Fiche d'espèce de Copépode
Calanoida ( Ordre )
    Diaptomoidea ( Superfamille )
        Tortanidae ( Famille )
            Tortanus ( Genre )
                Tortanus ( Sous-Genre )
Tortanus (Tortanus) barbatus  (Brady, 1883)   (F,M)
Syn.: Corynura barbata Brady, 1883 (p.71, Descr.F, figs.F); Giesbrecht, 1892 (p.772);
Corynura denticulata Giesbrecht, 1889; Maiphae & Sa-ardrit, 2011 (p.641, Table 2, 3, Rem.);
Tortanus denticulatus Giesbrecht & Schmeil, 1898 (p.158); Shih & Young, 1995 (p.74); Tortanus sp. M Dakin & Colefax, 1940 [After Mulyadi, 2004, p.166]
Ref.:
Giesbrecht & Schmeil, 1898 (p.158, Rem. F); A. Scott, 1909 (p.189, figs.F, Rem.); Sewell, 1912 (p.354, 377: Rem.); 1932 (p.399, Rem.); 1933 (p.29, Rem.); Früchtl, 1923 (p.456, Descr.M, fig.M, Rem.); Früchtl, 1924 b (p.15, 61, figs.F,M, Rem.); Steuer, 1926 a (p.61, figs.F,M, Rem., carte); Nicholls, 1944 (p.14, Rem.); C.B. Wilson, 1950 (p.345, fig.F); Kasturirangan, 1963 (p.66, figs.F); Goswami & Goswami, 1974 (p.109, fig. caryotypes); Greenwood, 1978 (p.18, figs.F,M, Rem.); Hulsemann, 1988 (p.656, Rem.); Ohtsuka, 1992 a (p.265); Kimmerer & al., 1985 (p.426); Bradford-Grieve, 1999 b (p.229, figs.F,M, Rem., figs.187, 195); Conway & al., 2003 (p.140, figs.F,M, Rem.); Mulyadi, 2004 (p.165, figs.F,M, Rem.); Phukham, 2008 (p.108, figs.F); Al-Yamani & al., 2011 (p.90, figs.F,M).
Espèce Tortanus (Tortanus) barbatus - Planche 1 de figures morphologiquesissued from : J.G. Greenwood in Proc. R. Soc. Qd, 1978, 89. [p.19, Fig.9].
Female (from Moreton Bay, E Australia): a, postrior metasome and urosome (dorsal); b, P5; c, aberrant left P5 with bifid denticle; d, posterior metasome and urosome with broad-caudal rami form (other specimen); e, P5 of same; f, posterior metasome and urosome of broad-caudal rami form (lateral left side).

Male: g, urosome (dorsal); h, P5 (anterior view); i, P5 (posterior view); j, thoracis segments 4 - 5 and urosome (lateral right side).Nota: Variability in the nature of the comb of denticles on the inner margin of the female left P5, and of lateral setae on the caudal rami.


Espèce Tortanus (Tortanus) barbatus - Planche 2 de figures morphologiquesissued from : A. Scott in Siboga-Expedition, 1909, XIX a. [Plate LV, Figs.16-18].
Female (from Patany Bay: Siam): 16, habitus (dorsal); 17, A1; 18, P5.


Espèce Tortanus (Tortanus) barbatus - Planche 3 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.31 , Figs.6, 13 ]. As Corynura denticulata.
Female: 6, A2 (anterior view); 13, P1 (posterior view).


Espèce Tortanus (Tortanus) barbatus - Planche 4 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.31 , Fig.20 ]. As Corynura denticulata.
Female: 20, P5 (posterior view).


Espèce Tortanus (Tortanus) barbatus - Planche 5 de figures morphologiquesissued from : Y. Al-Yamani, V. Skryabin, A. Gubanova, S. khvorov & I. Prusova in Marine Zooplankton Parctical Guide for the Northwestern Arabian Gulf, 2, 2011 [p.91, Fig.227 c].
Female (from Kuwait): c, P5.


Espèce Tortanus (Tortanus) barbatus - Planche 6 de figures morphologiquesissued from : Y. Al-Yamani, V. Skryabin, A. Gubanova, S. khvorov & I. Prusova in Marine Zooplankton Practical Guide for the Northwestern Arabian Gulf, 2, 2011 [p.91, Fig.228 c].
Male (from Kuwait): c, P5.


Espèce Tortanus (Tortanus) barbatus - Planche 7 de figures morphologiquesissued from : Y. Al-Yamani, V. Skryabin, A. Gubanova, S. khvorov & I. Prusova in Marine Zooplankton Practical Guide for the Northwestern Arabian Gulf, 2, 2011 [p.91, Fig.228 c].
Male (from Kuwait): c, P5.


