Liste des espèces et variétés par famille
Diaixidae Sars, 1902 ( Clausocalanoidea )
(1) Anawekia Othman & Greenwood, 1994
Rem.: Type: Anawekia spinosa Othman & Greenwood, 1994. Total: 3 spp.

[1] Anawekia bilobata  Othman & Greenwood, 1994   (F,M)    [Figs]

[2] Anawekia robusta  Othman & Greenwood, 1994   (F,M)    [Figs]

[3] Anawekia spinosa  Othman & Greenwood, 1994   (F,M)    [Figs]
(2) Byrathis Markhaseva & Ferrari, 2005
Rem.: Type: Xanthocalanus macrocephalon. Grice & Hulsemann,1970. Total: 7 spp. + 1 indet.

[1] Byrathis arnei  Schulz, 2006   (F, M)    [Figs]

[2] Byrathis divae  Markhaseva & Renz, 2011   (F)    [Figs]

[3] Byrathis laptevorum  (Markhaseva, 1998)   (F)    [Figs]

[4] Byrathis laurenae  Markhaseva & Ferrari, 2005       [Figs]

[5] Byrathis macrocephalon  (Grice & Hulsemann, 1970)   (F)    [Figs]

[6] Byrathis penicillatus  Markhaseva & Renz, 2011   (F)    [Figs]

[7] Byrathis volcani  Markhaseva & Ferrari, 2005   (F)    [Figs]

[8] Byrathis sp.  Markhaseva & Ferrari, 2005   (F)    [Figs]
(3) Diaixis Sars, 1902
Rem.: Type: Scolecithrix hibernica A. Scott, 1896.
formes plus ou moins hyperbenthiques. 8 spp.

[1] Diaixis asymmetrica  Grice & Hulsemann, 1970   (F,M)    [Figs]

[2] Diaixis centrura  Connell, 1981   (F,M)    [Figs]

Diaixis durani    Corral Estrada, 1972   
Ref.: Corral Estrada, 1972 a (p.51, Descr.M, figs.M); Andronov, 1979 (p.95, Descr.F, figs.F,M); Bradford & al., 1983 (p.123)
Loc: Espagne NW (Ria de Arosa, Galicie), Mauritanie, Médit. (Malaga)
Lg.: (379) F: 0,86-0,74; M: 0,74-0,68; (380) M: 0,76; {F: 0,74-0,86; M: 0,68-0,76}
Rem.: Othman & Greenwood, 1994 (p.988) estiment que la femelle décrite par Andronov n'appartient pas à la même espèce. Cf.: Diaixix pygmaea

[3] Diaixis gambiensis  Andronov, 1979   (F,M)    [Figs]

[4] Diaixis helenae  Andronov, 1979   (F,M)    [Figs]

[5] Diaixis hibernica  (A. Scott, 1896)   (F,M)    [Figs]

Diaixis hibernica hibernica    A. Scott, 1896   (F,M)
Ref.: Andronov, 1979 (p.97-99, figs.F); Bradford-Grieve & al., 1999 (p.881, 934, figs.F)
Loc: Afr.W, Canaries

Diaixis longiremis    ?   
Ref.: in Kovalev & Shmeleva, 1982 (p.84)
Loc: Adriatique
Rem.: Description originale inconnue.

Diaixis minor    ?   
Ref.: in Kovalev & Shmeleva, 1982 (p.84)
Loc: Médit. W
Rem.: Description originale inconnue.

[6] Diaixis pygmaea  (T. Scott, 1896)   (F,M)    [Figs]

[7] Diaixis tridentata  Andronov, 1974   (F)    [Figs]

[8] Diaixis trunovi  Andronov, 1979   (F,M)    [Figs]

Diaixis sp.    Duran, 1963   (M)
Ref.: Duran, 1963 (p.17, figs.M)
Loc: Medit. W (Baléares: Formentera)
Rem.: ? Cf. Diaixis durani
(4) Procenognatha Markhaseva & Schulz, 2010
Rem.: Type: Procenognatha semisensata. Total: 1 sp.

[1] Procenognatha semisensata  Markhaseva & Schulz, 2010   (F,M)    [Figs]
(5) Pseudophaenna Sars, 1902
Rem.: Type: Pseudophaenna typica Sars, 1903. Total: 1 sp.
Ce genre, placé parmi les Tharybidae, est transféré dans cette famille par Boxshall & Halsey (2004, p.96, 210).

[1] Pseudophaenna typica  Sars, 1902   (F,M)    [Figs]
(6) Ranthaxus Markhaseva & Schulz, 2010
Rem.: Total: 1 sp.

[1] Ranthaxus vermiformis  Markhaseva & Schulz, 2010   (F)    [Figs]
(7) Sensiava Markhaseva & Schulz, 2006
Rem.: Type species: Sensiava longiseta. Total: 3 spp. + 1 indet.
Inclus provisoirement par les auteurs dans la famille des Diaixidae.

