List species and varieties by family
Augaptilidae Sars, 1905 ( Arietelloidea )
(0) Alrhabdus Grice, 1973 (? Augaptilidae )
Rem.: Grice (1973, p.946) placed the genus among the Heterorhabdidae, until a male is found. Boxshall & Halsey (2004, p.68) placed the genus in Augaptilidae, as incertae sedis.
Type: Alrhabdus johrdeae Grice, 1973. Total: 1 sp.:

Diagnosis after Grice (1973, p.943):
1 - Prosome about twice the length of urosome.
2 - Head separate from 1st pedigerous somite, 4th and 5th pedigerous somites fused, posterior corner bifurcate.
3 - Urosome 4-segmented.
4 - Rostral plate provided with 2 strong spines.
5 - Labrum with cluster of ventrally projecting spines.
6 - Caudal rami asymmetrical, right ramus slightly longer than left and bears elongate seta.
7 - Seta on outer distolateral corner of basis of P3-P5.
8 - Modified seta on 1st and 2nd endopods of P5.
Gender feminine.

Diagnosis after Bradford-Grieve (1999 b, p.71) :
- Relatively large copepod. Male unknown
- Head and pedigerous segment 1 separate, pedigerous segments 4 and 5 separate, posterior corners bifurcate.
- Urosome 4-segmented.
- Rostral plate with 2 strong spines.
- Labrum with a cluster of ventrally projecting spines.
- Caudal rami asymmetrical, right ramus slightly longer than left and bearing an elongate seta.
- Md with a small palp, blade with 4 large , approximately evenly spaced teeth.
- Mx1 with a well-developed inner lobe 1, remainder of limb relatively small; outer lobe 1 with 5 setae, exopod with 6 setae, basipod 2 with 1 seta, and endopod with 3 setae.
- Mx2 with all lobes present, with setae and spines becoming thicker and stronger progressing from proximal to distal, including the terminal-most spine-like setae.
- Mxp basipodwith 8 spines and setae, basipod 2 with 3 marginal spines.
- P3-P5 basipod 2 with 1 seta on outer distolateral corner.
- P5 exopod segment 2 with a short spine, endopod segments 1 and 2 with inner edge setae modified into a lobate form witha sensory filament terminally.

[1] Alrhabdus johrdeae  Grice, 1973   (F)    [Figs]
(1) Augaptilina Sars, 1920
Rem.: Type: Augaptillina scopifera Sars, 1920. Total: 1 sp.

Diagnosis after Bradford-Grieve (1999 b, p.42): As for the family definition with the following additional characters:
1 - Urosome of female 3-segmented.
2 - Mandibular palp and Mx1 very reduced.
3 - Distal setae on Mx2 and Mxp form dense, elongate bunches resembling a brush.
4 - P1 endopod 2-segmented.

[1] Augaptilina scopifera  Sars, 1920   (F)    [Figs]
(2) Augaptilus Giesbrecht, 1889
Rem.: type : Hemicalanus longicaudatus Claus,1863. Total: 7 spp. (of which 1 doubtful).

Diagnosis after Bradford-Grieve (1999b, p.42): As for the family definition with the following additional characteristics:
- Cephalosome and pediger 1 separated, pedigers 4 and 5 fused.
- Posterolateral corners of last thoracic segment usually rounded.
- Rostral filaments fine.
- Female urosome 3-segmented, genital segment nearly always a little asymmetrical.
- Female A1 25-segmented
- Urosome male 5-segmented.
- Left A1 prehensile.
- A2 exopod rarely extending beyond endopod and with a reduced number of segments.
- Mandibular palp uniramous.
- Mx1 very reduced, consisting of a 3-segmented rod.
- Mx2 with proximal lobes rudimentary and with distal setae bearing characteristic shield-shaped appendages.- Mxp often with shield-shaped appendages.
- External spines on swimming legs more or less atrophied.
- Female P5 biramous with endopod and exopod 3-segmented each.

[1] Augaptilus anceps  Farran, 1908   (F)    [Figs]

[2] Augaptilus cornutus  Wolfenden, 1911   (F)    [Figs]

[3] Augaptilus glacialis  Sars, 1900   (F,M)    [Figs]

[4] Augaptilus lamellifer  Esterly, 1911   (F)    [Figs]

[5] Augaptilus longicaudatus  (Claus, 1863)   (F,M)    [Figs]

Augaptilus lucidus    Esterly, 1911   (M)
Rem.: Cf. Centraugaptilus lucidus.

Augaptilus macrodus    Esterly, 1911   (F)
Rem.: Cf: Centraugaptilus macrodus.

[6] Augaptilus megalurus  Giesbrecht, 1889   (F,M)    [Figs]

Augaptilus polaris    Harding, 1966   (F)
Ref.: Harding, 1966 (p.17, figs.F)
Loc: Arct. (80°N; 136°W)
Lg.: F: 2,65
Rem.: Cf. ?

Augaptilus pyramidalis    Esterly, 1911   (F)
Rem.: Cf. Centraugaptilus horridus.

