hyperbenthic. (220-235 m). For Fosshagen (1978, p;181) M. hepneri may be considered the most primitive member of the genus: A1 female 24; exopod of the P1 bears 1 spine on the outer margin of the 1st and 2nd segments; it is the only known species with both P5 present in the female and a 3-segmented right P5 in the male; and there is less reduction in M. hepneri than in the other species.