n’admettent pas la synonymie. Cf. Scolecithricella vittata |
|
[24] Scolecithricella tenuiserrata (Giesbrecht, 1892) (F,M) (**) [Figs] | |
|
|
[25] Scolecithricella timida Tanaka, 1962 (F) (**) [Figs] | |
|
|
[26] Scolecithricella tropica Grice, 1962 (F) (**) [Figs] | |
|
|
[27] Scolecithricella tydemani A. Scott, 1909 (F) (**) [Figs] | |
|
|
[28] Scolecithricella unispinosa Grice & Hulsemann, 1965 (F) (**) [Figs] | |
|
|
Scolecithricella vervoorti Park, 1980 (F,M) |
Syn.: | Amallophora altera Vervoort,1957 (p.94, figs.F) |
Rem.: | Cf. Mixtocalanus vervoorti |
|
[29] Scolecithricella vespertina Tanaka, 1955 (F) (**) [Figs] | |
|
|
[30] Scolecithricella vittata (Giesbrecht, 1892) (F,M) (*) [Figs] | |
|
|
Scolecithricella sp. Dakin & Colefax, 1940 |
Ref.: | Dakin & Colefax, 1940 (p.96, figs.F,M) |
Rem.: | Cf. Parundinella dakini |
|
[31] Scolecithricella sp. Tanaka, 1962 (M) [Figs] | |
|
|
[32] Scolecithricella sp. Grice, 1962 (M) [Figs] | |
|
|
[33] Scolecithricella sp. Grice & Hulsemann, 1967 (M) [Figs] | |
|
|
[34] Scolecithricella sp.1 Park, 1980 (M) (**) [Figs] | |
|
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[35] Scolecithricella sp.2 Park, 1980 (M) (**) [Figs] | |
|
|
[36] Scolecithricella sp.3 Park, 1980 (M) (**) [Figs] | |
|
|
(26) Scolecithrix Brady, 1883 | |
|
Rem.: | Type: Undina danae Lubbock,1856. Bradford & al. (1983, p.113) include 2 "sure" spp. (*), more 1 likely (**) in this genus. Transfers probable in other genera (***). Provisional total 13 spp.
Defintion from Bradfoird & al. (1983, p.112) : - Pedigerous segments 4 and 5 usually separate. - A1 female 19 or 20 segments ; A1 male 17-19 segments.. - Mx1 inner lobe (arthrite) with 1 seta on posterior surface ; endopod with all joinys fused. - P1 exopod segment 1 with or without external spine. - Mouthparts only very slightly reduced compared with female. - Female P5 absent or, if present, small and asymmetrical. - Male P5 tends to be uniramous on right ; biramous on left side.
Diagnosis from Park (1983, p.166) : Female: - Cephalosome and pediger 1 fused, pedigers 4 and 5 distinctly separate. - Urosome 4-segmented - Small caudal rami. - Rostrum biramous, eavh ramus tapering into a long filament. - A1 with 1st and 2nd, 8th through 12th segments fused, resulting in 20 free segments. - A2 with both rami well developed, exopod slightly longer than endopod, 1st exopodal segment without a seta. - Md with strong masticatory blade and roughly triangular basis bearing small 2-segmented endopod and large 5-segmented exopod. - Mx1 with 3 inner lobes and 1 outer lobe on coxa and endopod and exopod on basis. Endopod unsegmented and fused with basis. - Mx2 with 5 well-developed basipodal lobes and a small endopod. Endopod with 3 vermiform and 5 brush-form sensory filaments. - Mxp with strong coxa and basis followed by 5-segmented endopod. Coxa with a short brush-form sensory filament and 2 proximal setae modified into vermiform sensory filaments. P1 with 1-segmented endopod and 3-segmented exopod. Endopod produced distally into a prominent spiniform process along outer margin. 1st exopodal segment with or without an outer spine; 2nd and 3rd each with an outer spine. - P2 to P4 similar with 3-segmented exopod terminated with a strong spine. Basis with a prominent spiniform process distally along inner margin. Endopod produced distally into a sharp spiniform process along outer margin. 1st and 2nd exopodal segment each with an inner seta and an outer spine; 3rd with 4 inner setae and 3 outer spine of equal size. - Coxa with an inner seta in P2 and P3. - Endopod 2-segmented in P2 but 3-segmented in P3 and P4. - Posterior surface armed with strong spinules in P2 and P3, and with small spinules in P4. - P5 absent or if present small, unsegmented, and asymmetrical.
