Species Card of Copepod
Calanoida ( Order )
    Clausocalanoidea ( Superfamily )
        Clausocalanidae ( Family )
            Clausocalanus ( Genus )
Clausocalanus ingens  Frost & Fleminger, 1968   (F,M)
Syn.: Clausocalanus arcuicornis : De Decker, 1964 (part., p.15, 20, 30);
C. laticeps : Tanaka, 1960 (p.32, figs.F,M) data from Frost & Fleminger;
C. arcuicornis major Unterüberbacher, 1964 (part., p.19, Pl.27)
Ref.:
Frost & Fleminger, 1968 (p.39, figs.F,M, Rem.); Ramirez, 1969 (p.53, figs.F, Rem.); Björnberg & al., 1981 (p.628, figs.F,M); Bradford-Grieve, 1994 (p.114, figs.F,M, fig.101); Mazzocchi & al., 1995 (p.153, figs.F, Rem.); Bradford-Grieve & al., 1999 (p.878, 916, figs.F,M); Bucklin & al., 2003 (p.335, tab.2, figs.1, 4, Biomol)
Species Clausocalanus ingens - Plate 1 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [Pl.19, p.138-139; Pl.20, p.140-141; Pl.21, p.142-143].

Female: 1 a, habitus (right lateral view); 1 b, idem (dorsal view); 2 a, frontal region (right lateral); 2 b, rostrum ( (right lateral); 2 c, rostrum (anteroventral); 2 d, Th.4-5 (posterior part) and genital segment (right lateral); 2 z, genital segment (ventral view); 2 a-b taken from one specimen; 2 c-e from different specimens; 3 a, Th.4-5 (posterior part) and urosome with spermatophore attached (right lateral); 3 b, B2 of P2; 3 c, B2 of P3; 3 d-e, P5; 3 a taken from one specimen; 3 b-c from another specimen; 3 d-e from third and fourth specimens, respectively.


Species Clausocalanus ingens - Plate 2 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [Pl.22, p.144-145; Pl.23, p.146-147].

Male: 1 a, habitus (right lateral view); 1 b, idem (dorsal view); 1 c, frontal region (right lateral); 1 d, urosome (dorsal view); 1 e, urosome 4 (posterior part), urosome 5 and caudal rami (dorsal view); 1 a-b taken from one specimen; 1 c-e taken from different specimens; 2 a, Th.4-5 (posterior part) and urosome (left lateral view); 2 b, B2 of P2; 2 c, B2 of P3; 2 d, P5 (posterior); 2 e, P5 (left anterolateral); 2 a taken from one specimen; 2 b-c from another ; 2 d and e from third and fourth specimens, respectively.
Nota: 2nd segment of P5 less than 4.4 times as long as wide.


Species Clausocalanus ingens - Plate 3 of morphological figuresissued from : J.M. Bradford-Grieve in The Marine Fauna of New Zealand: Pelagic Calanoid Copepoda. National Institute of Water and Atmospheric Research (NIWA). New Zealand Oceanographic Institute Memoir, 102, 1994. [p.114, Fig.64].
Female: A, habitus (lateral left side); B, genital somite (lateral left side); C, rostrum (anterodorsal) view); D, basipod 2 of P3; E, P5.

Male: F, habitus (lateral left side); G, left caudal ramus (dorsal); H, basipod 2 of P3; I, P5.


Species Clausocalanus ingens - Plate 4 of morphological figuresIssued from: M.G. Mazzocchi, G. Zagami, A. Ianora, L. Guglielmo & J. Hure in Atlas of Marine Zooplankton Straits of Magellan. Copepods. L. Guglielmo & A. Ianora (Eds.), 1995. [p.154, Fig.3.26.1].
Female: A, habitus (dorsal); B, urosome (dorsal); C, habitus (lateral left side); D, urosome (lateral left side).
Nota: Proportional lengths of urosomites and furca 36:20:17:12:15 = 100. A1 almost reaching end of cephalothorax.


Species Clausocalanus ingens - Plate 5 of morphological figuresIssued from: M.G. Mazzocchi, G. Zagami, A. Ianora, L. Guglielmo & J. Hure in Atlas of Marine Zooplankton Straits of Magellan. Copepods. L. Guglielmo & A. Ianora (Eds.), 1995. [p.155, Fig.3.26.2].
Female (SEM preparation): B, urosome (dorsal); C, idem (ventral); D, P5.
Bars: B, C 0.100 mm; D 0.010 mm.


Species Clausocalanus ingens - Plate 6 of morphological figuresissued from : F.C. Ramirez in Contr. Inst. Biol. mar., Buenos Aires, 1969, 98. [p.56, Lam. IX, fig.62].
Female (from off Mar del Plata): 62, P5
Scale bar in mm: 0.02.


Species Clausocalanus ingens - Plate 7 of morphological figuresissued from : F.C. Ramirez in Contr. Inst. Biol. mar., Buenos Aires, 1969, 98. [p.50, Lam. VIII, figs.49-51].
Female (from off Mar del Plata): 49-51, basipodal segment 2 of P2-P4, respectively.
Scale bar in mm: 0.05.


Species Clausocalanus ingens - Plate 8 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.30, Table 2a].
Clausocalanus ingens Females: Measurements and ratios *.


