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Famille
Ryocalanidae Andronov, 1974
Ref.:
Andronov, 1974 a (p.1005); Razouls, 1982 (p.136); Bowman & Abele, 1982 (p.10); Park, 1986 (p.191, 194); Huys & Boxshall, 1991 (p.419); Andronov, 1991(p.133); Razouls, 1993 (p.309); Chihara & Murano, 1997 (p.898); Ohtsuka & Huys, 2001 (p.445, 461); Boxshall & Halsey, 2004 (p.15; 49; 182: Def.); Vives & Shmeleva, 2007 (p.995): Markhaseva, 2012 (p.148, Rem.); Renz & al., 2013 (p.247, 257); Renz & al., 2018 (p.30: phylogenetic relationships)
Bradford-Grieve J.M., (2002 onwards). Key to calanoid copepod families. Version 1 : 2 oct 2002. http://www.crustacea.net/crustace/calanoida/index.htm
Bradford-Grieve J.M., (2002 onwards). Key to calanoid copepod families. Version 1 : 2 oct 2002. http://www.crustacea.net/crustace/calanoida/index.htm
Rem.:
Andronov crée pour le genre monotypique R. infelix la famille des Ryocalanidae qui peut être définie par les segments 8 et 9 des A1 soudés, la structure et la segmentation des pattes comme dans la famille des Spinocalanidae, l'articulation de l'antennule géniculée entre les segments 22 et 23 (et non entre les segments 18 et 19 comme dans la majorité des autres genres).
2 G.: Ryocalanus, Yrocalanus.
2 G.: Ryocalanus, Yrocalanus.
Figures
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Planche 1
Issued from : J. Renz, E.L. Markhaseva & K. Schulz inZoosyst. Evol., 2013, 89 (2). [p.248, Fig.1]
Distribution of Ryocalanidae species in the world oceans.
1 - Ryocalanus infelix; 2 and 2a - Yrocalanus admirabilis; 3, 4, 5 - Yrocalanus asymmetricus, Yrocalanus bicornis, Ryocalanus bowmani; 6 - Ryocalanus spinifrons; 7 - Yrocalanus antarcticus; 8 - Ryocalanus brazilianus.
Issued from : J. Renz, E.L. Markhaseva & K. Schulz inZoosyst. Evol., 2013, 89 (2). [p.248, Fig.1]
Distribution of Ryocalanidae species in the world oceans.
1 - Ryocalanus infelix; 2 and 2a - Yrocalanus admirabilis; 3, 4, 5 - Yrocalanus asymmetricus, Yrocalanus bicornis, Ryocalanus bowmani; 6 - Ryocalanus spinifrons; 7 - Yrocalanus antarcticus; 8 - Ryocalanus brazilianus.
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Planche 2
Issued from : G.A. Boxshall & S.H. Halsey in An Introduction to Copepod Diversity. The Ray Society, 2004, No 166, Part. I. [p.182].
Armature formula of swimming legs P1 to P4.
- Female P5 absent. Male P5 asymmetrical; with coxae and intercoxal sclerites fused to form posteriorly bilobed transverse plate; right leg uniramous, comprising basis and 1-segmented exopod, both unarmed; left leg twice as long as right, biramous with 3-segmented exopod and minute, knob-like endopod; exopodal segments lacking spines and setae.
Nota: Proximal outer spine of 3rd exopodal segment of P1 often lost, as in Ryocalanus bicornis Markhaseva & Ferrari, giving formula I, I, 4. Inner margin of coxa of P4 ornamented with pair of large spinules (possibly spines) near base of inner seta.
Issued from : G.A. Boxshall & S.H. Halsey in An Introduction to Copepod Diversity. The Ray Society, 2004, No 166, Part. I. [p.182].
Armature formula of swimming legs P1 to P4.
- Female P5 absent. Male P5 asymmetrical; with coxae and intercoxal sclerites fused to form posteriorly bilobed transverse plate; right leg uniramous, comprising basis and 1-segmented exopod, both unarmed; left leg twice as long as right, biramous with 3-segmented exopod and minute, knob-like endopod; exopodal segments lacking spines and setae.
Nota: Proximal outer spine of 3rd exopodal segment of P1 often lost, as in Ryocalanus bicornis Markhaseva & Ferrari, giving formula I, I, 4. Inner margin of coxa of P4 ornamented with pair of large spinules (possibly spines) near base of inner seta.
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Planche 3
Issued from : G.A. Boxshall & S.H. Halsey in An Introduction to Copepod Diversity. The Ray Society, 2004, No 166, Part. I. [p.183, Fig.43].
Ryocalanidae: A, Ryocalanus admirabilis, habitus female; B, habitus male; C, distal tip of male A1. D,Ryocalanus spinifrons, female P1. E, Ryocalanus bowmani, male P5. F, Ryocalanus bicor,nis, coxa of female P4. [Andronov, 1992: A-C; Shimode, Toda & Kikuchi, 2000: D. Markhaseva & Ferrari, 1996: E-F]
Issued from : G.A. Boxshall & S.H. Halsey in An Introduction to Copepod Diversity. The Ray Society, 2004, No 166, Part. I. [p.183, Fig.43].
Ryocalanidae: A, Ryocalanus admirabilis, habitus female; B, habitus male; C, distal tip of male A1. D,Ryocalanus spinifrons, female P1. E, Ryocalanus bowmani, male P5. F, Ryocalanus bicor,nis, coxa of female P4. [Andronov, 1992: A-C; Shimode, Toda & Kikuchi, 2000: D. Markhaseva & Ferrari, 1996: E-F]
Genre
Ref.:
Tanaka, 1956 b (p.1); 1956 c (p.404); Razouls, 1982 (p.136); Razouls, 1993 (p.309); Bradford-Grieve, 1994 (p.93, Def.); Chihara & Murano, 1997 (p.898); Mauchline, 1998 (p.78: F; p.80: M); Shimode & al., 2000 (p.133: clé spp.); Bradford-Grieve, 2004 (p.287); Boxshall & Halsey, 2004 (p.184: Def., spp. Key); Vives & Shmeleva, 2007 (p.995); Markhaseva, 2012 (p.149, Rem.: 2 groups); Renz & al., 2013 (p.255, Rem.); Renéz & al., 2018 (p.11-12, Rem.)
Rem.:
Brodsky (1980: comm. pers.) indique que Ryocalanus devrait être orthographié comme Riocalanus. Type: Ryocalanus infelix Tanaka,1956 .
Après redéfinition du genre Ryocalanus le nombre des espèces est de 5.
Après redéfinition du genre Ryocalanus le nombre des espèces est de 5.
Genre
Ref.:
Renz & al., 2013 (p.257, Def., Rem.); Renz & al., 2018 (p.23: Rem.)
Rem.:
Type: Ryocalanus admirabilis Andronov, 1992. Total: 5 spp.