[7] Augaptilus spinifrons  Sars, 1907   (F,M)    [Figs]
Remark : The following species were originally included in the genus Augaptilus :
A. angustus Sars,1905 ( Cf. Euaugaptilus angustus ); A. antarcticus Wolfenden, 1911 (Cf. Euaugaptilus laticeps ); A. brevicaudatus Sars,1905 (Cf. Euaugaptilus squamatus ); A. bullifer Giesbrecht,1889 (Cf. Euaugaptilus bullifer ); A. californicus Esterly,1913 (Cf. Euaugaptilus squamatus ); A. clavatus Sars,1907 (Cf. Euaugaptilus clavatus ); A. cucullatus Sars,1905 (Cf. Centraugaptilus cucullatus ); A. depressus Esterly,1913 (Cf. Euaugaptilus filigerus ); A. elongatus Sars,1905 (Cf. Euaugaptilus elongatus ); A. facilis Farran,1908 (Cf. Euaugaptilus facilis ); Augaptilus fungiferus Steuer,1904 (F) [Ref.: Steuer,1904 (p.597,Descr.F); Wolfenden,1911 (p.336,figs.F); Matthews,1972 (p.35, Rem .)](Cf. Euaugaptilus magnus ); A. gibbus Wolfenden, 1904 (Cf. Euaugaptilus gibbus (Wolfenden, 1904); Augaptilus gibbus Sars,1905 [Ref.: Sars,1905 c (p.16,Rem.F);1907 a (p.3); Matthews,1972 (p.31)](Cf. Euaugaptilus gibbus (Wolfenden,1904)); A. gracilis Sars,1905 ( Cf. Euaugaptilus gracilis ); A. hecticus Giesbrecht,1889 (Cf. Euaugaptilus hecticus ); A. horridus Farran,1908 (F) (Cf. Centraugaptilus horridus ); A. laticeps Sars,1905 (Cf. Euaugaptilus laticeps ); A. latifrons Sars, 1907 (Cf. Euaugaptilus latifrons ); A. longicirrhus Sars,1905 (Cf. Euaugaptilus longicirrhus ); A. longimanus Sars,1905 (Cf. Euaugaptilus longimanus ); A. lucidus Esterly, 1911 (Cf. Centraugaptilus lucidus ); A. macrodus Esterly, 1911(Cf. Centraugaptilus rattrayi ); A. magnus Wolfenden,1904 (Cf. Euaugaptilus magnus ); A. mixtus Sars,1907 (Cf. Euaugaptilus mixtus ); A. nodifrons Sars,1905 (Cf. Euaugaptilus nodifrons ); A. oblongus Sars,1905 (Cf. Euaugaptilus oblongus ); A. palumbii Giesbrecht,1889 (Cf. Euaugaptilus palumboi ); A. placitus A. Scott,1909 (Cf. Euaugaptilus laticeps ); A. pyramidalis Esterly,1911 (Cf. Centraugaptilus horridus ); A. rattrayi T. Scott,1894 (Cf. Centraugaptilus rattrayi ); A. romanus Esterly, 1913 (Cf. Euaugaptilus filigerus ); A. rostratus Esterly,1906 (Cf. Euaugaptilus oblongus ); A. similis Farran,1908 (Cf. Euaugaptilus similis ); A. simplex Wolfenden,1911 (Cf. Euaugaptilus nodifrons ); Augaptilus simplex Esterly,1913 (Cf. Euaugaptilus nodifrons ); A. squamatus Giesbrecht,1889 (Cf. Euaugaptilus squamatus ); A. subfiligerus Wolfenden,1911 (Cf. Euaugaptilus oblongus ); A. tenuicaudis Sars,1905 (Cf. Euaugaptilus tenuicaudis ); A. tenuispinus Sars,1920 (Cf. Euaugaptilus tenuispinus ); A. truncatus Sars,1905 (Cf. Euaugaptilus truncatus ); A. validus A. Scott,1909 (Cf. Euaugaptilus validus ); A. zetesios Wolfenden,1902 (Cf. Augaptilus glacialis )
(3) Centraugaptilus Sars, 1920
Rem.: type: Augaptilus rattrayi rattrayi T. Scott, 1894 . Total: 6 spp.

Diagnosis after Bradford-Grieve (1999 b, p.42):
- As for the family definition with additional chatacteristics.
- The genus is separated from Augaptilus by having a strong bifurcate rostrum projecting forward.
- Cephalosome and pediger 1 separated, pedigers 4 and 5 fused.
- A1 short, not extending beyond prosome.
- Md elongate with 2 fine teeth and a smaller tooth between them.
- Mx1 inner lobe 1 with 2 long, hooked spines bearing shield-like appendages and 1 short seta; inner lobes 2 and 3 absent; endopod with 3 setae; exopod with 3 setae; outer lobe with 5 setae.
- Mx2 and Mxp powerful with distal setae strong and rolled up, carrying shield-shaped appendages.
- P1 exopod segment 2 without an outer distal spine.

