Species Card of Copepod
Harpacticoida ( Order )
    Clytemnestridae ( Family )
        Goniopsyllus ( Genus )
Goniopsyllus clausi  Huys & Conroy-Dalton, 2000   (F,M)
Syn.: Clytemnestra rostrata : Giesbrecht, 1892 (p.568, 771, figs.F,M); ? Monard, 1928; Vilela, 1965 (p.21, figs.M); 1968 (p.44, figs.F); Razouls, 1972 (p.96, Annexe: Fig.F); Huys & al., 1996 (p.300, figs.F,M); Boxshall & Huys, 1998 (p.781, figs.F,M)
Ref.:
Huys & Conroy-Dalton, 2000 (p.29, figs.F,M, Rem.); Huys & Boxshall, 2004 (p.278); Vives & Shmeleva, 2010 (p.97, figs.F,M, Rem.)
Species Goniopsyllus clausi - Plate 1 of morphological figuresissued from : Huys and Conroy-Dalton in Bull. nat. Mus. Lond., 2000, 66 (1). [p.30, Fig.19].
Female (from Bay of Cadiz, Spain): A, habitus (dorsal); B, habitus of ovigerous (lateral).
Scale bar in micrometers.


Species Goniopsyllus clausi - Plate 2 of morphological figuresissued from : Huys and Conroy-Dalton in Bull. nat. Mus. Lond., 2000, 66 (1). [p.31, Fig.20].
Female: A, A1; B, distal portion of A1 segment 6 (ventral; rudimentary element arrowed).

Male: C, A1 (ventral); D, A1 segments 3-6; E, A1 segment 78 (ventral; rudimentary element arrowed).
Scale bars in micrometers.


Species Goniopsyllus clausi - Plate 3 of morphological figuresissued from : Huys and Conroy-Dalton in Bull. nat. Mus. Lond., 2000, 66 (1). [p.32, Fig.21].
Female: A, A2 (outer); B, distal endopod segment of A2 (inner; rudimentary segment arrowed); C, Md; D, Md (mandibular gnathobase); F, Mx1 (posterior); G, Mx2 (posterior; exit of maxillary gland arrowed); H, oral area showing position of A2, labbrum, paragnethes (P), Md, Mx1, Mx2 and Mxp (Mxp); I, rostrum (dorsal)..

Male: E, Md (mandibular gnathobase).
Scale bars in micrometers.


Species Goniopsyllus clausi - Plate 4 of morphological figuresissued from : Huys and Conroy-Dalton in Bull. nat. Mus. Lond., 2000, 66 (1). [p.33, Fig.22].
Female: A, Mxp; B, distal portion of basis and endopod of Mxp (anterior); C, same (medial; D, same (posterior).

Male: Mxp (anterior); F, distal portion and endopod of Mxp (anterior); G, maxillipedal basis and endopod (medial); H, same (posterior).
Scale bars in micrometers.


Species Goniopsyllus clausi - Plate 5 of morphological figuresissued from : Huys and Conroy-Dalton in Bull. nat. Mus. Lond., 2000, 66 (1). [p.35, Fig.23].
Female: A, urosome (ventral); B, anal somite and left caudal ramus (dorsal); C, genital field (ventral); D, same (lateral); E, P1.
Scale bars in micrometers.


Species Goniopsyllus clausi - Plate 6 of morphological figuresissued from : Huys and Conroy-Dalton in Bull. nat. Mus. Lond., 2000, 66 (1). [p.36, Fig.24].
Female: A, P2 (anterior); B, P3 (anterior); C, P5 (anterior); D, aberrant P5 (anterior).
Scale bar in micrometers.


Species Goniopsyllus clausi - Plate 7 of morphological figuresissued from : Huys and Conroy-Dalton in Bull. nat. Mus. Lond., 2000, 66 (1). [p.37, Fig.25].
Female: B, P4 (anterior).

Male: A, habitus (dorsal); C, P5 (anterior.
Scale bars in micrometers.


Species Goniopsyllus clausi - Plate 8 of morphological figuresissued from : Huys and Conroy-Dalton in Bull. nat. Mus. Lond., 2000, 66 (1). [p.38, Fig.26, A-B].
Female: A, caudal ramus (lateral).

Male: B, urosome (ventral).
Scale bars in millimeters.


Species Goniopsyllus clausi - Plate 9 of morphological figuresissued from : Huys and Conroy-Dalton in Bull. nat. Mus. Lond., 2000, 66 (1). [p.39, Fig.27, C-D]. SEM photographs.
Female: C, genital double-somite (circular depression surronuding copulatory pore); D, genital aperture.


Species Goniopsyllus clausi - Plate 10 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [taf.45, Fig.22].
Male: abdominal segment 1 (ventral).


Species Goniopsyllus clausi - Plate 11 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.45, Fig.31].
Male: 31, A1 (over view).


Species Goniopsyllus clausi - Plate 12 of morphological figuresissued from : C. Razouls in Th. Doc. Etat Fac. Sc. Paris VI, 1972, Annexe. [Fig.100]. As Clytemnestra rostrata.
Female (from Banyuls: G du Lion): A, anal segment and caudal rami; B, P1; C, P2; D, P3; E, P4; F, P5.


Species Goniopsyllus clausi - Plate 13 of morphological figuresIssued from : K.H. Cho, W.-S. Kim & W. Lee in Proc. Biol. Soc. Washigton, 2010, 123 (2). [p.134, Table 1].
Diagnostic characteristics.

Nota: Compare with G. rostratus, G. dokdoensis, G. brasiliensis.

Compl. Ref.:
Tachibana & al., 2013 (p.545, Table 1, seasonal change 2006-2008)
NZ: 3

Distribution map of Goniopsyllus clausi by geographical zones
Loc:
Portugal (coast), Ibero-moroccan Bay (Bay of Cadiz), Mediterranean Sea (Banyuls, Napoli, Adriatic Sea), Japan (Tokyo Bay).

Type locality: 36°30' N, 7°20' W.
N: 5
Lg.:
(931) F: 0,979-1,067; M: 1,021; {F: 0,98-1,07; M: 1,02}
Rem.:

Identification characters after G.A. Boxshall & S.H. Halsey (2004, p.278):
1- Exopod of P1 with 3 distal setal elements; female A1 6-segmented.
2- Exopod of P1 with 3 distal setal elements; female A1 6-segmented.
2- Longer of 2 outer proximal bipinnate setae on female caudal ramus reaching well beyond distal margin of ramus.
3- Genital double-somite of female just longer than wide, with lateral margins slightly concave just posterior to mid-level.
Last update : 13/09/2015
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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2021. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed October 24, 2021]

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