Fiche d'espèce de Copépode
Calanoida ( Ordre )
    Clausocalanoidea ( Superfamille )
        Aetideidae ( Famille )
            Gaetanus ( Genre )
Gaetanus pileatus  Farran, 1903   (F,M)
Syn.: Gaetanus Caudani Canu, 1896 (? p.422, juv.M); Wolfenden, 1904 (p.132, figs.F, Rem.); 1905 a (p.8, fig.F); Pearson, 1906 (p.13, Rem.); A. Scott, 1909 (p.46, figs.F, Rem.); Giesbrecht & Schmeil, 1898 (p.33: Rem. juv. M); Pearson, 1906 (p.13); A. Scott, 1909 (p.46, figs.); Lysholm & Nordgaard, 1921 (p.14);
Gaetanus unicornis Esterly, 1906 a (p.57, figs.F); Wilson, 1936 c (p.90); Brodsky, 1950 (1967) (p.167, figs.F); Lee & al., 1971 (p.1150);
Gaetanus clarus Esterly, 1906 a (p.57, figs.M); Brodsky, 1950 (1967) (p.166, figs.M);
Gaetanus recticornis Wolfenden, 1911 (p.228, fig.F); C.B. Wilson, 1950 (p.233, figs.F);
Gaetanus kruppii : Sewell, 1947 (p.62, Rem.M, figs.M, no F)
Ref.:
Farran, 1903 (p.16, Descr.F, figs.F); 1908 b (p.35, Rem.); Wolfenden, 1911 (p.229, figs.F, Rem.); With, 1915 (p.104, figs.F); Sars, 1925 (p.56, figs.F, M); Farran, 1926 (p.250); Sewell, 1929 (p.103, figs.juv., Rem.); Rose, 1933 a (p.100, figs.F,M); Farran, 1936 a (p.89); Jespersen, 1940 (p.20); Sewell, 1947 (p.56, 57, 59); Vervoort, 1952 e (p.3, figs.F,M); Tanaka, 1957 b (p.175, figs.F); Vervoort, 1963 b (p.132, Rem.); Paiva, 1963 (p.33); Owre & Foyo, 1967 (p.45, figs.F,M, Rem.); Tanaka & Omori, 1970 (p.141); Park, 1975 a (p.10, 13, 20, figs.F,M, Rem. p.21-23); 1978 (p.139, figs.F); Bradford & Jillett, 1980 (p.55, figs.F,M); Gardner & Szabo, 1982 (p.230, figs.F,M); Baessa de Aguiar, 1987 (p.44, figs.F, Rem.F); Markhaseva, 1996 (p.211, figs.F, ?M); Bradford-Grieve & al., 1999 (p.879, 922, figs.F); Boxshall & Halsey, 2004 (p.59: fig.F); Vives & Shmeleva, 2007 (p.583, figs.F,M, Rem.)
Espèce Gaetanus pileatus - Planche 1 de figures morphologiquesissued from : Tanaka O. in Publ. Seto Mar. Biol. Lab., 1957, 6 (2). [Fig.43, p.176].
Female (from Sagami): a, habitus (dorsal aspect); b, head (lateral aspect); c, last thoracic segment and urosome (lateral aspect); d, third inner lobe with 3 setae and a conical process of Mx1; e, Mxp.; f, first basal joint of P4, with 16 lamellous bristles on the inner margin.

Nota Female:
- Cephalic spine has a characteristic shape.
- Last thoracic segment with lateral spines divergent, and extend to the middle of genital segment.
- Abdomen 4-segmented ; segments and caudal rami proportional lengths 44 : 17 : 14 : 9 : 16 = 100.
- Genital segment as long as wide, showing a sudden change in the continuity of the outline at the proximal 2/5 of the dorsal surface.
- Caudal rami as long as wide.
- A2 exopod about as long as endopod..
- Mx1 with 7 long and 2 short setae on outer lobe; 3rd inner lobe with 3 setae and a conical process (at apex fig.d).
- Mxp with 1st basal segment about as long as the 2nd; 1st segment with a lamellous process with a rounded tip on posterior margin.
- P1 exopod 2-segmented with no articulation between 1st and 2nd segments.
- P2 exopod 3-segmented and endopod 2-segmented; the number of teeth on terminal spine of P2 to P4 with 20, 20, and 23 respectively.
- P4 with coxa bearing about 16 lamellous bristles on the inner margin


Espèce Gaetanus pileatus - Planche 2 de figures morphologiquesissued from : T. Park in Antarctic Res. Ser. Washington, 1978, 27. [p.140, Fig.23].
Female: A, habitus (dorsal); B, forehead (lateral); C, posterior part of metasome and urosome (right lateral side); D, A2; E, Md; F, Mx1; G, Mx2.