[1] Centraugaptilus cucullatus  (Sars, 1905)   (F,M)    [Figs]

[2] Centraugaptilus horridus  (Farran, 1908)   (F,M)    [Figs]

[3] Centraugaptilus lucidus  (Esterly, 1911)   (M)    [Figs]

[4] Centraugaptilus macrodus  (Esterly, 1911)   (F)    [Figs]

[5] Centraugaptilus porcellus  Johnson, 1936   (F,M)    [Figs]

[6] Centraugaptilus rattrayi  (T. Scott, 1894)   (F,M)    [Figs]
(4) Euaugaptilus Sars, 1920
Rem.: Bradford-Grieve (1999 b) follows the position of Matthews (1972, p.63) concerning the synonymy of Neogaptilus. This genus is essentially bathypelagic.
Type: Augaptilus squamatus Giesbrecht, 1899. Total: 72 spp. + 1 unidentified.

Twenty-five characters used in the calculation of similarity coefficients have been tested by Matthews (1972) to discover which could be used in the diagnoses of the two groups. Since the sequence of species is associated with a tendency towards simplification of the structure and setation of the head appendages, a tendency which appears to have been followed along several different pathways, no single character was sufficient to define the groups. Several species, moreover, show affinities with both groups, so any definition must to some extent be arbitrary in its placing of the division between the groups. For this reason the groups have not been given generic or subgeneric rank.It is possible to classify the species based on the structure of Mx1 in 2 groups: ''affinis''' and ''qsquamatus Groups:
1 - "affinis'' Group:
Mx1 reduced with rarely an endopodite and 3rd endite (lobe) generally bears no setae ; in no case are an endopodite and a setose 3rd endite present together. The toyal number of setae and spines never exceeds 22, an dis usually considerably less.
The cup-shaped appendages on certain setae of Mx2 and Mxp almost always well developed.

2 - ''squamatus'' Group:
Mx1 endopodite often present. Generally the appendage with a total setae and spines 24 and 40.
The cup-shaped appendages on certain setae on Mx2 and Mxp often poorly developed or even absent.
E. angustus, E. grandicornis, E. rectus lack both endopodite in Mx1 and any setae on the 3rd endite, but in all three there are at least 26 setae and spines on the whole appendage.
E. marginatus, E. oblongus have only 21 and 23 setae and spines, respectively, on Mx1.

Diagnosis after Bradford-Grieve (1999 b, p.46): As for the family definition with the following additional characteristics:
- Cephalosome and pediger 1 separated , pedigers 4 and 5 fused.
- Posterolateral corners of last thoracic segment rounded
- Rostral filaments may be present or absent.
- Distinguished from Augaptilus by the greater development of Md and speciallay the Mx1 inner lobe 1 well-developed and provided with a number of strong spines terminally.
- Mandibular palp may be uni or biramous.
- Mx1 inner lobes 2 and 3 and basipod 2 with or without setae, endopod present or absent.
- Mxp may be elongated or more squat.
- P1 exopod and endopods may be 3-segmented or may have some degree of fusion between segments; basipod 2 seta present or absent; external edge spines on exopod variable.
- Female P5 either with both rami 3-segtmented or with some degree of fusion between segments.

After Bode & al. (2018, p.75) some species of this genus are ambush predators, which prey on small copepods by using specialized ''button setae'' on their maxilla and maxilliped (Matsuura & Nishida, 2000; Matsuura & al., 2010).

[1] Euaugaptilus affinis  Sars, 1920   (F)    [Figs]

[2] Euaugaptilus aliquantus  Park, 1993   (F)    [Figs]

[3] Euaugaptilus angustus  (Sars, 1905)   (F,M)    [Figs]

[4] Euaugaptilus antarcticus  (Wolfenden, 1911)   (F,M)    [Figs]

[5] Euaugaptilus atlanticus  Roe, 1975   (F,M)    [Figs]

[6] Euaugaptilus austrinus  Park, 1993   (F,M)    [Figs]

Euaugaptilus brevicaudatus    Sars, 1905   (F)
Syn.: Augaptilus brevicaudatus Sars, 1905 (p.12, Descr.F)
Ref.: Matthews, 1972 (p.28, 41)
Rem.: Cf. Euaugaptilus squamatus

[7] Euaugaptilus brevirostratus  Park, 1993   (F,M)    [Figs]

[8] Euaugaptilus brodskyi  Hulsemann, 1967   (F,M)    [Figs]

[9] Euaugaptilus bullifer  (Giesbrecht, 1889)   (F,M)    [Figs]

Euaugaptilus californicus    Esterly, 1913   (F)
Syn.: Augaptilus californicus Esterly, 1913 (p.186, Descr.F, figs.F); Brodsky, 1950 (1967) (p.371, figs.F,M)
Ref.: Matthews, 1972 (p.28, 41, Rem.)
Rem.: Cf. Euaugaptilus squamatus

[10] Euaugaptilus clavatus  (Sars, 1907)   (F)    [Figs]

[11] Euaugaptilus curtus  Grice & Hülsemann, 1967   (F,M)    [Figs]

Euaugaptilus depressus    Esterly, 1913   (F)
Syn.: Augaptilus depressus Esterly, 1913 (p.187, Descr.F, figs.F)
Ref.: Matthews, 1972 (p.29, 30)
Rem.: Cf. Euaugaptilus filigerus

[12] Euaugaptilus digitatus  Sars, 1920   (F,M)    [Figs]

