Species Card of Copepod
Calanoida ( Order )
    Clausocalanoidea ( Superfamily )
        Tharybidae ( Family )
            Tharybis ( Genus )
Tharybis macrophthalma  Sars, 1902   (F,M)
Ref.:
Sars, 1902 (1903) (p.68, figs.F,M); Lysholm & Nordgaard, 1921 (p.22); Rose, 1933 a (p.164, figs.F,M); Bradford & al., 1983 (p.126); Ferrari & Markhaseva, 1996 (p.284, figs.F,M); Andronov, 2002 (p.43, figs.F,M); Ferrari & Markhaseva, 2005 (p.46: tab.1); Boxshall & Halsey, 2004 (p.211, figs.F,M); Vives & Shmeleva, 2007 (p.859, figs.F,M, Rem.); Markhaseva & Renz, 2011 (p.67, Fig.M) ; Laakmann & al., 2019 (p.330, fig. 3, 4, phylogenetic relationships)
Species Tharybis macrophthalma - Plate 1 of morphological figuresissued from : G.O. Sars in An Account of the Crustacea of Norway. Vol. IV. Copepoda Calanoida. Published by the Bergen Museum, 1903. [Pl. XLV].
Female & Male.
R = rostrum (frontal view); Urs = urosome (dorsal).

Nota: Eye of female very large and conspicuous, oblong quadrangular in form, and occuring close to the dorsal surface, pigment bright red. A1 in the living animal are born in a somewhat unusual manner, exhibiting at the base an abrupt curve, and pointing obliquely bacwards.


Species Tharybis macrophthalma - Plate 2 of morphological figuresissued from : G.O. Sars in An Account of the Crustacea of Norway. Vol. IV. Copepoda Calanoida. Published by the Bergen Museum, 1903. [Pl. XLVI].
Female.
M = Md; m = Mx1; mp1 = Mx2; mp2 = Mxp.


Species Tharybis macrophthalma - Plate 3 of morphological figuresissued from : F.D. Ferrari & E.L. Markhaseva in Proc. Biol. Soc. Washington, 1996, 109 (2). [p.283, Fig.11].
Female and Male (from Raunefjord, Norway): A, Mx1 (anterior); B, Mx1 (tip of inner lobe 1, posterior); C, Mx2 (inner lobes 1-4 on syncoxa, posterior); D, Mx2 (inner lobe 5 on basis and innerl lobe 6 + exopod, anterior); E, Mxp (syncoxa); F, basis and endopod (setation of endopod not shown); G, Mxp (endopod, b= distal tip of basis); H, P1 (endopod (anterior).
Bars: 1 = 0.1 mm (E, F); 2 = 0.1 mm (A, B); 3 = 0.1 mm (C, D, G); 4 = 0.1 mm (H).

Nota: Mx1: Lateral lobe with 9 setae; exopod 1-segmented with 5 setae; basoendopod with sets of 3, 2, 2, and 3 setae; medial lobe 2 with 3 setae, medial lobe 3 with 4 setae, medial lobe 1 with 9 apical setae (5 thick with setules, 4 thin and unarmed) and 4 posterior setae. Mx2 with medial lobes 1-4 of coxa each with 4, 3, 3, 3 setae each; medial lobe 5 on basis with 4 sclerotized setae; medial lobe 2 + exopod an indistinctly segmented complex with 8 seta (terminal 3 Bradford's setae without setules, 4 Bradford's setae with apical setules and 1 sclerotized seta with setules). P1 with Von Vaupel Klein's organ on the lateral margin of ebndopod extended distally; denticles medial and lateral to the extension, and on the anterior face of endopod below the basal seta.


Species Tharybis macrophthalma - Plate 4 of morphological figuresissued from : V.N. Andronov in Arthropoda Selecta, 2002, 11 (1). [p.45, Figs.206-212]. After Sars, 1903.
Female: 206-207, habitus (dorsal and lateral, respectively); 208, rostrum; 209, urosome (dorsal); 210, P5.

Male: 211, habitus (right lateral side); 212, P5.


Species Tharybis macrophthalma - Plate 5 of morphological figuresissued from : V.N. Andronov in Arthropoda Selecta, 2002, 11 (1). [p.46, Figs.213-219].
Female (from NE Atlantic): 213, rostrum; 214, urosome (ventral); 215, idem (left lateral side); 216, P5.

Male: 217, P5; 218-219, distal end of exopod of left P5.


Species Tharybis macrophthalma - Plate 6 of morphological figuresissued from : E.L. Markhaseva & J. Renz in Zootaxa, 2011, 2889. [p.67, A]. Changed and schematized after Andronov (2002).
Male: A, P5 (Nota: right basis in grey).

Compl. Ref.:
Furnestin, 1960 (p.172); Mazza, 1966 (p.71); Matthews, 1967 (p.159, Table 1, Rem.); Hure & Krsinic, 1998 (p.53, 101); Vukanic, 2003 (139, tab.1); Markhaseva & Schnack-Schiel, 2003 (p.112, Rem.); Valdés & al., 2007 (p.104: tab.1); Belmonte, 2018 (p.273, Table I: Italian zones)
NZ: 3

Distribution map of Tharybis macrophthalma by geographical zones
Loc:
Morocco, Canary Is., Great Meteor Seamount (28-30°N,28° W), Azores, off NW Ireland, Norway (Christiana Fjord, off W coast, Raunefjorden (all the year), Trondheim Fjord), North Sea, NW Spain (Ria de Arosa), Medit. (G. of Lion, S Adriatic Sea)
N: 11
Lg.:
(65) F: 1,4; M: 1,3; {F: 1,40; M: 1,30}
Rem.: hyperbenthic (depth 30-200 m).
Last update : 06/02/2020
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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2021. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed October 21, 2021]

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