List species and varieties by family
Tortanidae Sars, 1902 ( Diaptomoidea )
(1) Tortanus Giesbrecht,1898; emend. Ohtsuka & Reid, 1998
Rem.: Type: Corynura gracilis Brady, 1883. 5 S.G.: Acutanus , Atortus, Boreotortanus, Eutortanus , Tortanus . Total: 47 spp. + 1 undet.

Diagnosis from Bradford-Grieve (1999 b, p.228):
- 1 median eye.
- No rostrum; a semicircular plate, thickly covered with short bristles, lies anterior to the labrum.
- Head and pediger segment 1 separate, pediger segments 4 and 5 fused or separate.
- Posterior corners of pediger segment 5 with rounded or pointed extremities which are small or absent in the male.
- Urosomal segments 2- or 3-segmented in female
- Genital segment female without seminal receptacles.
- Urosome 5-segmented in male, often asymmetrical, both because of asymmetry in individual segments and as a result of curvature along the longitudinal axis; urosome asymmetry less frequent in males.
- Female genital segment often asymmetrical, bearing a common genital aperture located medially on ventral surface, copulatory pore contained within median gemital aperture.
- Caudal rami sometimes asymmetrical, one larger than the other, often fused to the anal segmebt, with 6 setae.
- A1 12- to 15-segmented, prehensile on the right side in the male, its middle part widened and sometimes equipped with a denticulate plate.
- A2 coxa and basis separate, endopod and basis often fused; with exopod indistinctly 3-segmented, typically with an unarmed 1st segment, a long middle segment with up to 3 setae, and a short segment with 2 setae; endopod 2-segmented with 6 terminal setae.
- Md with an elongate palp; unarmed basis; endopod 2-segmented with 0 and 6 setae; exopod indistinctly 1- 4-segmented with 5 setae in total.
- Mx1 with much reduced number of lobes (inner lobe 1 one other the only ones present); inner lobe 1 with up to 13 spines and setae, the distal segment elongate bearing 3 powerful claw-like setae around the apex, and up to 7 short setae subapically.
- Mx2 with lobes 1-3 reduced; the remaining lobes bear claw-like setae.
- Mxp reduced with 2 long spines on the 1st segment; the distal part pof the limb bearing 3 or 4 setae on the inner margin and 1 on the outer margin of the distal segment.
- Swimming legs with3-segmented exopods; endopod of P1 2 or 3-segmented; P2-P4 with 2- segmented endopodss.
- Female P5 simple, uniramous, 2-3-segmented; coxa and coupler fused; basis with an outer seta; free exopodal segment typically with 1 outer margin spine and 3 spinous processes arranged around the margins; exopod segment sometimes in the form of a curved tapering process.
- Male P5 asymmetrical, uniramous; right leg 3-segmentd, 1st segment unarmed, 2nd segment often expanded into a medial or distal lamellate process armed with up to 3 setae, 3rd segment forming a subchela typically curved and armed with 3 or 4 small setae; left leg 4-segmented, 1st segment unarmed, 2nd and 3rd segments elongate, each with an inner and outer seta, 4th segment curved, typically armed with 2 setae on the inner margin, 1 apical seta and 2 on the outer margin; the inner seta on the 3rd segment often carried on a slender process.

Diagnosis from Mulyadi (2004, p.164):
- Cephalon without lateral hook.
- Rostrum absent, horse shoe-shaped plate fringe with hairs anterior to upper lip.
- Eye large, without lenses.
- Cephalon and pediger somite 1 separated, pediger somites 4 and 5 fused or separated.
- Corners of last thoracic somite pointed and sometimes asymmetrical in female, rounded in male.
- Urosome 2 or 3 somites and usually asymmetrical in female; 5 somites in male.
- Caudal rami usually very asymmetrical in female, less so in male.
- A1 17-segmented; male right A1 prehensile, thickened at middle.
- Endopod of A2 little longer than exopod.
- Basis of Md long; outermost tooth of gnathobase large and well separated from other teeth.
- Mx1 reduced to basal segment bearing short broad proximal lobe and long narrow distal lobe, the latter with 2 long and 1 shorter apical setae.
- Mx2 with much reduced proximal lobe.
- Mxp with well-developed basal segment armed with strong setae; endopod much reduced.
- P1-P4 with 3 exopodal segment, 2 or 3 endopodal segments in P1, 2 endopodal segments in P2-P4.
- P5 uniramous in both sexes, 2-3-segmented and sometimes asymmetrical in female; right P5 male forming a chela.