[13] Euaugaptilus diminutus  Park, 1970   (F)    [Figs]

[14] Euaugaptilus distinctus  (Brodsky, 1950)   (F)    [Figs]

[15] Euaugaptilus elongatus  (Sars, 1905)   (F,M)    [Figs]

[16] Euaugaptilus facilis  (Farran, 1908)   (F,M)    [Figs]

[17] Euaugaptilus fagettiae  Björnberg, 1975   (F)    [Figs]

[18] Euaugaptilus farrani  Sars, 1920   (F)    [Figs]

[19] Euaugaptilus fecundus  Tanaka & Omori, 1974   (F,M)    [Figs]

Euaugaptilus filiger    Claus, 1863   
Syn.: Hemicalanus filigerus Claus,1863 (p.179)
Rem.: L'appellation filiger paraît grammaticalement la plus correcte, mais le Code de Nomenclature Zoologique impose de conserver l'appellation d'origine.
Cf. Euaugaptilus filigerus

[20] Euaugaptilus filigerus  (Claus, 1863)   (F,M)    [Figs]

[21] Euaugaptilus fosaii  Pineda-Polo, 1979   (F)    [Figs]

[22] Euaugaptilus fundatus  Grice & Hülsemann, 1967   (F)    [Figs]

Euaugaptilus fungiferus    Steuer, 1904   (F)
Syn.: Augaptilus fungiferus Steuer, 1904 (p.597, Descr.F); non Wolfenden, 1911 (p.336)
Ref.: Vervoort, 1957 (p.139, Rem.); Matthews, 1972 (p.31)
Rem.: Cf. Euaugaptilus magnus

[23] Euaugaptilus gibbus  (Wolfenden, 1904)   (F,M)    [Figs]

Euaugaptilus gibbus    Sars, 1905   (F,M)
Syn.: Augaptilus gibbus Sars, 1905 c (p.16, Descr.F)
Ref.: Matthews, 1972 (p.31)
Rem.: Cf. Euaugaptilus gibbus (Wolfenden,1904)

[24] Euaugaptilus gracilis  (Sars, 1905)   (F)    [Figs]

[25] Euaugaptilus graciloides  Brodsky, 1950   (F)    [Figs]

[26] Euaugaptilus grandicornis  Sars, 1920   (F)    [Figs]

[27] Euaugaptilus hadrocephalus  Park, 1993   (F)    [Figs]

[28] Euaugaptilus hecticus  (Giesbrecht, 1889)   (F,M)    [Figs]

[29] Euaugaptilus hulsemannae  Matthews, 1972   (F)    [Figs]

[30] Euaugaptilus humilis  Farran, 1926   (F)    [Figs]

[31] Euaugaptilus hyperboreus  Brodsky, 1950   (F,M)    [Figs]

[32] Euaugaptilus indicus  Sewell, 1932   (F)    [Figs]

[33] Euaugaptilus laticeps  (Sars, 1905)   (F,M)    [Figs]

[34] Euaugaptilus latifrons  (Sars, 1907)   (F)    [Figs]

Euaugaptilus longiantennalis    Park, 1970   (F)
Ref.: Park, 1970 (p.533, figs.F); Matthews, 1972 (p.34, 36)
Rem.: Cf.Euaugaptilus marginatus

[35] Euaugaptilus longicirrhus  (Sars, 1905)   (F,M)    [Figs]

[36] Euaugaptilus longimanus  (Sars, 1905)   (F,M)    [Figs]

[37] Euaugaptilus longiseta  Grice & Hülsemann, 1965   (F,M)    [Figs]

[38] Euaugaptilus luxus  Tanaka & Omori, 1974   (F)    [Figs]

[39] Euaugaptilus magnus  (Wolfenden, 1904)   (F,M)    [Figs]

Euaugaptilus magnus magnus    Vervoort, 1965   
Syn.: Euaugaptilus magnus : Sewell, 1947 (p.213)
Ref.: Vervoort, 1965 (p.139, Rem.); Matthews, 1972 (p.35)
Loc: G. de Guinée, Japon
Lg.: (16) F: 7,82-6,6; M: 7,9
Rem.: Cf. Euaugaptilus magnus

Euaugaptilus magnus fungiferus    Sewell, 1947   
Ref.: Sewell, 1947 (p.214, figs.F, Rem.juv.); Vervoort, 1957 (p.139, Rem.); Matthews, 1972 (p.35, Rem.)
Loc: Mer Arabe
Lg.: (11) F: 7,408-7,083
Rem.: Cf.Euaugaptilus magnus

[40] Euaugaptilus malacus  Grice & Hülsemann, 1967   (F)    [Figs]

[41] Euaugaptilus marginatus  Tanaka, 1964   (F)    [Figs]

[42] Euaugaptilus matsuei  Tanaka & Omori, 1967   (F)    [Figs]

[43] Euaugaptilus maxillaris  Sars, 1920   (F,M)    [Figs]

[44] Euaugaptilus mixtus  (Sars, 1907)   (F)    [Figs]