Male: - Body similar to that of female. - Cephalosome and pediger 1 fused, pedigers 4 and 5 distinctly separate. - Urosome 5-segmented, 5th segment (anal somite) almost telescoped in preceding segment. - Rostrum biramous , each ramus tapering into a long filament as in female. - A1 with 8th through 12th segments fused as in female, but 1st and 2nd separate; 20th and 21th separate on left but fused on right. - A2, Md, Mx1, Mx2 and Mxp agree in morphological details with those of female and almost as well developed as in female. P1 to P4 similar to those of female except that outer exopodal spines relatively small; - P5 asymmetrical each with 2-segmented basipod followed in left leg by 3-segmented exopod and 1-segmented endopod and in right only by 2-segmented exopod. |
|
Scolecithrix abyssalis Giesbrecht, 1888 (F,M) |
Ref.: | Giesbrecht, 1888; 1892 (p.266, figs.F); Bradford, 1973 (p.142) |
Rem.: | Cf. Scolecithricella abyssalis |
|
[1] Scolecithrix aculeata Esterly, 1913 (F) (***) [Figs] | |
|
|
Scolecithrix acutus Wolfenden, 1911 (F) |
Ref.: | Wolfenden, 1911 (p.253); Farran, 1926 (p.266); Vervoort, 1965 (p.61) |
Rem.: | Cf. Lophothrix latipes |
|
[2] Scolecithrix aequalis Wolfenden, 1911 (F) (***) [Figs] | |
|
|
Scolecithrix ancorarum Oliveira, 1946 (F) |
Ref.: | Oliveira, 1946 (p.460, figs.F); Bradford & al., 1983 (p.113) |
Rem.: | Cf. Paracalanus parvus |
|
Scolecithrix angusta Esterly, 1911 (F) |
Ref.: | Esterly, 1894 b (p.52) |
Rem.: | Cf. Lophothrix latipes |
|
Scolecithrix atlanticus Wolfenden, 1904 (F) |
Ref.: | Van Breemen, 1908 a (p.75, Rem.F) |
Rem.: | Cf. Brachycalanus atlanticus |
|
Scolecithrix auropecten Giesbrecht, 1892 (F) |
Ref.: | Giesbrecht, 1892 (p.266) |
Rem.: | Cf. Archescolecithrix auropecten |
|
Scolecithrix birshteini Brodsky, 1955 (F) |
Ref.: | Brodsky, 1955 a (p.198, Descr.F, figs.F); Bradford, 1973 (p.141); Bradford & al., 1983 (p.113); Park, 1983 (p.166) |
Loc: | Kouriles-Kamtchatka |
Lg.: | (27) F: 2,62-2,1 |
Rem.: | Cf. Pseudoamallothrix birshteini |
|
Scolecithrix birshteini major Brodsky, 1955 (F) |
Ref.: | Brodsky, 1955 a (p.199, Rem.F, figs.F) |
Rem.: | Cf. Pseudoamallothrix profunda |
|
Scolecithrix birshteini minor Brodsky, 1955 (F) |
Ref.: | Brodsky, 1955 a (p.199, Rem.F, figs.F) |
|
[3] Scolecithrix bradyi Giesbrecht, 1888 (F,M) (*) [Figs] | |
|
|
Scolecithrix brevicornis Sars, 1900 (F) |
Ref.: | Sars, 1900 (p.46) |
Rem.: | Cf. Scaphocalanus brevicornis |
|
Scolecithrix chelifer Thompson, 1903 (juv. M) |
Ref.: | I.C. Thompson, 1903 a (p.21, Descr. juv.M, figs.M); Bradford, 1983 (p.61) |
Loc: | off Irlande W |
Rem.: | Cf. Cornucalanus chelifer |
|
Scolecithrix chelipes Giesbrecht, 1896 (M) |
Ref.: | Giesbrecht, 1896 (1897) (p.321) |
Rem.: | Cf. Macandrewella chelipes |
|
Scolecithrix cristata Giesbrecht, 1895 (F) |
Ref.: | Giesbrecht, 1895 c (p.252) |
Rem.: | Cf. Scaphocalanus magnus |
|
Scolecithrix ctenopus Giesbrecht, 1888 (F,M) |
Ref.: | Giesbrecht, 1892 (p.266, 285, Descr.M, figs.M) |
Rem.: | Bradford (1973) considère cette forme comme le type d'un groupe d'espèces dont le genre, qui reste à définir, est intermédiaire entre Scolecithrix , Scolecithricella et Amallothrix . Vyshkvartzeva (1999 (2000), p.219) en fait le type du nouveau genre Scolecitrichopsis. |
|
Scolecithrix cuneifrons Willey, 1918 (F,M) |
Ref.: | Willey, 1918 (1919) (p.194); Farran, 1936 a (p.102) |
Rem.: | Cf. Scottocalanus securifrons |
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Scolecithrix curta Farran, 1926 (F) |
Ref.: | Farran, 1926 (p.259) |
Rem.: | Cf. Scaphocalanus curtus |
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[4] Scolecithrix danae (Lubbock, 1856) (F,M) (*) [Figs] | |
|
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Scolecithrix dentata Giesbrecht, 1892 (F,M) |
Ref.: | Giesbrecht, 1892 (p.266) |
Rem.: | Cf. Scolecithricella dentata |
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Scolecithrix dubia Giesbrecht, 1892 (M) |
Ref.: | Giesbrecht, 1892 (p.266); De Decker & Mombeck, 1964 (p.14) |
Rem.: | Tanaka (1962, p.44, 45) et Campaner (1984 a, p.176) ne sont pas en accord sur la synonymie. Cf. Scolecithricella dentata |
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Scolecithrix echinata Farran, 1905 (F) |