Key to subgenera from Ohtsuka & Reid (1998, p.775) :
1 – Right caudal ramus of female bearing large acute outer process ; right caudal seta II of male much thicker and longer than left ; 2 nd endopod segment of leg P4 bearing 7 setae …….. Tortanus (Boreotortanus) Ohtsuka & Reid, 1998.
1’ – Caudal rami of both sexes not as above ; 2 nd endopod segment of leg P4 with 6 setae ……… 2.
2 – Mxp syncoxa beraing 5 setae ……. 3.
2’ - Mxp syncoxa bearing only 2 setae …… 4.
3 – In both sexes, dorsal terminal triangular process present on both caudal rami ; prosomal corners of female not produced ; caudal rami of male exyremely slender, as long as or longer than urosomites compined, 2 nd endopod segments of legs P2 and P3 each bearing 7 setae …….. T. (Tortanus) Giesbrecht, 1898.
3’ – In both sexes, dorsal terminal triangular process absent on both caudal rami ; prosomal corners of female produced posteriorly into winglike process ; caudal rami of male shorter than urosomites compbined ; 2 nd endopod segments of legs P2 and P3 each bearing 8 setae ……T. (Eutortanus) Smirnov, 1935.
4 – In both sexes, pedigers 4 and 5 separate ; female P5 with with 1-segmented exopodbearing either 4 spines or 3 spines plus 1 process ; in right P5 of male, exopod arising proximally from basis …….. T. (acutanus) Ohtsuka, 1992.
4’ – In both sexes, pedigers 4 and 5 fused ; female exopod of P5 either reduced or 1-segmented, tapering distally with spines/processes ; in right P5 of male, exopod arising distally from basis …….. T. (Atortus) Ohtsuka, 1992.

[1] Tortanus (Atortus) ampliramus  Ohtsuka, El-Sherbiny & Ueda, 2000   (F,M)    [Figs]

[2] Tortanus (Atortus) andamanensis  Nishida, Anandavelu & Padmavati, 2015   (F,M)    [Figs]

[3] Tortanus (Acutanus) angularis  Ohtsuka, 1992   (F,M)    [Figs]

[4] Tortanus (Tortanus) barbatus  (Brady, 1883)   (F,M)    [Figs]

[5] Tortanus (Atortus) bilobus  Mulyadi, Nishida & Ohtsuka, 2017   (F,M)    [Figs]

[6] Tortanus (Atortus) bonjol  Othman, 1987   (F,M)    [Figs]

[7] Tortanus (Atortus) bowmani  Othman, 1987   (F,M)    [Figs]

[8] Tortanus (Atortus) brevipes  A. Scott, 1909   (F)    [Figs]

Tortanus bumpusi    Wheeler, 1901   (F,M)
Syn.: Corynura bumpusii Wheeler, 1901 (p.184, figs.F,M)
Rem.: Cf. Tortanus discaudatus

[9] Tortanus (Atortus) capensis  Grindley, 1978   (F,M)    [Figs]

[10] Tortanus (Acutanus) compernis  Gonzalez & Bowman, 1965   (F,M)    [Figs]

Tortanus denticulatus    Giesbrecht, 1889   (F)
Syn.: Corynura denticulata Giesbrecht,1899; 1892 (p.525, 772, figs.F)
Ref.: Giesbrecht & Schmeil, 1898 (p.158); Sewell, 1912 (p.377, Rem.); Früchtl, 1924 b (p.15, Rem.); Greenwood, 1978 (p.18, Rem.); Hulsemann, 1988 (p.656, Rem.)
Loc: Mer Rouge
Rem.: Cf. Tortanus barbatus

Tortanus denticulatus    Shen & Lee, 1963   (F,M)
Ref.: Shen & Lee, 1963 (p.595, figs.F,M); Bowman, 1971 a (p.521); Hulsemann, 1988 a (p.656, Rem.)
Loc: Chine
Rem.: Cf. Tortanus sheni

[11] Tortanus (Eutortanus) derjugini  Smirnov, 1935   (F,M)    [Figs]

[12] Tortanus (Eutortanus) dextrilobatus  Chen & Zhang, 1965   (F,M)    [Figs]

[13] Tortanus (Atortus) digitalis  Ohtsuka & Kimoto, 1989   (F,M)    [Figs]

[14] Tortanus (Boreotortanus) discaudatus  (Thompson & Scott, 1897)   (F,M)    [Figs]