Euaugaptilus mixtus    Brodsky, 1950   (F,M)
Ref.: Brodsky, 1950 (1967) (p.380, figs.F,M); Tanaka, 1974 b (p.58, figs.F); Matthews, 1972 (p.37)
Rem.: non E. mixtus (Sars,1907), nom praeoc.: Cf. Euaugaptilus brodskyi

[45] Euaugaptilus modestus  Brodsky, 1950   (M)    [Figs]

[46] Euaugaptilus niveus  Tanaka & Omori, 1974   (F, ?M)    [Figs]

[47] Euaugaptilus nodifrons  (Sars, 1905)   (F,M)    [Figs]

[48] Euaugaptilus nudus  Tanaka, 1964   (F)    [Figs]

[49] Euaugaptilus oblongus  (Sars, 1905)   (F,M)    [Figs]

[50] Euaugaptilus pachychaeta  Matthews, 1972   (F)    [Figs]

[51] Euaugaptilus pacificus  Matthews, 1972   (F,M)    [Figs]

[52] Euaugaptilus palumbii  (Giesbrecht, 1889)   (F,M)    [Figs]

[53] Euaugaptilus parabullifer  Brodsky, 1950   (F)    [Figs]

[54] Euaugaptilus paroblongus  Matthews, 1972   (F)    [Figs]

[55] Euaugaptilus penicillatus  Sars, 1920   (F,M)    [Figs]

[56] Euaugaptilus perasetosus  Park, 1993   (F)    [Figs]

Euaugaptilus perodiosus    Tanaka & Omori, 1974   (F)
Ref.: Tanaka & Omori, 1974 (p.246, Descr.F, figs.F); Boxshall, 1985 (p.306, Rem.)
Loc: Japon (Izu)
Rem.: Cf. Eaugaptilus placitus

[57] Euaugaptilus placitus  (A. Scott, 1909)   (F,M)    [Figs]

Euaugaptilus propinquus    Sars, 1920   (F)
Ref.: compl: Deevey & Brooks, 1977 (tab.2); Vives, 1982 (p.294
Loc: Açores-Canaries, off Madère, off Portugal, off Maroc-Mauritanie, Mer des Sargasses, Japon
Rem.: Hulsemann (1967) et avec doute Tanaka & Omori (1974) d’une part et Roe (1975) d’autre part ne s'accordent pas sur la synoymie entre cette forme et E. mixtus. Cf. Euaugaptilus mixtus (Sars,1907)

Euaugaptilus pseudaffinis    Brodsky, 1950   (F)
Ref.: Brodsky, 1950 (1967) (p.380, Descr.F, figs.F); Grice & Hülsemann, 1967 (p.36, Rem.); Matthews, 1972 (p.40)
Loc: Pacif. NW, Station Knot
Lg.: (22) F: 5,1
Rem.: ? Cf. Euaugaptilus affinis

[58] Euaugaptilus quaesitus  Grice & Hülsemann, 1967   (F)    [Figs]

[59] Euaugaptilus rectus  Grice & Hülsemann, 1967   (F,M)    [Figs]

[60] Euaugaptilus rigidus  (Sars, 1907)   (F,M)    [Figs]

[61] Euaugaptilus roei  Matthews, 1972   (F)    [Figs]

[62] Euaugaptilus sarsi  Grice & Hülsemann, 1965   (F)    [Figs]

[63] Euaugaptilus similis  (Farran, 1908)   (F)    [Figs]

Euaugaptilus similis    Brodsky, 1950   (F,M)
Ref.: Brodsky, 1950 (1967) (p.378, figs.F,M); Matthews, 1972 (p.22, 41, Rem.)
Rem.: non Euaugaptilus similis (Farran,1908), nom. praec.: Cf. Euaugaptilus pacificus

Euaugaptilus simulans    Sars, 1924   (F)
Ref.: Sars, 1925 (p.284, figs.F); Matthews, 1972 (p.41, 44, Rem.)
Rem.: Cf. Euaugaptilus vicinus

[64] Euaugaptilus squamatus  (Giesbrecht, 1889)   (F,M)    [Figs]

Euaugaptilus subfiligerus    Wolfenden, 1911   (F)
Syn.: Augaptilus subfiligerus Wolfenden, 1911 (p.343)
Ref.: Matthews, 1972 (p.38, 42)
Rem.: Cf. Euaugaptilus oblongus

[65] Euaugaptilus sublongiseta  Park, 1970   (F)    [Figs]

[66] Euaugaptilus tenuicaudis  (Sars, 1905)   (F)    [Figs]

[67] Euaugaptilus tenuispinus  Sars, 1920   (F)    [Figs]

[68] Euaugaptilus truncatus  (Sars, 1905)   (F)    [Figs]

[69] Euaugaptilus unisetosus  Park, 1970   (F)    [Figs]

[70] Euaugaptilus validus  (A. Scott, 1909)   (F,M)    [Figs]

[71] Euaugaptilus vescus  Park, 1970   (F)    [Figs]

[72] Euaugaptilus vicinus  Sars, 1920   (F)    [Figs]