Ref.: | Farran, 1905 (p.37) |
Rem.: | Cf. Scaphocalanus echinatus |
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[5] Scolecithrix elephas Esterly, 1913 (F) (**) [Figs] | |
|
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Scolecithrix emarginata Farran, 1905 (F) |
Ref.: | Farran, 1905 (p.36) |
Rem.: | Cf. Amallothrix emarginata |
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Scolecithrix falcifer Farran, 1926 (F) |
Ref.: | Farran, 1926 (p.262) |
Rem.: | Cf. Amallothrix falcifer |
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Scolecithrix fowleri Farran, 1926 (F,M) |
Rem.: | Cf. Bradfordiella fowleri |
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Scolecithrix fultoni T & A. Scott, 1898 (F,M) |
Rem.: | Cf. Stephos fultoni |
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Scolecithrix glacialis Giesbrecht, 1902 (F,M) |
Ref.: | Giesbrecht, 1902 (p.25) |
Rem.: | Cf. Scolecithricella minor |
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Scolecithrix globiceps Farran, 1908 (F) |
Ref.: | Farran, 1908 b (p.54) |
Rem.: | Cf. Amallothrix gracilis |
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Scolecithrix gracilipes Farran, 1908 (F) |
Ref.: | Farran, 1908 b (p.52, figs.F) |
Loc: | Irlande W |
Rem.: | Cf. Scaphocalanus major ou Scaphocalanus brevicornis |
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Scolecithrix hibernica A. Scott, 1896 (F,M) |
Ref.: | Vervoort, 1965 (p.87) |
Rem.: | Cf. Diaixis hibernica |
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Scolecithrix incisa Farran, 1929 (F) |
Syn.: | Scolecithricella incisa : Vervoort,1965 b (p.397) |
Ref.: | Farran, 1929 (p.244, Descr.F, figs.F); Sewell, 1948 (p.574); Bradford, 1973 (p.143); Bradford & al., 1983 (p.78) |
Loc: | Antarct. (Cap Evans) |
Lg.: | f. Pseudoamallothrix incisa
|
|
Scolecithrix inornata Esterly, 1906 (F,M) |
Syn.: | Amallothrix inornata : Davis, 1949 (p.44, figs.F); Brodsky, 1950 (1967) (part., p.260, Descr.F,M, figs.F,M, Rem.); Vaupel Klein, 1970 (p.20); Morioka, 1972 a (p.314); Marlowe & Miller, 1975 (p.839); Vives, 1982 (p.282); Yamaguchi & al., 2002 (p.1007, tab.1) |
Ref.: | Esterly, 1906 a (p.67, Descr.F, figs.F); Bradford, 1973 (p.143, Rem.); Bradford & al., 1983 (p.78, Rem.) |
Loc: | off Mauritanie (in Vives, 1982, p.292), off Hokkaido SE, Pacif. N, Station Knot, Mer d'Okhotsk, Station 'P', San Diego |
Lg.: | (17) F: 4,3; (22) F: 4,3-3,8; M: 3,7-3,6; (59) F: 4,3-4,2; (205) F: 3,9 |
Rem.: | Synonyme probable de Amallothrix emarginata . Cf. Pseudoamallothrix inornata |
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Scolecithrix laminata Farran, 1926 (F) |
Ref.: | Farran, 1926 (p.265, Descr.F, figs.F) |
Rem.: | Cf. Pseudoamallothrix laminata |
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Scolecithrix latipes T. Scott, 1894 (F,M) |
Ref.: | T. Scott, 1894 b (p.52) |
Rem.: | Cf. Lophothrix latipes |
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Scolecithrix longicornis T. Scott, 1894 (F) |
Ref.: | T. Scott, 1894 b (p.50) |
Rem.: | Cf. Scolecithrix ctenopus |
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Scolecithrix longifurca Giesbrecht, 1888 (F) |
Ref.: | Giesbrecht & Schmeil, 1898 (p.45) |
Rem.: | Cf. Scaphocalanus longifurca |
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[6] Scolecithrix longipes Giesbrecht, 1892 (F) (**) [Figs] | |
|
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Scolecithrix longirostris Esterly, 1913 (F) |
Syn.: | Scolecithricella longirostris : Sewell, 1948 (p.557, 563) |
Ref.: | Esterly, 1913 (p.185, Dsecr.F, figs.F); Brodsky, 1950 (1967) (p.273, figs.F); Bradford & al., 1983 (p.113, Rem.) |
Ref. compl.: | Suarez-Morales & Gasca, 1998 a (p.111) |
Loc: | Californie |
Lg.: | (144) F: 2,07 |
Rem.: | Forme juvénile ?, sp. douteuse. |
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[7] Scolecithrix magnus Wolfenden, 1911 (F) (**) [Figs] | |
|
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Scolecithrix major T. Scott, 1894 (F) |
Ref.: | T. Scott, 1894 b (p.52) |
Rem.: | Cf. Scaphocalanus major |
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Scolecithrix marginata Giesbrecht, 1888 (F) |
