Species Card of Copepod
Calanoida ( Order )
    Arietelloidea ( Superfamily )
        Augaptilidae ( Family )
            Heteroptilus ( Genus )
Heteroptilus attenuatus  (Sars, 1905)   (F, ? M)
Syn.: Pontoptilus attenuatus Sars,1905 c (p.20, Rem.F);
? Heteroptilus sp. : Sewell, 1947 (p.243, figs.M).
Ref.:
Sars, 1925 (p.324, figs.F); Rose, 1933 a (p.237, figs.F); Vives & Shmeleva, 2007 (p.259, figs.F, Rem.)
Species Heteroptilus attenuatus - Plate 1 of morphological figuresIssued from : G.O. Sars in Résult. Camp. Scient. Prince Albert I, 69, pls.1-127 (1924). [Pl.CXVI, figs.1-15].
Female: 1, habitus (dorsal); 2, idem (lateral left side); 3, forehead (lateral); 4, rostral prominence; 5, A2; 6, Md; 7, right Md (masticatory edge); 8, labrum wiith falciform process; 9, Mx1; 10, Mx2; 11, Mxp; 12, P1; 13, P3; 14, P5; 15, distal segments of urosome and caudal rami (dorsal).


Species Heteroptilus attenuatus - Plate 2 of morphological figuresissued from : R.B.S. Sewell in The John Murray Expedition, 1933-34, Scientific Reports, VIII (1), 1947. [p.244, Fig.67]. As Heterorhabdus sp. With doubt.
Male (from N Arabian Sea): A, left A1; B, A2; C, Md (mandibular palp and base of biting blade); D, curved process on posterior lip of mouth; E, Mx1; F, Mx2; G, P1; H, P3; I, right P5; J, left P5.
Nota: Proportional lengthsof the prosome and urosome as 75 to 25. Proportional lengths of the various segments of the body (cephalon to caudal rami) as 394:115:71:71:99:51:41:38:33:18:69 = 100. Posterior thoracic margin of 5th segment is produced backwards in a wing, the apex of which is rounded and bears a minute hair or seta. Left A1 modified into grasping organ.; the 17th segment is produced in a straight horizontal spine that overhangs the 18th segment.; segments 19 to 21 fused and the combined joint bears 2 straight spines, of which the distal is the longer.; segments 22 and 23 fused, 24 and 25 separate. The proportional lengths of the exopod and endopod of A2 as 53 to 47.Md asymmetrical; on the right side the biting edge is produced on its anterior aspect in a long curved tooth (similar to that in the female H. attenuatus; in left side the biting edge bears a series of 5 teeth and 1 seta. In Mx2, the 5th lobe bears 6 setae, of which 4 are serrated, and the 6th is a stout curved spine with a few teeth on its margin.

Compl. Ref.:
Sewell, 1948 (p.505); C.B. Wilson, 1950 (p.239); Lozano Soldevilla & al., 1988 (p.60); Suarez-Morales & Gasca, 1998 a (p.108); Chaouadi & Hafferssas, 2018 (p.913, Table II: occurrence).
NZ: 6

Distribution map of Heteroptilus attenuatus by geographical zones
Loc:
W Medit. (Alboran Sea, Habibas Islands), Bay of Biscay, Canary Is., Azores, Caribbean Sea, N Arabian Sea, Pacif. ( off E Tuamotu Is., off E Easter Is.)
N: 4
Lg.:
(1) F: 5,7; {F: 5,70}
Rem.: bathypelagic? (0-4800 m).
Populations possibly isolated at the time of the closure of the isthmus of Panama.
Perhaps cited for the first time in the Alboran Sea by Chaouadi & Hafferssas ( 2018 (p.913, Table II).
Last update : 28/02/2020
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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2021. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed October 24, 2021]

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