Species Card of Copepod
Calanoida ( Order )
    Arietelloidea ( Superfamily )
        Heterorhabdidae ( Family )
            Paraheterorhabdus ( Genus )
                Paraheterorhabdus ( Sub-Genus )
Paraheterorhabdus (Paraheterorhabdus) medianus  (Park, 1970)   (F,M)
Syn.: Heterorhabdus medianus Park, 1970 (p.477, 525, figs.F, no M); Heptner, 1972 a (p.60);
Heterochäta vipera Giesbrecht, 1889; 1892 (p.373, 383, 773, figs.M, non F);
Heterorhabdus vipera : Giesbrecht & Schmeil, 1898 (p.116, Rem. M, no F)
Ref.:
Park, 2000 (p.81, figs.F,M, Rem.)
Species Paraheterorhabdus (Paraheterorhabdus) medianus - Plate 1 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 2000, 31. [p.204, Fig.52].
Female: a, habitus (left side); b, urosome (dorsal); c, d, forehead (left, dorsal, respectively); e, f, genital somite (left, ventral, respectively); g, masticatory edge of left Md (posterior); h, masticatory edge of right Md (posterior); i, left Mx2 (posterior).


Species Paraheterorhabdus (Paraheterorhabdus) medianus - Plate 2 of morphological figuresissued from : T. Park in Bull. Scripps Inst. Oceanogr. Univ. California, San Diego, 2000, 31. [p.205, Fig.53].
Female: a, right Mxp (anterior); b, exopod of P1 (anterior); c, exopod of P5 (anterior).

Male: d, urosome (dorsal); e, P5 (anterior); f, second exopodal segment of left P5 (anterior); g, right P5 (endopod omitted), posterior; h, basipod of right P5 (anterior).


Species Paraheterorhabdus (Paraheterorhabdus) medianus - Plate 3 of morphological figuresIssued from : T. S. Park in Bull. Mar. Sc., 1970, 20 (2). [p.522, Figs.253-254]. As Heterorhabdus medianus.
Female (from Caribbean Sea & G. of Mexico): 253, habitus (lateral right side); 254, idem (dorsal).
Nota: Head and 1st thoracic segment separate, 4th and 5th fused. caudal rami asymmetrical.


Species Paraheterorhabdus (Paraheterorhabdus) medianus - Plate 4 of morphological figuresIssued from : T. S. Park in Bull. Mar. Sc., 1970, 20 (2). [p.524, Figs.255-261]. As Heterorhabdus medianus.
Female: 255, right Md (biting edge of the mandibular blade); 256, left Md (idem); 257, Mx1; 258, Mx2; 259, Mxp; 260, P1 (anterior); 261, P5 (anterior).
Nota: 2nd inner lobe and basis of Mx1, each with 1 seta; endopod with 5 setae. Penultimate lobe of Mx2 elongate and not covered by spinules.


Species Paraheterorhabdus (Paraheterorhabdus) medianus - Plate 5 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf. 20, Fig.6]. As Heterochäta vipera.
Male: 6, exopod 3 of P3.


Species Paraheterorhabdus (Paraheterorhabdus) medianus - Plate 6 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892, 19 , Atlas von 54 Tafeln. [Taf.20, Figs.32, 33]. As Heterochäta vipera.
Male: 32, left P5 (anterior view); 33, right P5 (anterior view).
Nota: Same scale.


Species Paraheterorhabdus (Paraheterorhabdus) medianus - Plate 7 of morphological figuresParaheterorhabdus (Paraheterorhabdus) medianus female:
1 - Left caudal ramus distinctly longer than right and completely fused with anal segment (Fig.52-b)
2 - Dorsally, posterolateral corners of prosome rounded.


Species Paraheterorhabdus (Paraheterorhabdus) medianus - Plate 8 of morphological figuresParaheterorhabdus (Paraheterorhabdus) medianus female:
1 - Left caudal ramus distinctly longer than right and completely fused with anal segment; setation in both rami as in female (Fig.53-d).
2 - Basis of right P5 with wide, rectangular inner lobe.
3 - Exopod of right P5 with wide bilobed distal end (Fig.53-e); 2nd lobe of Mx2 (Fig.52-i) with a long seta.

NZ: 6

Distribution map of Paraheterorhabdus (Paraheterorhabdus) medianus by geographical zones
Loc:
off W South Africa, Caribbean Sea, G. of Mexico, off Cape Verde Is., Indian (E equatorial), E Pacif. tropical (N-S), Pacif. tropical (central S-N), Salomon Is.

Type locality: Caribbean Sea and Gulf of Mexico.
N: 3
Lg.:
(46) M: 2,6; (88) F: 2,6; 2,56; (824) F: 2,88-2,48; M: 2,84-2,6; {F: 2,48-2,88; M: 2,60-2,84}
Rem.: Mesopelagic.
For Park (1970, p.527) this species shows somewhat intermediate characteristics between P. vipera (Giesbrecht, 1889) and P. robustus (Farran. 1908).
Last update : 27/01/2015
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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2021. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed October 20, 2021]

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