Species Card of Copepod
Calanoida ( Order )
    Clausocalanoidea ( Superfamily )
        Clausocalanidae ( Family )
            Clausocalanus ( Genus )
Clausocalanus ingens  Frost & Fleminger, 1968   (F,M)
Syn.: Clausocalanus arcuicornis : De Decker, 1964 (part., p.15, 20, 30);
C. laticeps : Tanaka, 1960 (p.32, figs.F,M) data from Frost & Fleminger;
C. arcuicornis major Unterüberbacher, 1964 (part., p.19, Pl.27)
Ref.:
Frost & Fleminger, 1968 (p.39, figs.F,M, Rem.); Ramirez, 1969 (p.53, figs.F, Rem.); Björnberg & al., 1981 (p.628, figs.F,M); Bradford-Grieve, 1994 (p.114, figs.F,M, fig.101); Mazzocchi & al., 1995 (p.153, figs.F, Rem.); Bradford-Grieve & al., 1999 (p.878, 916, figs.F,M); Bucklin & al., 2003 (p.335, tab.2, figs.1, 4, Biomol)
Species Clausocalanus ingens - Plate 1 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [Pl.19, p.138-139; Pl.20, p.140-141; Pl.21, p.142-143].

Female: 1 a, habitus (right lateral view); 1 b, idem (dorsal view); 2 a, frontal region (right lateral); 2 b, rostrum ( (right lateral); 2 c, rostrum (anteroventral); 2 d, Th.4-5 (posterior part) and genital segment (right lateral); 2 z, genital segment (ventral view); 2 a-b taken from one specimen; 2 c-e from different specimens; 3 a, Th.4-5 (posterior part) and urosome with spermatophore attached (right lateral); 3 b, B2 of P2; 3 c, B2 of P3; 3 d-e, P5; 3 a taken from one specimen; 3 b-c from another specimen; 3 d-e from third and fourth specimens, respectively.


Species Clausocalanus ingens - Plate 2 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [Pl.22, p.144-145; Pl.23, p.146-147].

Male: 1 a, habitus (right lateral view); 1 b, idem (dorsal view); 1 c, frontal region (right lateral); 1 d, urosome (dorsal view); 1 e, urosome 4 (posterior part), urosome 5 and caudal rami (dorsal view); 1 a-b taken from one specimen; 1 c-e taken from different specimens; 2 a, Th.4-5 (posterior part) and urosome (left lateral view); 2 b, B2 of P2; 2 c, B2 of P3; 2 d, P5 (posterior); 2 e, P5 (left anterolateral); 2 a taken from one specimen; 2 b-c from another ; 2 d and e from third and fourth specimens, respectively.
Nota: 2nd segment of P5 less than 4.4 times as long as wide.


Species Clausocalanus ingens - Plate 3 of morphological figuresissued from : J.M. Bradford-Grieve in The Marine Fauna of New Zealand: Pelagic Calanoid Copepoda. National Institute of Water and Atmospheric Research (NIWA). New Zealand Oceanographic Institute Memoir, 102, 1994. [p.114, Fig.64].
Female: A, habitus (lateral left side); B, genital somite (lateral left side); C, rostrum (anterodorsal) view); D, basipod 2 of P3; E, P5.

Male: F, habitus (lateral left side); G, left caudal ramus (dorsal); H, basipod 2 of P3; I, P5.


Species Clausocalanus ingens - Plate 4 of morphological figuresIssued from: M.G. Mazzocchi, G. Zagami, A. Ianora, L. Guglielmo & J. Hure in Atlas of Marine Zooplankton Straits of Magellan. Copepods. L. Guglielmo & A. Ianora (Eds.), 1995. [p.154, Fig.3.26.1].
Female: A, habitus (dorsal); B, urosome (dorsal); C, habitus (lateral left side); D, urosome (lateral left side).
Nota: Proportional lengths of urosomites and furca 36:20:17:12:15 = 100. A1 almost reaching end of cephalothorax.


Species Clausocalanus ingens - Plate 5 of morphological figuresIssued from: M.G. Mazzocchi, G. Zagami, A. Ianora, L. Guglielmo & J. Hure in Atlas of Marine Zooplankton Straits of Magellan. Copepods. L. Guglielmo & A. Ianora (Eds.), 1995. [p.155, Fig.3.26.2].
Female (SEM preparation): B, urosome (dorsal); C, idem (ventral); D, P5.
Bars: B, C 0.100 mm; D 0.010 mm.


Species Clausocalanus ingens - Plate 6 of morphological figuresissued from : F.C. Ramirez in Contr. Inst. Biol. mar., Buenos Aires, 1969, 98. [p.56, Lam. IX, fig.62].
Female (from off Mar del Plata): 62, P5
Scale bar in mm: 0.02.


Species Clausocalanus ingens - Plate 7 of morphological figuresissued from : F.C. Ramirez in Contr. Inst. Biol. mar., Buenos Aires, 1969, 98. [p.50, Lam. VIII, figs.49-51].
Female (from off Mar del Plata): 49-51, basipodal segment 2 of P2-P4, respectively.
Scale bar in mm: 0.05.


Species Clausocalanus ingens - Plate 8 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.30, Table 2a].
Clausocalanus ingens Females: Measurements and ratios *.


Species Clausocalanus ingens - Plate 9 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.31, Table 2b].
Clausocalanus ingens males: Measurements and ratios *.
r = sample range; m = sample mean; n = number of specimens measured; s = sample standard deviation.


