Species Card of Copepod
Calanoida ( Order )
    Clausocalanoidea ( Superfamily )
        Scolecitrichidae ( Family )
            Heteramalla ( Genus )
Heteramalla sarsi  Roe, 1975   (F)
Syn.: Heteramalla dubia (F) : Sars, 1907 a (p.17, Rem.F); A. Scott, 1909 ( p.86, figs.F); Sars, 1925 (p.142, figs.F); Sewell, 1929 (p.176); Rose, 1933 a (part., p.135, figs.F); Wilson, 1942 a (part., p.189); Grice & Hulsemann, 1965 (p.235); Vervoort, 1965 (part., p.28, F, Rem.); Vives, 1982 (p.291); Lozano Soldevilla & al., 1988 (p.58);
? H. dubia : Wilson, 1942 a (p.189); Grice & Hulsemann, 1967 (p.16);
no H. dubia : Rose, 1929 (p.26); no Massuti Alzamora, 1940 (p.6); 1942 (p.110, Rem.); Mazza, 1966 (p.70)
Ref.:
Roe, 1975 (p.342, Rem.); Bradford & al., 1983 (p.88); Vyshkvartzeva, 2005 (p.158, Redescr.F, Rem.); Vives & Shmeleva, 2007 (p.743, figs.F, Rem.)
Species Heteramalla sarsi - Plate 1 of morphological figuresIssued from : G.O. Sars in Résult. Camp. Scient. Prince Albert I, 69, pls.1-127 (1924). [Pl.XXXIX, figs.1-16]. As Heteramalla dubia.
Female: 1, habitus (dorsal); 2, idem (lateral left side); 3, forehead, lateral); 4, rostrum (frontal view); 5, A1; 6, A2; 7, Md; 8, Mx1; 9, Mxp; 10, two forms of sensory filaments; 11, Mxp; 12, P1; 13, P2; 14, P3; 15, P4; 16, P5.


Species Heteramalla sarsi - Plate 2 of morphological figuresissued from : A. Scott in Siboga-Expedition, 1909, XIX a. [Plate XXXIII, Figs.1-9]. As Heteramalla dubia.
Female (from Halmahera Sea): 1, habitus (dorsal); 2, forehead (lateral); 3, last thoracic and genital segments (left side); 4, rostrum; 5, A1; 6, Mx2 (distal portion); 7, P2; 8, part of terminal spine (exopodite of P2); 9, P5.


Species Heteramalla sarsi - Plate 3 of morphological figuresIssued from N.V. Vyshkvartzeva in Zoosyst. Rossica, 2005, 13 (2). [p. 159, Figs.1-9].
Female (from 45°20'N,153°54'E): 1, habitus (lateral); 2, forehead (lateral); 3, rostrum; 4, Thoracic segment 5 and urosome (right lateral); 5, genital double-somite (ventral); 6, A1; 7, distal segments of A1; 8, Md; 9, Md (masticatory edge of gnathobase).


Species Heteramalla sarsi - Plate 4 of morphological figuresIssued from N.V. Vyshkvartzeva in Zoosyst. Rossica, 2005, 13 (2). [p. 160, Figs.10-13].
Female: 10, A2; 11, Mx1; 12, Mx2; 13, Mxp.


Species Heteramalla sarsi - Plate 5 of morphological figuresIssued from N.V. Vyshkvartzeva in Zoosyst. Rossica, 2005, 13 (2). [p. 161, Figs.14-19].
Female: 14, P1; 15, P2 (exopodal segments 2-3 detached); 16, P3; 17, apical spine of P3; 18, P4; 19, P5.


Species Heteramalla sarsi - Plate 6 of morphological figuresIssued from N.V. Vyshkvartzeva in Zoosyst. Rossica, 2005, 13 (2). [p. 162, Table 1].
Sampling localities, size and morphological variation of Heteramalla sarsi Roe, 1975.Nota: Female specimens described from the North Pacific, generally agreewith Sars' (1924-25) description of his Atlantic specimens, except that they lack a seta on inner lobe 2 of Mx1 (this feature conforms with Roe's, 1975 Atlantic specimens, but possibly it is a variable feature in this species) and bear 1 plumose and 1 aesthetasc-like seta on inner lobe 3 of Mxp syncoxa (feature described by Vyshvartzeva for the first time). Sars did not describe Mxp, but the illustration shows only 1 seta with a tapering tip, which is transformed in the Pacific sample by a aesthetasc-like seta on inner lobe 3. There are some other discrepancies in the descriptions of H. sarsi.
It should be mentioned that Massuti Alzamora (1940) misindentified another species as 'Heteramalla dubia'; actually, his material fits neither H. sarsi nor Scopalatum dubium.

Compl. Ref.:
Deevey & Brooks, 1977 (p.256, Table 2, station "S")
NZ: 6 + 1 doubtful

Distribution map of Heteramalla sarsi by geographical zones
Loc:
off Bermuda: Station ‘’ S’’ (32°10’N, 64°30’W), G. of Guinea, off Morocco-Mauritania, off NW Canary Is., W Indian, Indonesia-Malaysia, off E Philippines, off E Kuril Is., ? [ Hawaii, off Is. Christmas NE, off S Easter Is. (in C.B. Wilson, 1942 a, 1950) ]
N: 8
Lg.:
(1) F: 3,7; (3) F: 3,65-3,41; (5) F: 3,78; (941) F: 3,41-3,78; {F: 3,41-3,78}
Rem.: Bathypelagic. Sargasso Sea: 1500-2000 m (Deevey & Brooks, 1977, station "S");
In the faunistic listings where the sex is not always specified, the locality records are preceded by ?
Last update : 23/03/2015
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