[1] Sensiava longiseta  Markheseva & Schulz, 2006   (M, ?F)    [Figs]

[2] Sensiava peculiaris  Markheseva, 2014   (F)    [Figs]

[3] Sensiava secunda  Markheseva, 2014   (F, ?M)    [Figs]

[4] Sensiava sp.  Markheseva, 2014   (F)    [Figs]
(8) Thoxancalanus Markhaseva, Laakmann & Renz, 2013
Rem.: Type: Thoxancalanus spinatus. Total: 1 sp.

[1] Thoxancalanus spinatus  Markhaseva, Laakmann & Renz, 2013 (2014)   (F)    [Figs]
(9) Vensiasa Markhaseva, 2015
Rem.: Type species: Vensiasa incerta Markhaseva, 2015.

Diagnosis from Markhaseva (2015, p.183):
Female:
- Cehalosome and pediger 1, pedigers 4 and 5 incompletely separate.
- posterior corners, in dorsal view, triangular ; as rounded lobes, in lateral view.
- Rostrum as a poorly developed plate with 2 filaments.-
A1 24 free segments.
- Coxa of A2 with 1 seta; basis with 2 setae; endopodal segment 1 with 1 seta, exopod setal formula 0,0-0-1, 1, 1, 1, 1, 1, 3.
- Gnathobase of Md with 7-8 teeth; exopod 5-segmented with 1,1,1,1, and 2 setae; endopod segment 1 with 3 setae, 2nd endopod with 9 setae; basis with 3 setae.
- Mx1 praecoxal arthrite with 9 terminal spines, 2 posterior setae and 1 anterior seta; coxal endite with 3 setae, coxal epipodite with 8-9 setae; proximal basal endite with 3 setae, distal basal endite with 2-3 setae; endopod with 5 setae, exopod with 8 setae.
- Mx2 praecoxal endite bearing 5 setae and attenuation, coxal and basal endites with 3 setae each; enditic-like lobe of proximal endopodal segment with 4 setae; 2 setae of distal basal endite and 2 setae of enditic-like lobe equal in length and curved distally; endopod with 6 worm-like (w) and 2 brush-like (br) sensory setae, 3 terminal worm-like setae long, 3 proximal worm-like setae short and very thin and 2 brush-like setae: 1 thick with brush well developed and 1 thin with poorly developed brush.
- Mxp syncoxa with 1, 2 and 3 setae on praecoxal endites.
- P1 to P4 typical clausocalanoidean setation and segmentation.
- P5 uniramous, symmetrical, 3-segmentesd, with 4 setal elements terminally.

Male:
- Rostrum as in female.
- Cephalosome and pediger 1, pedigers 4 and 5 separate.
- Prosome posterior corners as short rounded lobes.
- Right A1 23-segmented; left 24 free segments; ancestral segments X-XI fused.
- Oral appendages sexually dimorphic and moderately reduced compared to females.
- Setatio of A2 asin female.
- Number of setae on Md basis and endopod segment 2 as in female, endopod segment 1 with 2 setae.
- Armament of Mx1 differs from female in: praecoxal arthrite with 1 posterior seta; coxal epipodite with 7 setae; proximal basal endite with 2 setae, and exopod with 7 setae.
- Mx2 praecoxal endite with 4 setae, other lobes with the same setal numbers as in female.
- Mxp setation of syncoxa as in female, distal endopod setae more than 1.6 times longer than basis.
- P1 to P4 as in female.
- P5 nearly as long as urosome; protopods asymmetrical, right leg uniramous, 3-segmented; left leg biramous, endopod longer than 3-segmented exopod.

The new genus Vensiasa is defined among ''Bradfordians'' by the morphology of the sensory setae on the maxillary (Mx2) endopod (6w + 2br), their structure is an apomorphy for the genus: the 3 proximal worm-like setae are remarkably short and thin, 1 brush-like seta is very thin with poorly developed brush and 1 brush-like seta is very thick, brush well developed. Mx1 endopod with 5 setae (vs 7-12 in other diaixids) and male's P5 protopods asymmetrical; left P5 endopod longer than 3-segmented exopod (vs in other diaixids if protopods asymmetrical, then left endopod rudimentary or absent).
The new genus appears to be more closely related to the genus Sensiava.

[1] Vensiasa incerta  Markhaseva, 2015   (F)    [Figs]

[2] Vensiasa sp.  Markhaseva, 2015   (F,M)    [Figs]
(10) Xancithrix Markhaseva, 2012
Rem.: Type: Xancithrix ohmani. Total: 1 sp.

[1] Xancithrix ohmani  Markhaseva, 2012   (F,M)    [Figs]

 Toute utilisation de ce site pour une publication sera mentionnée avec la référence suivante :

Razouls C., Desreumaux N., Kouwenberg J. et de Bovée F., 2005-2024. - Biodiversité des Copépodes planctoniques marins (morphologie, répartition géographique et données biologiques). Sorbonne Université, CNRS. Disponible sur http://copepodes.obs-banyuls.fr [Accédé le 28 mars 2024]

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