Species Card of Copepod
Calanoida ( Order )
    Arietelloidea ( Superfamily )
        Heterorhabdidae ( Family )
            Heterorhabdus ( Genus )
Heterorhabdus prolatus  Park, 2000   (F,M)
Syn.: Heterorhabdus papilliger: Grice, 1962 (p.222, figs.M, no F), after Park (2000, p.110)
Ref.:
Park, 2000 (p.109, figs.F,M, Rem.)
Species Heterorhabdus prolatus - Plate 1 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 2000, 31. [p.230, Fig.78].
Female: a, habitus (left side); b, c, urosome (left, dorsal, respectively); d, e, forehead (left, dorsal, respectively); f, g, h, genital somite (left, ventral, right, respectively).

Nota:
- Prosome length 70 % length of body and about 2.3 times length of urosome.
- Forehead angular and produced ventrad into a small rostrum bearing 2 slender filaments. dorsally, midanterior tubercular process (fig.78 e) tapering into a pointed tip.
- Cephalosome 46 % length of prosome, with a cervical groove halfway along dorsal margin
- Posterolateral corner of prosome rounded.
- Length ratios of 4 urosomal somites and left caudal ramus 38.8 : 18.7 : 13.7 : 8.8 : 20.0 = 100.
- Left caudal ramus extending beyond posterior end of right ramus by about 1/5 its length( as measured along medial margin). Dorsal appendicular seta of left ramus almost twice as long as that of right ramus.; 4th marginal seta of left ramus much thicker than other marginal setae and longer than body. Dorsally, each caudal ramus with a relatively large pore.
- A1 reaching close to posterior end of caudal ramus.


Species Heterorhabdus prolatus - Plate 2 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 2000, 31. [p.231, Fig.79].
Female: a, left Mx2 (posterior); b, P5 (anterior).
Nota: Mx2: Falcate spine of 5th lobe with proximal 54 % of medial margin serrated with spinules; falcate spine of 6th lobe with proximal 63 % of medial margin serrated with spinules. (compare with H. spinifer fig.76 g).

Male: c, right P5 (endopod omitted), anterior; d, left P5 (anterior); e, exopod of P5 (anterior).

Nota male:
- Prosome 71 % length of body; urosome 30 % length of body.
- First 4 urosomal somites armed with relatively large spinules along posterior margins.
- Caudal rami and setation as in female.


Species Heterorhabdus prolatus - Plate 3 of morphological figuresissued from : G.D. Grice in Fish. Bull. Fish and Wildl. Ser., 1962, 61. [p.221, Pl.24, Figs.13-16]. As Heterorhabdus papilliger.
Male (from equatorial Pacific): 13, habitus (lateral); 14, Mx2; 15, P5; 16, distal end of protrusion of 2nd exopodal segment of right P5.


Species Heterorhabdus prolatus - Plate 4 of morphological figuresHeterorhabdus prolatus female:
1 - See key to species groups of Heterorhabdus: ''papilliger'' Group (p.90, 102).
2 - Midanterior tubercular process of forehead produced into an angular process (Fig.78-e) and not produced into a spiniform process. P5 with inner marginal seta on 1st endopodal segment (Fig.79-b).
3 - Laterally, genital prominence extending far shorter to posterior end of somite (Fig.78-b).


Species Heterorhabdus prolatus - Plate 5 of morphological figuresHeterorhabdus prolatus male:
1 - See key to species groups of Heterorhabdus: ''papilliger'' Group (p.90, 102).
2 - Midanterior tubercular process of forehead angular and not produced into a spiniform process. Basal inner lobe of left P5 rather poorly developed and angular distally (Fig.79-d)
3 - In right P5, medial projection of 2nd exopodal segment not terminally expanded, somewhat truncate, with nearly parallel proximal and distal sides (Fig.79-c).

NZ: 4

Distribution map of Heterorhabdus prolatus by geographical zones
Loc:
S Mozambique, W Indian, NE Indian (equatorial), Indonesia-Malaysia, China Seas (South China Sea), Pacif. (equatorial: E-W), E Pacif. tropical

Type locality: 14°46'S, 93°37'W.

For Park (2000, p.110), this species is apparently an equatorial species of the Indo-Pacific.
N: 2
Lg.:
(101) M: 2,11-1,7; (824) F: 2,14-1,78; M: 2,14-1,8; {F: 1,78-2,14; M: 1,70-2,14}
Rem.: "Papilliger" Group.
Last update : 20/05/2017
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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2021. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed October 21, 2021]

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