Espèce Gaetanus pileatus - Planche 3 de figures morphologiquesissued from : T. Park in Antarctic Res. Ser. Washington, 1978, 27. [p.141, Fig.24].
Female: A, Mxp; B, P1; C, P2; D, P3; E, medial part of coxa of P4 (posterior). P1-3: legs (anterior).


Espèce Gaetanus pileatus - Planche 4 de figures morphologiquesissued from : E.L. Markhaseva in Proc. Zool. Inst. RAN, St. Petersburg, 1996, 268. [p.213, Fig.168].
Female (from SE Pacif.). Ce: forehead (dorsal and lateral, respectively); CP4: coxopod of P4 (partial).


Espèce Gaetanus pileatus - Planche 5 de figures morphologiquesissued from : E.L. Markhaseva in Proc. Zool. Inst. RAN, St. Petersburg, 1996, 268. [p.214, Fig.169 ].
Male (from Indian: off NW Cocos island)


Espèce Gaetanus pileatus - Planche 6 de figures morphologiquesissued from : T. Park in Bull. Mar. Sci., 1975, 25 (1). [p.21, Fig.6].
Female: A, B, forehead, lateral (showing a variability of the cephalic process).
Male: C, forehead (lateral); D, posterior part of metasome and urosome (left side, lateral); E, figth legs (posterior); F, last two exopodal segments of left fifth leg (medial).


Espèce Gaetanus pileatus - Planche 7 de figures morphologiquesissued from : J.M. Bradford & J.B. Jillett in Mem. N.Z. Oceanogr. Inst., 86, 1980. [p.57, Fig.37].
Female: A, habitus (lateral left side); B, idem (dorsal); C, basipod 1 of Mxp; D, exopod 1 of P1; E, inner edge of basipod 1 of P4.

Male: F, habitus (lateral left side); G, idem (dorsal); H, P5.
Nota: P5 figured by Sars (1925, plate 17, fig.16) is identical to that found on a specimen of G. latifrons and not like P5 on the present male. We suggest there has been some confusion between the figures on plate 17.


Espèce Gaetanus pileatus - Planche 8 de figures morphologiquesIssued from: G.O. Sars in Résult. Camp. Scient. Prince Albert I, 69, pls.1-127 (1924). [Pl.XVII, figs. 3-6].
Female: 3, habitus (dorsal); 4, idem (lateral left side).

Male: 5, habitus (lateral left side); 6, P5.


Espèce Gaetanus pileatus - Planche 9 de figures morphologiquesissued from : R.B.S. Sewell in The John Murray Expedition, 1933-34, Scientific Reports, VIII (1), 1947. [p.63, Fig.10, A, C-F]. As Gaetanus kruppi.
Male (from Arabian Sea): A, A1; C, A2; D, Mx1; E, Mx2; F, P5.
Nota: Proportional lengths of cephalothorax and abdomen as 75 to 25. The proportional lengths of the various segments (cephalon to caudal rami) as 540:71:65:75:42:74:48:42:10:32 = 1000. Head and 1st thoracic segment fused, 4th and 5th fused.


Espèce Gaetanus pileatus - Planche 10 de figures morphologiquesissued from : A. Scott in Siboga-Expedition, 1909, XIX a. [Plate VIII, Figs.9-15]. As Gaetanus caudani.
Female (from Indonesia-Malaysia): 9, habitus (dorsal); 10, forehead (lateral); 11, last thoracic and genital segments (left side); 12, rostrum; 13, Mxp; 14, P1; 15, P4 (basal portion only).
Nota: The lamella on the basal segment of Mxp is narrowed at the apex and deeply excavated. The posterior margin of the first three abdominal segments is fringed with fine spines.