Euaugaptilus sp.    Grice & Hulsemann, 1965   (M)
Ref.: Grice & Hülsemann, 1965 (p.249, Descr.M, figs.M); Matthews, 1972 (p.28, 44); Tanaka & Omori, 1974 (p.239, 241, Rem.)
Loc: Atlant. NE (30-60°)
Lg.: (226) M: 3,07
Rem.: Cf. Euaugaptilus niveus

Euaugaptilus sp.1    Grice & Hulsemann, 1967   (F)
Ref.: Grice & Hulsemann, 1967 (p.19, 35, Descr.F, figs.F); Matthews, 1972 (p.30, 44)
Rem.: Cf. Euaugaptilus facilis

[73] Euaugaptilus sp.2  Grice & Hülsemann, 1967   (M)    [Figs]

Euaugaptilus sp.3    Grice & Hulsemann, 1967   (M)
Ref.: Grice & Hulsemann, 1967 (p.19, 37, Descr.M, figs.M) ; Matthews, 1972 (p.45); Roe, 1975 (p.351, Rem.)
Loc: Indien SW
Lg.: (67) M: 6,16; {M: 6,16}
Rem.: Cf. Euaugaptilus fecundus

Euaugaptilus sp.1    Tanaka & Omori, 1974   (juv. F)
Ref.: Tanaka & Omori, 1974 (p.260, juv.F 5)
Loc: Japon (Izu)
Lg.: (68) F: 3,19

Euaugaptilus sp.2    Tanaka & Omori, 1974   (juv. F?)
Ref.: Tanaka & Omori, 1974 (p.262, figs. juv 4 F?)
Loc: Japon (Izu)
Lg.: (68) 4,79.
(5) Frankferrarius Markhaseva, 2013
Rem.: Type: Frankferrarius admirabilis Markhaseva, 2013. 1 sp.
Provisionally placed in the Augaptilidae family by Markhaseva (2013, p.1263). For the author the species F. admirabilis shares with this family:
1- the general pattern of segmentation and setation of its swimming legs although P3 endopod segment 3 armament is 2, 2 and 3 setae (versus 2, 2 and 4 setae in augaptilids);
2 - a single genital operculum positioned medially on the ventral part of the genital double-somite;
3 - the female P5 has a 3-segmented exopod with armament of I-O, I-1, II, I, 3;
4- the male left P5 is of the augaptilid type. Nevertheless, the species type differs from other augaptilids in being without setae on Mxp basis and in details of morphology and setation of oral parts and basis of P1. This new genus probably should be placed in a new family but this must await a revision of the Augaptilidae.

[1] Frankferrarius admirabilis  Markhaseva, 2013   (F,M)    [Figs]
(6) Haloptilus Giesbrecht, 1898
Rem.: sp. type: Hemicalanus plumosus Claus, 1863. Total: 26 spp. (of which 2 doubtful) + 2 unidentified.
Park (1988, p.2) divides the genus into 3 groups based on the numbers of teeth on the mandibular cutting edge, the number of endopodal segments on Mx1 and the number of setae of Mx1 and Mx2. 1- Primitive group represented by H. fons, H. angusticeps, H. caribbeanensis, H. furcatus, H. major. 2- Specialized group: H. acutifrons, H. longicirrus, H. longicornis, H. mucronatus, H. H. ocellatus, H. ornatus, H. oxycephalus, H. paralongicirrus, H. plumosus, H. pseudooxycephalus, H. spiniceps, H. tenuis. 3- Neither obviously primitive nor fully specialized with regard to these characters: H. austini, H. chierchiae, H. validus.

Diagnosis after Bradford-Grieve (1999 b, p.58): As in the family with the following additional characteristics:
- Female urosome 4-segmented, symmetrical.
- Male urosome 5-segmented.
- Caudal rami and female mouthparts with long plumose setae.
- Male mouthparts atrophied.
- Female A1 25-segmented.
- Male A1 shorter than female and well supplied with aesthetascs, geniculate on left and 21-segmented, with a 4-segmented distal part.
- A2 endopod much longer than exopod.
- Mandible blade forked, endopod and exopod elongate.
- Mx1 well developed with inner lobes 2 and 3, endopod small, exopod elongate.
- Mx2 elongate with reduced lobes, distal setae not much longer than proximal setae.
- Mxp endopod thick.
- P5 exopod segment 2 with inner seta pointed.
- Female P5 with last inner seta of exopodal segment 3 of usual kength.
- Male P5 exopod and endopod 3-segmented and with both terminal exopod segments hooked, without a pincer, right and left alike.