Ref.: | Giesbrecht, 1888; 1892 (p.266, 285, figs.F); Vyshkvartzeva, 1999 (2000) (p.234) |
Rem.: | Cf. Scolecithricella marginata. |
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[8] Scolecithrix medius Wolfenden, 1911 (F) (**) [Figs] | |
|
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Scolecithrix minor Brady, 1883 (F,M) |
Ref.: | Brady, 1883 (p.58) |
Rem.: | Cf. Scolecithricella minor |
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[9] Scolecithrix mollis Esterly, 1913 (F) (**) [Figs] | |
|
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Scolecithrix nicobarica Sewell, 1929 (F,M) |
Ref.: | Sewell, 1929 (p.209, figs.F,M); Vyshkvartzeva, 1999 (2000) (p.234) |
Rem.: | Genre mal assuré. Cf. Scolecithricella nicobarica |
|
[10] Scolecithrix obscura Esterly, 1913 (F) (**) [Figs] | |
|
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Scolecithrix ovata Farran, 1905 (F) |
Ref.: | Farran, 1905 (p.37) |
Rem.: | Cf. Pseudoamallothrix ovata |
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[11] Scolecithrix pacifica Esterly, 1905 (F) (**) [Figs] | |
|
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Scolecithrix persecans Giesbrecht, 1895 (F,M) |
Ref.: | Giesbrecht, 1895 c (p.253) |
Rem.: | Cf. Scottocalanus persecans |
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Scolecithrix polaris Wolfenden, 1911 (F,M) |
Ref.: | Wolfenden, 1911 (p.252); Farran, 1929 (p.208, 243) |
Rem.: | Pour Brodsky, 1950 (1967, p.260) cette espèce est synonyme de Scolecithrix inornata contrairement à Park, 1980 (p.61, 66) suivi par Bradford & al., 1983 (p.82). Cf. Amallothrix emarginata |
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[12] Scolecithrix porrecta Giesbrecht, 1888 (F) (**) [Figs] | |
|
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Scolecithrix profunda Giesbrecht, 1892 (F,M) |
Ref.: | Giesbrecht, 1892 (p.266) |
Rem.: | Cf. Scolecithricella profunda |
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Scolecithrix pygmaea T. Scott, 1899 (F,M) |
Rem.: | Cf. Diaixix pygmaea |
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Scolecithrix (Amallophora) robusta T. Scott, 1894 (F) |
Ref.: | T. Scott, 1894 b (p.56) |
Rem.: | Cf. Amallothrix robusta |
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Scolecithrix römeri Mrazek, 1902 (F,M) |
Ref.: | Mrazek, 1902 (p.522) |
Rem.: | Cf. Scolecithricella minor |
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Scolecithrix scotti Giesbrecht, 1893 (M) |
Ref.: | Giesbrecht, 1897 b (p.254) |
Rem.: | Cf. Scopalatum dubia |
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Scolecithrix securifrons T. Scott, 1894 (F,M) |
Ref.: | T. Scott, 1894 b (p.47, figs.F,M); Cleve, 1904 a (p.197) |
Rem.: | Cf. Scottocalanus securifrons (F) & Scottocalanus helenae (M) |
|
Scolecithrix similis T. Scott, 1894 (M) |
Syn.: | Scolecithrix ( Amallophora ) dubia var. similis T. Scott, 1894 b (p.56); Giesbrecht & Schmeil, 1898 (p.46) |
Rem.: | Cf. Scaphocalanus major |
|
Scolecithrix similis Wolfenden, 1904 (F) |
Syn.: | Scolecithricella similis : Lysholm & al., 1945 (p.28) |
Ref.: | Wolfenden,1904 (p.119, Descr.F, figs.F); Van Breemen, 1908 a (p.73, Rem.F, fig.F); Rose, 1942 (p.149: Rem.) |
Loc: | Féroé, Atlant.N, Canaries |
Lg.: | (229) F: 1,5 |
Rem.: | Probable à Scolecithricella. Cf. ? |
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Scolecithrix subdentata Esterly, 1905 (F) |
Ref.: | Esterly, 1905 (p.167) |
Rem.: | Cf. Scolecithricella ovata |
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Scolecithrix tenuipes T. Scott, 1894 (M) |
Syn.: | Scolecithricella tenuipes : A. Scott, 1909 (p.92, Rem.M); Sewell, 1948 (p.516, 517, 523, 546) |
Ref.: | T. Scott, 1894 b (p.48, Descr.M, figs.M); Giesbrecht & Schmeil, 1898 (p.47, Rem.M); Thompson & Scott, 1903 (p.234, 245); Bradford & al., 1983 (p.104) |
Loc: | Congo, G. de Guinée, Mer Rouge, Ceylan, Indonésie-Malaisie (in A. Scott, 1909) |
Lg.: | (57) M: 1,4 |
Rem.: | Probable dans le G. ' Scolecithrix ctenopus '. Cf. Scolecitrichopsis tenuipes |
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Scolecithrix tenuiserrata Giesbrecht, 1892 (F,M) |
Ref.: | Giesbrecht, 1892 (p.266, 284, figs.F,M); Vyshkvartzeva, 1999 (2000) (p.234) |
Rem.: | Probable dans le G. Scolecithricella . |