Espèce Tortanus (Tortanus) barbatus - Planche 8 de figures morphologiquesissued from : N. Phukham in Species diversity of calanoid copepods in Thai waters, Andaman Sea (Master of Science, Univ. Bangkok). 2008. [p.191, Fig.65].
Female (from W Malay Peninsula): a, c, habitus (dorsal and lateral, respectively); b, P5.

Body length after the drawing: F = 1.430 mm.


Espèce Tortanus (Tortanus) barbatus - Planche 9 de figures morphologiquesissued from : Mulyadi in Published by Res. Center Biol., Indonesia Inst. Sci. Bogor, 2004. [p.165, Fig.91].
Female (from Java): a, habitus (dorsal); b, P5.
Nota: P5 asymmetrical; right leg 2-segmented, shorter than left one, exopodal segment (distal) short and naked; left leg long and stout, inner margin armed with a tuft of 5 curved lamelliform teeth at middle part.

Male: c, habitus (dorsal); d, P5.
Nota : A1 18-segmented, right side geniculate, anterior margins of segments 8 and 11 with row of small teeth, segments 13 to 15 with row of fine teeth, knee-joint between segments 14 and 15. P5: right leg 2-segmented, basal segment wider and triangular, provided with 2 finger-like process beset with nodulated protuberances and 1 seta; terminal segment strongly curved and arches over the first, bearing 4 small inner setae. Left leg with 1 basal and 2 exopodal segments, exopodal segment 1 with 1 outer seta; exopodal segment 2 (distal) with 2 small spines (1 near distal end and the other slightly posterior to it), 2 inner setae, and several distinct spinules near apex arranged in 7 or 8 semi-circular rows.

Ref. compl.:
Sewell, 1948 (p.324); Kott, 1957 (p.6, 15); Kimmerer & McKinnon, 1985 (p.149); Madhupratap & Haridas, 1986 (p.105, tab.1); Othman & al., 1990 (p.561, 564, Table 1); Gajbhiye & al., 1991 (p.189); Shih & Young, 1995 (p.74); Ramaiah & al., 1996 (p.3); Ramaiah & Nair, 1997 (tab.1); Mauchline, 1998 (tab.8); Greenwood & al., 2002 (p.17, Table 2); Rakhesh & al., 2008 (p.154, Table 5: abundance vs stations); McKinnon & al., 2008 (p.844: Tab.1, p.848: Tab. IV); Cornils & al., 2010 (p.2076, Table 3); Cornils & al., 2010 (p.2076, Table 3); Maiphae & Sa-ardrit, 2011 (p.641, Table 2, 3, Rem., as T. barbotus); Chew & Chong, 2011 (p.127, Table 3, abundance vs location); Johan & al., 2012 (p.647, Table 1, fig.2, salinity range); Varadharajan & Soundarapandian, 2013 (p.2: as Tartanus barbatus occurrence vs stations); Rakhesh & al., 2013 (p.7, Table 1, abundance vs stations); Jagadeesan & al., 2013 (p.27, Table 3, 5, fig.11, seasonal variation); Anjusha & al., 2013 (p.40, Table 3, abundance & feeding behavior)
NZ: 5

Carte de distribution de Tortanus (Tortanus) barbatus par zones géographiques
Loc:
Red Sea, Arabian Gulf (Kuwait), Indian, Madagascar, India (Bombay, Burhabalanga estuary, Gulf of Mannar, Palk Bay, Godavari region, Kakinada Bay, Pointcalimere), Andaman Sea, Perai River estuary (Penang), W Malay Peninsula, Straits of Malacca (Sangga estuary, upper estuary to offshore), G. of Thailand, Indonesia-Malaysia, Java (Cilacap Bay, off Labuan, off Surabaya), SW Celebes, Aru (Philippines), Viet-Nam (Cauda Bay), China Seas (East China Sea, South China Sea), E & W Australia (S & E, Brisbane River estuary, Melbourne, SE, Shark Bay, North West Cape, G. of Carpentaria)
N: 42
Lg.:
(5) F: 1,6; (81) F: 1,43; (334) F: 1,43-1,36; (625) M: 1,12-1,05; (638) F: 1,44-1,32; (991) F: 1,25-2,1; M: 1,05-1,12; (1085) F: 1,15-1,5; M: 0,9-1,2; (1122) F: 1,59; M: 1,0; {F: 1,15-2,10; M: 0,90-1,20}
Rem.: néritique, épipélagique
Voir aussi les remarques en anglais
Dernière mise à jour : 18/05/2018

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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2020. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Sorbonne Université, CNRS. Disponible sur http://copepodes.obs-banyuls.fr [Accédé le 28 septembre 2020]

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