[1] Haloptilus aculeatus  (Brady, 1883)   (F)    [Figs]

[2] Haloptilus acutifrons  (Giesbrecht, 1892)   (F,M)    [Figs]

[3] Haloptilus angusticeps  Sars, 1907   (F,M)    [Figs]

[4] Haloptilus austini  Grice, 1959   (F)    [Figs]

[5] Haloptilus bulliceps  Farran, 1926   (M,F juv.)    [Figs]

[6] Haloptilus caribbeanensis  Park, 1970   (F)    [Figs]

[7] Haloptilus chierchiae  (Giesbrecht, 1889)   (F,M)    [Figs]

[8] Haloptilus fertilis  (Giesbrecht, 1892)   (M)    [Figs]

[9] Haloptilus fons  Farran, 1908   (F)    [Figs]

[10] Haloptilus furcatus  Sars, 1920   (F)    [Figs]

[11] Haloptilus longiceps  Tanaka, 1964   (F)    [Figs]

[12] Haloptilus longicirrus  Brodsky, 1950   (F)    [Figs]

[13] Haloptilus longicornis  (Claus, 1863)   (F,M)    [Figs]

Haloptilus longimanus    Wolfenden, 1906   (F)
Ref.: Pseudhaloptilus longimanu

[14] Haloptilus major  Wolfenden, 1911   (F)    [Figs]

[15] Haloptilus mucronatus  (Claus, 1863)   (F,M)    [Figs]

[16] Haloptilus ocellatus  Wolfenden, 1905   (F,M)    [Figs]

Haloptilus orientalis    Brady, 1883   (juv. F)
Syn.: Hemicalanus orientalis Brady, 1883 (p.45, figs.juv.F); Giesbrecht, 1892 (p.384)
Ref.: Giesbrecht & Schmeil, 1898 (p.120); Matthews, 1972 (p.52); Park, 1988 (p.3)
Loc: Mer de Banda
Rem.: espèce indéterminable.

[17] Haloptilus ornatus  (Giesbrecht, 1892)   (F,M)    [Figs]

[18] Haloptilus oxycephalus  (Giesbrecht, 1889)   (F,M)    [Figs]

[19] Haloptilus pacificus  Chiba, 1956   (F)    [Figs]

[20] Haloptilus paralongicirrus  Park, 1970   (F,M)    [Figs]

[21] Haloptilus plumosus  (Claus, 1863)   (F,M)    [Figs]

[22] Haloptilus pseudooxycephalus  Brodsky, 1950   (F)    [Figs]

[23] Haloptilus setuliger  Tanaka, 1964   (F)    [Figs]

[24] Haloptilus spiniceps  (Giesbrecht, 1892)   (F,M)    [Figs]

Haloptilus spinifrons    Sars, 1900   (F)
Syn.: Hemicalanus spinifrons Sars, 1900 (p.95, figs.F)
Ref.: Matthews, 1972 (p.48)
Loc: Arct.
Rem.: Cf. Haloptilus acutifrons

[25] Haloptilus tenuis  Farran, 1908   (F,M)    [Figs]

[26] Haloptilus validus  Sars, 1920   (F)    [Figs]

Haloptilus sp.    Tanaka & Omori, 1971   (juv. F)
Ref.: Tanaka & Omori, 1971 (p.258, figs.juv.F)
Loc: Japon (Izu)
Lg.: (90) juv.F: 4,33

[27] Haloptilus sp.  Arcos, 1975   (M)    [Figs]

[28] Haloptilus sp.  Ali-Khan & Ali-Khan, 1984   (F)    [Figs]
(7) Heteroptilus Sars, 1920
Rem.: sp. type: Pontoptilus attenuatus. 2 spp. + 1 unidentified.

Diagnosis after Bradford-Grieve (1999 b, p.66): As in the family with the following additional characters. The genus is distinguished by the structure of the mandibles which resemble those of Heterorhabdus with right mandibular blade much larger than the left and carrying an extremely large sctthe-like hook.
- Urosome of female 4-segmented. P1 with a 2-segmented endopod.
- Female P5 with a 1-segmented endopod.
- Male P5 with 3-segmented endopods.

[1] Heteroptilus acutilobus  (Sars, 1905)   (F,M)    [Figs]

[2] Heteroptilus attenuatus  (Sars, 1905)   (F, ? M)    [Figs]

[3] Heteroptilus sp.  Sewell, 1947   (M)    [Figs]

Heteroptilus sp.    Vervoort, 1957   (M)
Ref.: Vervoort,1957 (p.141)
Loc: Antarct. (Pacif. SW)
Rem.: espèce non identifiable.
Neoaugaptilus Brodsky, 1950
Rem.: type: Neoaugaptilus distinctus. Cf. Euaugaptilus
Neoaugaptilus distinctus    Brodsky, 1950   (F)
Ref.: Brodsky, 1950 (1967) (p.385, Descr.F, figs.F); Matthews, 1972 (p.4, 29, 63, Rem.)
Loc: Pacif. NW
Rem.: Cf. Euaugaptilus distinctus