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Scolecithrix tolli Jashnov, 1947 |
Rem.: | Cf. Derjuginia tolli |
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Scolecithrix tumida T. Scott, 1894 (F) |
Ref.: | T. Scott, 1894 b (p.52) |
Rem.: | Cf. Scolecithricella abyssalis |
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[13] Scolecithrix valens Farran, 1926 (F) (**) [Figs] | |
|
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Scolecithrix valida Farran, 1908 (F) |
Ref.: | Farran, 1908 b (p.55, Descr.F, figs.F); 1929 (p.208, 244, Rem.); Vyshkvartzeva, 1999 (2000) (p.235, Rem.); Razouls & al., 2000 (p.343, Appendix) |
Lg.: | (35) F: 4,06 |
Rem.: | bathypélagique. Probable dans le G. Amallothrix . Localisations incertaines du fait d’une confusion possible par certains auteurs entre cette espèce et Amallothrix paravalida et Scolecithrix valens. La forme sous cette dénomination signalée en Antarctique par Farran (1929) ne correspond à aucune des deux espèces citées précédemment. |
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Scolecithrix vittata Giesbrecht, 1892 (F,M) |
Ref.: | Giesbrecht, 1892 (p.266); Vyshkvartzeva, 1999 (2000) (p.234) |
Rem.: | Cf. Scolecithricella vittata |
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Scolecithrix vorax Esterly, 1911 (F) |
Ref.: | Esterly, 1911 (p.327) |
Rem.: | Cf. Scopalatum vorax |
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" Scolecithrix ctenopus “ Group: Scolecitrichopsis |
|
Ref.: | Sars, 1925 (p.175, Rem.); Bradford, 1973 (p.146, Déf.); Campaner, 1979 (p.86); Bradford & al., 1983 (p.103, 109, Rem.); Vyshkvartzeva, 1999 (2000) (p.218, Rem.) |
Rem.: | type Scolecithrix ctenopus Giesbrecht,1888. Bradford & al., 1983 considèrent que les 10 espèces suivantes pourraient appartenir à ce goupe : |
|
Scolecithrix ctenopus Giesbrecht, 1888 (F,M) |
Ref.: | Giesbrecht, 1892 (p.266) |
Rem.: | Cf.: Scolecitrichopsis ctenopus. |
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Scolecithrix magnus Wolfenden, 1911 (F) |
Ref.: | Wolfenden, 1911 (p.258) |
Rem.: | Probable à Amallothrix |
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Scolecithricella (Amallothrix) marquesae Vervoort, 1965 (F) |
Ref.: | Vervoort, 1965 (p.74) |
Rem.: | Non in Scolecithricella, non in Amallothrix. Cf. Scolecitrichopsis tenuipes |
|
Scolecithricella pseudoculata Campaner, 1979 (F) |
Ref.: | Campaner, 1979 (p.82) |
Rem.: | Cf. Scolecitrichopsis pseudoculata |
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Scolecithricella spinacantha Wilson, 1942 (F) |
Ref.: | C.B. Wilson, 1942 a (p.208) |
Rem.: | Probable dans ce groupe. Probale comme Scolecitrichopsis spinacantha |
|
Scolecithrix tenuipes T. Scott, 1894 (M) |
Ref.: | T. Scott, 1894 b (p.48) |
Rem.: | Probable dans ce groupe. Cf. Scolecitrichopsis tenuipes |
|
Xanthocalanus alvinae Grice & Hulsemann, 1970 (F) |
Ref.: | Grice & Hulsemann, 1970 (p.189) |
Rem.: | Probable dans ce groupe. Cf. Scolecitrichopsis alvinae |
|
Xanthocalanus difficilis Grice & Hulsemann, 1965 (F) |
Ref.: | Grice & Hulsemann, 1965 (p.235) |
Rem.: | Non Xanthocalanus . Probable dans ce groupe. Probable comme Scolecitrichopsis difficilis |
|
Xanthocalanus distinctus Grice & Hulsemann, 1970 (M) |
Ref.: | Grice & Hulsemann, 1970 (p.190) |
Rem.: | Probable dans ce groupe. Cf. Scolecitrichopsis distinctus |
|
Xanthocalanus elongatus Grice & Hulsemann, 1970 (F,M) |
Ref.: | Grice & Hulsemann, 1970 (p.190) |
Rem.: | Non Xanthocalanus . Probable dans ce groupe. Cf. Scolecitrichopsis elongatus |
|
" Scolecithrix fowleri " Group: |
|
Ref.: | Bradford, 1973 (p.146, Def.) |
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Scolecithrix fowleri Farran, 1926 (F,M) |
Ref.: | Farran, 1926 (p.261) |
Rem.: | Probable dans ce Groupe |
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Scolecithrix grata Grice & Hulsemann, 1967 (F) |
Ref.: | Grice & Hulsemann, 1967 (p.27) |
Rem.: | Genre et famille à préciser. Provisoirement dans ce Groupe |
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(27) Scolecitrichopsis Vyshkvartzeva, 2000 | |
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Rem.: | type: Scolecithrix ctenopus Giesbrecht,1888. Total: 7 spp. [+ 3 probable].
