Species Card of Copepod
Calanoida ( Order )
    Clausocalanoidea ( Superfamily )
        Stephidae ( Family )
            Stephos ( Genus )
Stephos grievae  Krsinic, 2015   (F,M)
Krsinic, 2015 (p.1, Descr. F, M, figs. F, M, Rem.)
Species Stephos grievae - Plate 1 of morphological figuresIssued from : F. Krsinic in Mar. Biodiv. Rec., 2015, 8. [p.3, Fig.1].
Female (from 42°47'42.2''N, 17.5°23'22.3''E): A-B, habitus (dorsal and lateral, respectively); C, rostral area (ventral).

Nota: ratio of prosome and urosome as 2.9 : 1.
- Rostral area, with 2 sensillae medially, rostrum represented by rounded knob.
- Urosome 4 free somites; proportional lengths of urosomites and caudal rami 37.4 : 18.6 : 17 : 13.5 : 13.5 = 100.
- Caudal rami symmetrical, 1.25 times longer than wide; ramus with 4 terminal setae corresponding to setae III-VI, seta VII small, pinnate, setae I and II absent

Species Stephos grievae - Plate 2 of morphological figuresIssued from : F. Krsinic in Mar. Biodiv. Rec., 2015, 8. [p.4, Fig.2].
Female: A, A1 segments 1-15; B, A1 segments 16-24; C, urosome (dorsal); D, genital double-somite (ventral).

Nota: Genital double-somite about as long as wide, symmetrical, with proximolateral margins produced in dorsal aspect. External genital area symmetrical in venytral view, paired egg-laying duct, paired gonopores situated medially near posterior edge, covered by single operculum.
- A1 symmetrical, 24-segmented, reaching almost to end of 3rd urosomite.

Species Stephos grievae - Plate 3 of morphological figuresIssued from : F. Krsinic in Mar. Biodiv. Rec., 2015, 8. [p.5, Fig.3].
Female: A, A2; B, Md; C, Mx1; D, Mxp; E, Mx2.

Nota: A2 with exopod (7-seggmented) longer than endopod (2-segmented).
- Md gnathobase cutting edge with isolated unicuspid tooth, 8 heterogeneous teeth and 1 dorsal spinulose seta. Endopod 2-segmented (1st segment with 4 unequal setae and terminal segment truncate bearing 11 setae). Exopod 5-segmented (setal formula: 1, 1, 1, 1, 2).
Mx1 praecoxal arthrite bearing 9 stout marginal spines and 4 elements on posterior surface; coxal epipodite with 9 and coxal endite with 4 setae; proximal basal endite with 3 setae, distal basal endite with 5 setae, basal exite unarmed; exopod bearing 11 setae; endopod articulated to basis, indistinctly 3-segmented, with setal formula 3, 4, 7.
- Mx2 praecoxal endites with 6 and 3 setae, respectively; coxal endites each with 3 setae; allobasis with proximal endite powerfully developed with 3 setae, 1 stouter than rest; endopod 3-segmented, with setal formula 1, 1, 4.
- Mxp praecoxal endite with 1 seta and row of spinules; syncoxal endites, setal formula 2, 3, 3on medial margin, distal seta thorn-like; distal margin with row of spinules; basis with longitudinal row of spinules and with 3 setae, free endopod 6-segmented, armature formula 2, 4, 4, 3, 3+1, 4.

Species Stephos grievae - Plate 4 of morphological figuresIssued from : F. Krsinic in Mar. Biodiv., 2015, 8. [p.6, Fig.4].
Female: A, P1; B, P2; C, P3; D, P4; E, P5.

Nota: P5 symmetrical, uniramous, 3-segmented, proximal segment separated fom intercoxal sclerite; 3rd segment very long, armed with setules distally along both margin; distal segment armed with single small spine on anterior surface and spiniform process laterally.

Species Stephos grievae - Plate 5 of morphological figuresIssued from : F. Krsinic in Mar. Biodiv. Rec., 2015, 8. [p.2].
Female and Male : Armature formula in swimming legs P1 to P4.

Species Stephos grievae - Plate 6 of morphological figuresIssued from : F. Krsinic in Mar. Biodiv. Rec., 2015, 8. [p.7, Fig.5].
Male: A-B, habitus (dorsal and lateral, respectively); C, urosome (dorsal).Nota: Rostral area as in female.
- Prosome: urosome ratio = 2.85: 1.
- Urosome 5 somites; proportional lengths of urosomites 30 : 16.5 : 20 : 18.5 : 15 = 100.
- genital somite slightly asymmetrical viewved dorsally, wider than long; genital aperture located on left side.
Caudal rami as in female.
Armature of A2 and swimming legs P1 to P4 as in female.

Species Stephos grievae - Plate 7 of morphological figuresIssued from : F. Krsinic in Mar. Biodiv. Rec., 2015, 8. [p.8, Fig.6].
Male: A, A1 segments 1-15; B, A1 segments 16-24; C, right P5; D, E, F, left P5 (F: segments 4 and 5).
-P5 uniramous and asymmetrically. Right leg 4-segmented and very long; esgment 2 (basis) longer than coxa, segment 4 very elongate, distal part is more robust than proximal, hook-like distally with thickened knee-shaped projection and inner margin with small denticles. left leg 5-segmented, segments unarmed, segment 2 (basis) elongate and robust, segments 3 and 4 shorter and of similar length, segment 5 crescent -shaped, longer than segment 2, articulated to segment 4 at approximately 1/4 of length of segment, shorter part in form of heavy, tapering processes, while longer part forms a club-shaped curved extension.

NZ: 1

Distribution map of Stephos grievae by geographical zones
Lenga Pit, Mljet Island (S Adriatic)

Type locality: 42° 47' 42.2'' N, 17.5° 23' 22.3'' E.
N: 1
(1196)* F: 0, 60-0,67; M: 0,55-0,62.
* : excluding caudal rami.
Rem.: Lenga Pit (Mljet Island) is about 100 m from the shoreline, there is a small 9 m deep lake at the end of the cave (Salinity in the water column below depths of 3 m is between 21.00 and 39.00, temperature approximately 15.5°C.
For Krsinic (2015, p.5) the species with male P5 is typical of species group IV (see Bradford-Grieve, 1999).
Last update : 12/05/2016
    to add your remark

 Any use of this site for a publication will be mentioned with the following reference :

Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2024. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed July 21, 2024]

© copyright 2005-2024 Sorbonne University, CNRS

CNRS   Observatoire Océanologique de Banyuls sur Mer - Laboratoire Arago
Sorbonne Université


Version française
English version



On the WEB of CNRS


Marine Planktonic Copepods

Marine Planktonic Copepods


Imprimer Contact Accueil Plan du site Accès restreint Retour Une du Labo Imprimer Contact Plan du site Crédits Téléchargez les Plug-Ins