Pachyptilus Sars, 1920
Rem.: Cf. Pseudhaloptilus (this genus is considered questionable by Bradford-Grieve et al., 1999, p. 942, but admitted by Boxshall & Halsey, 2004, p.68 who consider Pachyptilus to be a junior synonym).
Pachyptilus abbreviatus    Sars, 1905   (F)
Syn.: Pontoptilus abbreviatus Sars, 1905 c (p.19, Rem.F); Farran, 1908 b (p.81, Rem.F); Lysholm & Nordgaard, 1921 (p.27)
Ref.: Sars, 1925 (p.319, figs.F); Rose, 1929 (p.36); 1933 a (p.236, figs.F
Rem.: Cf. Pseudhaloptilus abbreviatus.
Pachyptilus eurygnathus    Sars, 1920   (F)
Syn.: non P. eurygnathus : Sewell, 1947 (p.239)
Ref.: Sars, 1920 c (p.18, Rem.F); 1925 (p.321, figs.F); Rose, 1933 a (p.236, figs.F)
Rem.: Cf. Pseudhaloptilus eurygnathus.
Pachyptilus lobatus    Sars, 1920   (F)
Ref.: Sars, 1920 c (p.19, Rem.F); 1925 (p.322, figs.F
Rem.: Cf. Pseudhaloptilus lobatus.
Pachyptilus longimanus    Wolfenden, 1906   (F)
Syn.: Haloptilus longimanus Wolfenden, 1906 (p.29, Descr.F, figs.F)
Ref.: Bradford-Grieve & al., 1999 (p.883, 942, figs.F)
Loc: Atlant. S
Lg.: (10) F: 6,72
Rem.: Cf. Pseudhaloptilus longimanus.
Pachyptilus pacificus    Johnson, 1936   (F,M)
Syn.: P. eurygnathus : Sewell, 1947 (p.239, figs.F, Rem.)
Ref.: M.W. Johnson, 1936 (p.65, Descr.F, figs.F); Brodsky, 1950 (1967) (p.392, figs.F); Tanaka, 1964 b (p.88, fig.F, Rem.M);
Rem.: Cf. Pseudhaloptilus pacificus.
(8) Pontoptilus Sars, 1905
Rem.: Type species: Pontoptilus muticus Sars, 1905. Total: 6 spp.

Diagnosis after Bradford-Grieve (1999 b, p.66): As in the family with the following additional characters:
- Urosome female 4-segmented.
- A2 exopod longer than endopod.
- Mx1 with a large inner lobe 1 and all parts present with slender, curved setae.
- P5 female with endopod 2-segmented.

Nota: Body bulky. Cephalosome and pediger 1 separated, pedigers 4 and 5 fused. Posterolateral corners of last thoracic segment symmetrical, rounded or pointed .

Pontoptilus abbreviatus    Sars, 1905   (F)
Ref.: Sars, 1905 c (p.19, Rem.F); Farran, 1908 b (p.81, Rem.)
Rem.: Cf. Pachyptilus abbreviatus

Pontoptilus acutilobus    Sars, 1905   (F)
Ref.: Sars, 1905 c (p.21, Rem.F)
Rem.: Cf. Heteroptilus acutilobus

Pontoptilus attenuatus    Sars, 1905   (F)
Ref.: Sars, 1905 c (p.20, Rem.F)
Rem.: Cf. Heteroptilus attenuatus

[1] Pontoptilus lacertosus  Grice & Hulsemann, 1967   (F)    [Figs]

[2] Pontoptilus mucronatus  Sars, 1905   (F,M)    [Figs]

[3] Pontoptilus muticus  Sars, 1905   (F)    [Figs]

[4] Pontoptilus ovalis  Sars, 1907   (F)    [Figs]

[5] Pontoptilus pertenuis  Sars, 1907   (F)    [Figs]

[6] Pontoptilus robustus  Sars, 1905   (F,M)    [Figs]
(9) Pseudaugaptilus Sars, 1907
Rem.: Type: Isochaeta longisetosus. Total: 3 spp. (of which 1 doubtful).

For Bradford-Grieve (1999 b, p.68) this genus differs from the genus Euaugaptilus by a 4-segmented urosome in female.
- Rostral filaments long and thin.
- Mandibular blade long and thin and denticulate distally.
- P5 in both sexes with exopod without outer edge spines; and differing little between the sexes.

[1] Pseudaugaptilus longiremis  Sars, 1907   (F,M)    [Figs]

[2] Pseudaugaptilus orientalis  Tanaka, 1964   (F)    [Figs]

[3] Pseudaugaptilus polaris  Brodsky, 1950   (F)    [Figs]

Pseudaugaptilus sp.    Minoda, 1971   (M)
Ref.: Minoda, 1971 (p.41, Rem.M)
Loc: Mer de Béring
Lg.: (208) M: 5,1.
(10) Pseudhaloptilus Wolfenden, 1911
Rem.: Type: Pontoptilus abbreviatus Sars, 1905.Total: 5 spp.

Diagnosis after Bradford-Grieve (1999 b, p.66): As in the family with the following additional characters:
- Genus distinguished from Pontoptilus by the markedly compact body.
- Presence of a well-developed, bifurcate rostrum.
- Extraordinary development of the anterior lip.
- Md blade elarged and axe-shaped.
- P5 female with 1-segmented endopods.

[1] Pseudhaloptilus abbreviatus  (Sars, 1905)   (F)    [Figs]

[2] Pseudhaloptilus eurygnathus  (Sars, 1920)   (F)    [Figs]

[3] Pseudhaloptilus lobatus  (Sars, 1920)   (F)    [Figs]

[4] Pseudhaloptilus longimanus  (Wolfenden, 1906)   (F)    [Figs]

[5] Pseudhaloptilus pacificus  (Johnson, 1936)   (F,M)    [Figs]

 Any use of this site for a publication will be mentioned with the following reference :

Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2024. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at [Accessed February 27, 2024]

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