Diagnosis from Vyshkvartzeva (2000, p.219) : - Pedigerous somites 4 end 5 separate or fused (if fused, , a well marked notch on ventro-lateral margin indicates the border of each somite). - Posterolateral corners of last prosomal somite produced distally, triangular, sometimes with a spine-like process. Rostrum as a short plate with 2 thin filaments (in S. difficilis, conical, without filaments). - A1 female 24-segmented (8th and 9th segments of the ancestral fused). A1 male asymmetrical, left one 21-segmented (8-12th segments fused), right one 20-segmented (additional to those of left, 20th and 21th segments fused). - A2 exopod of female with 4 medial setae (in segments 3-4), of male with 5 medial setae (in segments 2-6). Exopod slightly longer than endopod. - Mx1: inner lobe 1 with 1-2 posterior setae; inner lobe 3 with 2-3 setae; exopod usually with 7 setae (in S. tenuipes with 6 setae; in S. difficilis with 10 setae). - Mx2: endopod distally with 3 worm-like and 5 small, subequal, brush-like sensory filaments; inner lobe 5 with 3 sclerotized setae (frequently 2 of them stout, hook-like, with denticles) and 1 worm-like sensory seta or with 2 sclerotized and 2 sensory setae; on inner lobes 2-4 sometimes 1 or 3 sclerotized setae transformed into worm-like one. (Mx2 of S. difficilis wiyh differing armament. - Male mouthparts almost not reduced compared with those of female.. P1: exopod al segments 1-3 usually with long external spine each (S. ctenopus without external spine on exopodal segment 1). - Outer distal corner of endopodal segment 1 of P2 not produced, rounded; external spine on exopodal segment 1 of P2 short, not longer than half of exopodal segment 2.. - Posterior surfaces of P2-P4 endopod and exopod segments usually with numerous spines and spinules; posterior surfaces of endopodal segments 2-3 and exopodal segments 2-3 of P4 sometimes with flat, large, lancet-like spines; P4 segments of protopod with numerous spines and spinules; anterior surface of endopodal segment 2 of P3 with long spines distally. - P5 female uniramous, 2-3-segmented, of variable shape; common basal segment long; surface of all or distal segments with spinules. - P5 male uniramous, strongly asymmetrical; left leg much longer than the right one, 5-segmented; 4 proximal segments subcylindrical; 4th segment sometimes with long spines along inner margin, 5th segment short; right leg of 3-5 segments, not reaching beyond the middle of 2nd proximal segments of left leg. |
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[1] Scolecitrichopsis alvinae (Grice & Hulsemann, 1970) (F) [Figs] | |
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[2] Scolecitrichopsis ctenopus (Giesbrecht, 1888) (F,M) [Figs] | |
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Scolecitrichopsis difficilis Grice & Hulsemann, 1965 (F) |
Syn.: | Xanthocalanus difficilis Grice & Hulsemann, 1965 (p.235, Descr.F, figs.F) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.221) |
Rem.: | Probable dans ce genre |
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[3] Scolecitrichopsis distinctus (Grice & Hulsemann, 1970) (M) [Figs] | |
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[4] Scolecitrichopsis elenae (Schulz, 2005) (F,M) [Figs] | |
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[5] Scolecitrichopsis elongatus (Grice & Hulsemann, 1970) (F,M) [Figs] | |
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Scolecitrichopsis polaris Brodsky, 1950 (F) |
Syn.: | Xanthocalanus polaris Brodsky, 1950 (1967) (p.229, Descr.F, figs.F) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.221) |
Rem.: | Probable dans ce genre. |
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[6] Scolecitrichopsis pseudoculata (Campaner, 1979) (F) [Figs] | |
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Scolecitrichopsis spinacantha Wilson, 1942 (F) |
Syn.: | Scolecithricelle spinacantha Wilson, 1942 a (p.208, Descr.F, figs.F) |
Ref.: | Vyshkvartzeva, 1999 (2000) (p.221) |
Rem.: | Probable dans ce genre. |
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[7] Scolecitrichopsis tenuipes (T. Scott, 1894) (F,M) [Figs] | |
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(28) Scolecocalanus Farran, 1936 | |
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Rem.: | Total: 5 spp.
Definition from Bradford & al. (1983, p.113), after Farran, 1936 and C.B. Wilson, 1950 : - Rostral spines large and tapered. - Lenticular thickeningt at base of rostrum (as in Macandrewella. - Genital segment asymmetrical in dorsal view). - P4 exopod segments 3 and 4 bear longitudinal row of spinules on anterior surface. - Female P5 present only on left side ; short basal segment bearing a long curved spine. - Male right P5 with basis swollen ; endopod well developed , extending almost as far as exopod ; left leg endopod well developed. |
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[1] Scolecocalanus galeatus Farran, 1936 (F) [Figs] | |
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[2] Scolecocalanus infrequens (Tanaka, 1969) (F) [Figs] | |
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[3] Scolecocalanus lobatus Farran, 1936 (F) [Figs] | |
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[4] Scolecocalanus spinifer C.B. Wilson, 1950 (F,M) [Figs] | |
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[5] Scolecocalanus stocki Vervoort, 1990 (F,M) [Figs] | |
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(29) Scopalatum Roe, 1975 | |
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Rem.: | type: Scopalatum gibbera Roe,1975. Total: 5 spp. + 1 undet.
Nota after Boxshall & Halsey (2004, p.190): - Mx2 female with 1, male with 2 greatly enlarged brush-like sensory setae.
Diagnosis after Vives & Shmeleva (2007, p.804) : - Cephalosome and pediger segment 1 fused, pedigers 4 and 5 fused or separte. - A2 exopod slightly longer than endopod. - Mx2 endopod showing 5 setae as powder-puff and 3 flattened vermiform setae ; in female, one of the seta is brush-shape and very large ; and 2 in male. - P1 endopod 1-segmented ; P2 endopod 2-segmentd ; P3 and P4 endopod 3-segmented - P2 with large external terminal spines ; exopod segments 2 and 3 and endopod segment 2 bear spines on the surface. - Coxa of P3 with group of thick and large spines ; endopod segments 2 and 3 bear large spine on the surface ; but are small and rare on the same segments of P4. Exopodal segments lacking of spine. |
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[1] Scopalatum dubia T. Scott, 1894 (M) [Figs] | |
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[2] Scopalatum farrani Roe, 1975 (F) [Figs] | |
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[3] Scopalatum gibbera Roe, 1975 (F) [Figs] | |
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[4] Scopalatum smithae Grice, 1962 (F) [Figs] | |
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[5] Scopalatum vorax (Esterly, 1911) (F) [Figs] | |
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[6] Scopalatum sp. Bradford, Haakonssen, Jillett, 1983 (F) [Figs] | |
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(30) Scottocalanus Sars, 1905 | |
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Rem.: | Type: Scolecithrix securifrons T. Scott,1894. 15 spp. (with 1 doubtful) + 1 undet.