Species Clausocalanus ingens - Plate 9 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.31, Table 2b].
Clausocalanus ingens males: Measurements and ratios *.
r = sample range; m = sample mean; n = number of specimens measured; s = sample standard deviation.


Species Clausocalanus ingens - Plate 10 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.26, Fig.9].
Ratio of length-to-width of 2nd segment of P5 male (ordinate plotted against length of 2nd segment of left P5 (abscissa) for Clausocalanus ingens.


Species Clausocalanus ingens - Plate 11 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.143, Pl.21, b, c, d-e].
Female: b, basipodite 2 of P2; c, basipodite 2 of P3; d-e, P5 (from different specimens).


Species Clausocalanus ingens - Plate 12 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.47, Pl.23, a, d, e].
Male: a, posterior part of last thoracic segment and urosome (left lateral); d, P5 (posterior); e, P5 (left anterolateral).


Species Clausocalanus ingens - Plate 13 of morphological figuresissued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, 1968, 12. [p.147, Pl.23, b, c].
Male: b, basipodite 2 of P2; c, basipodite 2 of P3.


Species Clausocalanus ingens - Plate 14 of morphological figuresIssued from : O. Tanaka in Spec. Publs. Seto mar. biol. Lab., 10, 1960 [Pl. XI, 1-8]. As Clausocalanus laticeps.
Female (from Sub-Antarctic): 1, forehead (dorsal); 2,


Species Clausocalanus ingens - Plate 15 of morphological figuresIssued from : O. Tanaka in Spec. Publs. Seto mar. biol. Lab., 10, 1960 [p.32]. As Clausocalanus laticeps.
Female: Proportional lengths of segments of A1.


Species Clausocalanus ingens - Plate 16 of morphological figuresssued from : O. Tanaka in Spec. Publs. Seto mar. biol. Lab., 10, 1960 [Pl. XI, 9-12]. As Clausocalanus laticeps.
Male (from Sub-Antarctic): 9, habitus (lateral); 10, P2; 11, distal segment of left P5; 12, right P5.


Species Clausocalanus ingens - Plate 17 of morphological figuresIssued from : O. Tanaka in Spec. Publs. Seto mar. biol. Lab., 10, 1960 [p.33]. As Clausocalanus laticeps.
Male: Proportional lengths of segments of A1.

Compl. Ref.:
Björnberg, 1973 (p.311, 385); Nyan Taw & Ritz, 1979 (p.196); De Decker, 1984 (p.315, 343: carte); Razouls & al., 2000 (p.343, tab. 3, Appendix); Ward & al., 2003 (p.121, tab.4); Lo & al., 2004 (p.89, tab.1); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical, Fig.4, 6, 7, Tabe 4); Hidalgo & al., 2012 (p.134, Table 2); Zakaria & al., 2016 (p.1, Table 1, Rem.)
NZ: 9

Distribution map of Clausocalanus ingens by geographical zones
Species Clausocalanus ingens - Distribution map 4issued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [p.40, Chart 3, a].
Occurrence of C. ingens in samples examined; closed circles represent samples examined; open circles represent samples in which adults were found; bars through open circles represent samples from which specimens were removed for measurements.
Species Clausocalanus ingens - Distribution map 5issued from : B. Frost & A. Fleminger in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 1968, 12. [p.41, Chart 3, b].
Occurrence of C. ingens in samples examined; closed circles represent samples examined; open circles represent samples in which adults were found; bars through open circles represent samples from which specimens were removed for measurements.
Loc:
sub-Antarct. (N South Georgia, SW & SE Pacif., Straits of Magellan (Central part and Pacific area), S Pacif., S Australia, Tasmania , New Zealand (N, SE North Island), China Seas (East China Sea), Taiwan (N: Mienhua Canyon), Chile, off Santiago, S Indian, South Africa (E & W), SW Atlant., S Brazil , E Medit. (W Egyptian coast in Zakaria & al., 2016 (Table 1)
N: 16
Lg.:
(30) F: 1,9-1,44; M: 1,08-0,99; (36) F: 1,84;1,67; (66) F: 1,62; M: 1,07; {F: 1,44-1,90; M: 0,99-1,08}

The mean female size is 1.694 mm (n = 5; SD = 0.1832) and the mean male size is 1.047 (n = 3; SD = 0.0493). The size ratio (male : female) is 0.64.
Rem.: epipelagic.
Sampling depth (sub-Antarct.) : 0-500 m.
For Frost & Fleminger (1968, p.41) C. ingens female is distinguished from laticeps by the length-width ratio of the caudal ramus and the form and spacing of basipodite 2 spiniform processes 2 and 3 of P3. C. ingens male differs from laticeps in the Prosome/Urosomal segment 2 length ratio, the form of basipodite 2 spiniform processes of P3, and the length-width ratio of the caudal ramus. C. ingens was confused with laticeps by Tanaka (1960), and it is likely that the two species have been confused by others.
Lo & al. (2004) seem to indicate for the first time the presence of this species in the NW Pacific (north of Taiwan), and Zakaria & al. (2016) in the East Mediterranean Sea (Egyptian coast).
Last update : 18/08/2018
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