[15] Tortanus (Acutanus) ecornatus  Ohtsuka & Reid, 1998   (F,M)    [Figs]

[16] Tortanus (Atortus) erabuensis  Ohtsuka, Fukuura, Go, 1987   (F,M)    [Figs]

[17] Tortanus (Tortanus) forcipatus  Giesbrecht, 1889   (F,M)    [Figs]

[18] Tortanus (Atortus) giesbrechti  Jones & Park, 1968   (F,M)    [Figs]

[19] Tortanus (Tortanus) gracilis  (Brady, 1883)   (F,M)    [Figs]

[20] Tortanus (Atortus) indonesiensis  Mulyadi, Nishida & Ohtsuka, 2017   (M)    [Figs]

[21] Tortanus (Atortus) insularis  Ohtsuka & Conway, 2003   (F,M)    [Figs]

[22] Tortanus (Eutortanus) komachi  Itoh, Ohtsuka & Sato, 2001   (F,M)    [Figs]

[23] Tortanus (Atortus) longipes  Brodsky, 1948   (F,M)    [Figs]

[24] Tortanus (Atortus) lophus  Bowman, 1971   (F,M)    [Figs]

[25] Tortanus (Atortus) lukmani  Mulyadi, Nishida & Ohtsuka, 2017   (M)    [Figs]

[26] Tortanus (Atortus) manadoensis  Mulyadi, Nishida & Ohtsuka, 2017   (M)    [Figs]

[27] Tortanus (Atortus) magnonyx  Ohtsuka & Conway, 2005   (F,M)    [Figs]

[28] Tortanus (Atortus) minicoyensis  Francis & Bijoy Nandan, 2019   (F,M)    [Figs]

[29] Tortanus (Atortus) murrayi  A. Scott, 1909   (F,M)    [Figs]

[30] Tortanus (Atortus) nishidai  Ohtsuka, El-Sherbiny & Ueda, 2000   (M)    [Figs]

[31] Tortanus (Atortus) omorii  Mulyadi, Nishida & Ohtsuka, 2017   (M)    [Figs]

[32] Tortanus (Atortus) processus  Mulyadi, Nishida & Ohtsuka, 2017   (M)    [Figs]

[33] Tortanus (Atortus) recticaudus  (Giesbrecht, 1889)   (F,M)    [Figs]

[34] Tortanus (Atortus) rubidus  Tanaka, 1965   (F,M)    [Figs]

[35] Tortanus (Atortus) ryukyuensis  Ohtsuka & Kimoto, 1989   (F,M)    [Figs]

[36] Tortanus (Atortus) scaphus  Bowman, 1971   (F,M)    [Figs]

Tortanus (Atortus) scaphus variant       
Ref.: McKinnon, 1993 (p.1, figs.F, Rem.)
Loc: (860) F: 2,62; { F: 2,6

[37] Tortanus (Acutanus) setacaudatus  Williams, 1906   (F,M)    [Figs]

[38] Tortanus (Eutortanus) sheni  Hulsemann, 1988   (F,M)    [Figs]

[39] Tortanus (Atortus) sigmoides  Nishida, Anandavelu & Padmavati, 2015   (M)    [Figs]

[40] Tortanus (Atortus) sinicus  Chen, 1983   (F,M)    [Figs]

[41] Tortanus (Eutortanus) spinicaudatus  Shen & Bai, 1956   (F,M)    [Figs]

[42] Tortanus (Atortus) sulawesiensis  Mulyadi, Nishida & Ohtsuka, 2017   (F,M)    [Figs]

[43] Tortanus (Atortus) taiwanicus  Chen & Hwang, 1999   (F,M)    [Figs]

[44] Tortanus (Eutortanus) terminalis  Ohtsuka & Reid, 1998   (F,M)    [Figs]

[45] Tortanus (Atortus) tropicus  Sewell, 1932   (F,M)    [Figs]

[46] Tortanus (Atortus) tumidus  Chen, Hwang & Yin, 2004   (F,M)    [Figs]

[47] Tortanus (Eutortanus) vermiculus  Shen, 1955   (F,M)    [Figs]

Tortanus (Atotus) sp.    Ohtsuka, 1992   
Ref.: Ohtsuka, 1992 b (p.265, nom. nud.)
Loc: Sud Yemen

[48] Tortanus (Atortus) vietnamicus  Nishida & Cho, 2005   (F,M)    [Figs]

[49] Tortanus sp.  Mulyadi, 2004   (F)    [Figs]

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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2024. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at [Accessed May 30, 2024]

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