Sano & al. (2013, p.23) note that the genus is considerd omnivore/detritivore in previous studies, and their chemo-sensory setae are considered to be involved in the detection of detritus (see in Nishida & Ohtsuka, 1997).
Diagnosis after Bradford & al. (1983, p.115) and Vives & Shmeleva (2007, p.807) : - Head usually with crest ; Pedigegous segments 4 and 5 partly or completely fused. - Rostrum large, bifurcate with notch more or less marked, with or without short processes. - Corners of last thoracic segment pôinted sometimes slightly rounded. - Mx1 inner lobe 1 (arthrite) with 3 setae on posterior surface ; inner lobe 3 with 3 setae ; endopod segments 2 and 3 with 4 setae ; - P1 exopod segment 1 with external spine. - P5 female imperfectly 3-segmented ; terminal segment wide, with long sub-apical spine directed backwards on the inner edge, and very small apical spine. - Male P5 asymmetrical, biramous on both sides ; right leg endopod usually well developed, reaching exopod segment 2 ; left leg endopod small, terminal part of exopod complex, prehensile, formed from distal part of segment 2 and small segment 3. |
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Scottocalanus acutus Sars, 1905 (F) |
Ref.: | Sars, 1905 c (p.7) |
Rem.: | Cf. Scottocalanus securifrons |
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Scottocalanus australis Farran, 1936 (F) |
Syn.: | Scottcalanus australis : Tanaka, 1961 a (p.146, figs.F) |
Ref.: | Farran, 1936 a (p.101, Descr.F, figs.F); Owre & Foyo, 1967 (p.63, figs.F); Deevey & Brooks, 1977 (tab.2); Park, 1983 (p.198, 199, Rem.); He & al., 1992 (p.250); Vives, 1982 (p.292); Suarez & Gasca, 1991 (tab.2); Shih & Young, 1995 (p.73); Suarez-Morales & Gasca, 1998 a (p.111) |
Loc: | Australie (Grande Barrière), mers de Chine S, E, Japon (Izu), G. du Mexique, Floride, off Madère, Baie Ibéro-marocaine |
Lg.: | (34) F: 3,96; (108) F: 3,9 |
Rem.: | Pour Tanaka (1961 a), Owre & Foyo (1967) et Shih & Young ( 1995) cette espèce n’est pas synonyme de S. helenae contrairement à Vervoort, 1965 (p.45, 54), Bradford & al., 1983 (p.115) et Mulyadi, 1994 (p.385). Vives (1982) signale ces deux espèces respectivement à Madère-Baie Ibéro-marocaine et aux Canaries, de même que Suarez (1992) et Suarez-Morales & Gasca, (1998 a) dans le Golfe du Mexique, Shih & Young (1995) dans les mers de Chine. Cf. Scottocalanus helenae |
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[1] Scottocalanus backusi Grice, 1969 (F) [Figs] | |
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[2] Scottocalanus corystes Owre & Foyo, 1967 (F,M) [Figs] | |
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Scottocalanus cuneifrons Willey, 1919 (F,M) |
Ref.: | Willey, 1918 (1919) (p.194, figs.F,M) |
Rem.: | Cf. Scottocalanus securifrons |
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[3] Scottocalanus dauglishi Sewell, 1929 (F,M) [Figs] | |
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[4] Scottocalanus farrani A. Scott, 1909 (F,M) [Figs] | |
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[5] Scottocalanus helenae (Lubbock, 1856) (F,M) [Figs] | |
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Scottocalanus infrequens Tanaka, 1969 (F) |
Ref.: | Tanaka, 1969 (p.271, Descr.F, figs.F); Park, 1983 (p.198, 199); Bradford & al., 1983 (p.115); Vyshkvartzeva, 2001 (p.80: Rem.) |
Loc: | Philippines, off I. Honshu, off Californie, I. Reine Charlotte |
N: | 1 |
Lg.: | (105) F: 4,76 |
Rem.: | Cf. Scolecocalanus infrequens |
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[6] Scottocalanus investigatoris Sewell, 1929 (M) [Figs] | |
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[7] Scottocalanus longispinus A. Scott, 1909 (F, ?M) [Figs] | |
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[8] Scottocalanus persecans (Giesbrecht, 1895) (F,M) [Figs] | |
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[9] Scottocalanus rotundatus Tanaka, 1961 (F,M) [Figs] | |
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[10] Scottocalanus securifrons (T. Scott, 1894) (F,M) [Figs] | |
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[11] Scottocalanus sedatus Farran, 1936 (F) [Figs] | |
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[12] Scottocalanus setosus A. Scott, 1909 (F) [Figs] | |
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[13] Scottocalanus terranovae Farran, 1929 (F,M) [Figs] | |
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[14] Scottocalanus thomasi A. Scott, 1909 (F,M) [Figs] | |
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[15] Scottocalanus thori With, 1915 (F,M) [Figs] | |
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Scottocalanus sp. Brodsky, 1950 (F) |
Syn.: | Scottcalanus securifrons (F) : Tanaka,1937 (p.259, Rem.F, figs.F) |
Ref.: | Brodsky, 1950 (1967) (p.244, Rem.F, figs.F, Rem.) |
Loc: | Japon (Suruga) |
N: | 1 |
Lg.: | (22) F: 4; {F: 4,00} |
Rem.: | Vervoort (1965, p.36, 37) ne partage pas l’opinion de Brodsky. |
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[16] Scottocalanus sp. Morris, 1970 (M) [Figs] | |
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(31) Undinothrix Tanaka, 1961 | |
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Rem.: | Type species: Undinothrix spinosa. Total: 1 sp.