Species Clausocalanus ingens - Plate 10 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.26, Fig.9].
Ratio of length-to-width of 2nd segment of P5 male (ordinate plotted against length of 2nd segment of left P5 (abscissa) for Clausocalanus ingens.


Species Clausocalanus ingens - Plate 11 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.143, Pl.21, b, c, d-e].
Female: b, basipodite 2 of P2; c, basipodite 2 of P3; d-e, P5 (from different specimens).


Species Clausocalanus ingens - Plate 12 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.47, Pl.23, a, d, e].
Male: a, posterior part of last thoracic segment and urosome (left lateral); d, P5 (posterior); e, P5 (left anterolateral).


Species Clausocalanus ingens - Plate 13 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.147, Pl.23, b, c].
Male: b, basipodite 2 of P2; c, basipodite 2 of P3.


Species Clausocalanus ingens - Plate 14 of morphological figuresIssued from : O. Tanaka in Spec. Publs. Seto mar. biol. Lab., 10, 1960 [Pl. XI, 1-8]. As Clausocalanus laticeps.
Female (from Sub-Antarctic): 1, forehead (dorsal); 2,


Species Clausocalanus ingens - Plate 15 of morphological figuresIssued from : O. Tanaka in Spec. Publs. Seto mar. biol. Lab., 10, 1960 [p.32]. As Clausocalanus laticeps.
Female: Proportional lengths of segments of A1.


Species Clausocalanus ingens - Plate 16 of morphological figuresssued from : O. Tanaka in Spec. Publs. Seto mar. biol. Lab., 10, 1960 [Pl. XI, 9-12]. As Clausocalanus laticeps.
Male (from Sub-Antarctic): 9, habitus (lateral); 10, P2; 11, distal segment of left P5; 12, right P5.


Species Clausocalanus ingens - Plate 17 of morphological figuresIssued from : O. Tanaka in Spec. Publs. Seto mar. biol. Lab., 10, 1960 [p.33]. As Clausocalanus laticeps.
Male: Proportional lengths of segments of A1.

Compl. Ref.:
Björnberg, 1973 (p.311, 385); Nyan Taw & Ritz, 1979 (p.196); De Decker, 1984 (p.315, 343: carte); Razouls & al., 2000 (p.343, tab. 3, Appendix); Ward & al., 2003 (p.121, tab.4); Lo & al., 2004 (p.89, tab.1); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical, Fig.4, 6, 7, Tabe 4); Hidalgo & al., 2012 (p.134, Table 2); Zakaria & al., 2016 (p.1, Table 1, Rem.)
NZ: 9

Distribution map of Clausocalanus ingens by geographical zones
Species Clausocalanus ingens - Distribution map 4issued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [p.40, Chart 3, a].
Occurrence of C. ingens in samples examined; closed circles represent samples examined; open circles represent samples in which adults were found; bars through open circles represent samples from which specimens were removed for measurements.
Species Clausocalanus ingens - Distribution map 5issued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [p.41, Chart 3, b].
Occurrence of C. ingens in samples examined; closed circles represent samples examined; open circles represent samples in which adults were found; bars through open circles represent samples from which specimens were removed for measurements.
Loc:
sub-Antarct. (N South Georgia, SW & SE Pacif., Straits of Magellan (Central part and Pacific area), S Pacif., S Australia, Tasmania , New Zealand (N, SE North Island), China Seas (East China Sea), Taiwan (N: Mienhua Canyon), Chile, off Santiago, S Indian, South Africa (E & W), SW Atlant., S Brazil , E Medit. (W Egyptian coast in Zakaria & al., 2016 (Table 1)
N: 16
Lg.:
(30) F: 1,9-1,44; M: 1,08-0,99; (36) F: 1,84;1,67; (66) F: 1,62; M: 1,07; {F: 1,44-1,90; M: 0,99-1,08}

The mean female size is 1.694 mm (n = 5; SD = 0.1832) and the mean male size is 1.047 (n = 3; SD = 0.0493). The size ratio (male : female) is 0.64.
Rem.: epipelagic.
Sampling depth (sub-Antarct.) : 0-500 m.
For Frost & Fleminger (1968, p.41) C. ingens female is distinguished from laticeps by the length-width ratio of the caudal ramus and the form and spacing of basipodite 2 spiniform processes 2 and 3 of P3. C. ingens male differs from laticeps in the Prosome/Urosomal segment 2 length ratio, the form of basipodite 2 spiniform processes of P3, and the length-width ratio of the caudal ramus. C. ingens was confused with laticeps by Tanaka (1960), and it is likely that the two species have been confused by others.
Lo & al. (2004) seem to indicate for the first time the presence of this species in the NW Pacific (north of Taiwan), and Zakaria & al. (2016) in the East Mediterranean Sea (Egyptian coast).
Last update : 18/08/2018
    to add your remark
  

 Any use of this site for a publication will be mentioned with the following reference :

Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2021. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed October 23, 2021]

© copyright 2005-2021 Sorbonne University, CNRS

Webmaster
CNRS   Observatoire Océanologique de Banyuls sur Mer - Laboratoire Arago
Sorbonne Université

 

Version française
English version

 

Search

On the WEB of CNRS

 


Marine Planktonic Copepods

Marine Planktonic Copepods

 

Imprimer Contact Accueil Plan du site Accès restreint Retour Une du Labo Imprimer Contact Plan du site Crédits Téléchargez les Plug-Ins