Definition from Tanaka (1961 a, p.153) after female of the type species: - Head and 1st pedigerous segment fused, 4th and 5th pediger segments incompletely separate. - Posterolateral corners of last thoracic segment produced into a triangular expansion. - Rostrum bifurcate, pointed at apex, to which a filament-like spines are attached. - Urosome 4-segmented. - Caudal rami furnished with ordinary 4 setae and a short appendicular seta. - A1 23-segmented. - P1 3-segmented exopod and 1-segmented endopod. - Mx1 with 5 setae on the exopod. - Mx2 as in Lophothrix Giesbrecht, with sensory appendages on the distal lobes. - Mxp 1st basal segment with a slender bud-like sensory appendage about the middle of the anterior margin; the proximal outer margin of the 2nd basal segment rather coarsely serrated, the distal 2 segments of endopod with each a well developed outer marginal seta. - P1 3-segmented exopod and 1-segmented endopod; the segment of the exopod with no outer edge spine. - P2 3-segmented exopod and 2-segmented endopod. - P3 and P4 have each 3-segmented exopod and endopod. - Posterior surface of the exopod and endopod of P2 to P4 furnished with spinules. - P5 symmetrical, cpmposed of 3 segments; the terminal segment furnished with 3 spines, of which the apical two are very strong and coarsely denticulated.
Male always unknown (C. R., 2017) |
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[1] Undinothrix spinosa Tanaka, 1961 (F) [Figs] | |
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(32*) Xantharus Andronov, 1981 | |
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Rem.: | Type: Xantharus formosus Andronov,1981. Total; 4 spp. For Markhaseva, Laakmann & Renz (2013, p.20) this genus must be transfered in Diaixidae family.
Diagnosis from Schulz (2006, p.55, emend.) : - Rostrum produced ventrally, bifurcate, triangular-shaped, with 2 short, slender filaments. - Cephalosome and pediger 1 fused, pedigers 4 and 5 with suture line visible at least dorsally. - Posterolateral corners of prosome produced into angular or rounded lappets. - Urosome with anal somite very short (nearly ''invisible''). - Caudal rami usually with 7 setae including minute seta I at midlength, seta II at laterodistal edge and seta VII subdistally on ventral surface. - A1 female 24-segmented, symmetrical, usually relatively short, not extending to posterior border of prosome; of greater lengths in male, with ancestral segmentsXXII and XXIII comparatively small, segment XXII hardly 0.5 times length of XXIII, and both segments each having 1 exceptionally long seta in female. - Male A1 dimorph, left 24-segmented, right one 23-segmented, due to fusion of ancestral segments XXII and XXIII. - A2 with exopod slightly longer than endopod and bearing 9 setae; endopodal segment 1 distinctly longer than terminal exopodal compound segment. - Md basis with 3-4 setae. - Mx1 with 2-3 posterior setae on praecoxal arthrite; coxal and 1st basal endite with 2 and 4 setae. - Mx2 with 4-5 setae on 1st praecoxal endite; endopod with 8 sensory setae including 5 brush-like and 3 vermiform; 1 sclerotized seta may be present. - Mxp syncoxa with setal formula 1, 2, 3, 3; 1 seta of lobe 3 sensory with very small brush-like tip. - Male mouthparts not reducede. - Sementation of legs as typical of superfamily. - P1 with subterminal outer lobehaving angular shape distally and long endopod; exopodal segments 1-3 each with 1 large outer spine. - P2-P4 with fine spinulation on posterior surfaces of endopods. - P4 with spinulation on posterior surfaces of both rami. - P2-P4 each with long, slender terminal spine bearing coarsely to densely serrate outer margin. - Female P5 symmetrical, uniramous, typically 3-segmented but basis and exopod may be fused, exopod armed with 2 apical spine-like processes and 0-1 subapical inner spine. - Male P5 biramous, slightly asymmetrical, 5-segmented, with rudimentary, unarmed endopods or leg uniramous; both legs with protopods about subequal in size, coxa with posterior surface spinulation distally; exopopds 3-segmented, 1st exopodal segmens each with 1 small outer distal spine; 3rd exopodal segment of right leg bifurcated in proximal third, with each branch tapering to unequal lengths. |
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[1] Xantharus cryeri Bradford-Grieve, 2005 (F) [Figs] | |
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[2] Xantharus formosus Andronov, 1981 (F,M) [Figs] | |
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[3] Xantharus renatehaassae Schulz, 1998 (F) [Figs] | |
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[4] Xantharus siedleckii Schulz & Kwasniewski, 2004 (F) [Figs] | |
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