Espèce Gaetanus pileatus - Planche 11 de figures morphologiquesissued from : G.P. Farran in Scient. Invest. Fish. Brch, Ireland, 1901 (2), Appendix 7, 1903. [Plate XVII, Figs.3-8].
Female (from off Cleggan, Irelandl): 3, A2; 4, Md (cutting edge); 5, Mx2; 6, Mx1 (lower lobes omitted); 7, Mxp; 8, P1 (anterior); 8, P1 (anterior).


Espèce Gaetanus pileatus - Planche 12 de figures morphologiquesissued from : R.N. Wolfenden in J. mar. biol. Ass. U.K, 1904, VII (N.S.). [Pl.IX, Fig.20]. As Gaetanus caudani.
Female (from NE Atlantic): 20, habitus (dorsal).


Espèce Gaetanus pileatus - Planche 13 de figures morphologiquesissued from : R.N. Wolfenden in J. mar. biol. Ass. U.K, 1904, VII (N.S.). [Pl.IX, Figs.21, 22]. As Gaetanus caudani.
Female (from NE Atlantic): 21, basipod of P4; 22, P1.


Espèce Gaetanus pileatus - Planche 14 de figures morphologiquesissued from : R.N. Wolfenden in Die Marinen Copepoden der Deutschen Südpolar-Expedition 1901-1903, 1911. [p.228, Fig.16].As Gaetanus recticornis.
Female (from S Atlantic: off Trinidad Island): a, habitus (lateral, left side); b, posterior part cephalothorax and urosome; c, basal segment of Mxp.


Espèce Gaetanus pileatus - Planche 15 de figures morphologiquesissued from : R.N. Wolfenden in Die Marinen Copepoden der Deutschen Südpolar-Expedition 1901-1903, 1911. [p.230, Fig.17]. As Gaetanus recticornis
Female: a, habitus (lateral, right side); b, Mxp; c, basipod of P4.


Espèce Gaetanus pileatus - Planche 16 de figures morphologiquesissued from : R.N. Wolfenden in Die Marinen Copepoden der Deutschen Südpolar-Expedition 1901-1903, 1911. [Pl.XXVI, Fig.13]. As Gaetanus recticornis
Female: 13, P1.


Espèce Gaetanus pileatus - Planche 17 de figures morphologiquesissued from : R.N. Wolfenden in Die Marinen Copepoden der Deutschen Südpolar-Expedition 1901-1903, 1911. [Pl.XXVII, Figs.1-2].
Female:1, habitus (dorsal); 2, P1.


Espèce Gaetanus pileatus - Planche 18 de figures morphologiquesissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1906, 3 (5). [Pl.9, Fig.5]. As Gaetanus clarus.
Male (from San Diego, California): 5, forehead (lateral).


Espèce Gaetanus pileatus - Planche 19 de figures morphologiquesissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1906, 3 (5). [Pl.14, Fig.90]. As Gaetanus clarus.
Male: 90, P5 (right foot at left of figure).


Espèce Gaetanus pileatus - Planche 20 de figures morphologiquesissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1906, 3 (5). [Pl.9, Fig.3]. As Gaetanus unicornis.
Female (from San Diego, California): 3, forehead (lateral).

Nota: A1 24-segmentedreacing beyond the caudal rami by at least the last 4 segments.


Espèce Gaetanus pileatus - Planche 21 de figures morphologiquesissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1906, 3 (5). [Pl.12, Fig.54]. As Gaetanus unicornis.
Female: 54, 1st basal of Mxp.
The lamella on the 1st basal of Mxp is present, but does not have the rod-like extension of G. brevicornis or G. miles.
Nota
Nota: Exopod of P1 distinctly 2-segmented. 1st basal of P4 with tube-like bristles.


Espèce Gaetanus pileatus - Planche 22 de figures morphologiquesissued from : C.O. Esterly in Univ. Calif. Publs Zool., 1906, 3 (5). [Pl.13, Fig.76]. As Gaetanus unicornis.
Female: distal portion of inner margin of 1st basal of P4.


Espèce Gaetanus pileatus - Planche 23 de figures morphologiquesissued from : C.B. Wilson in Bull. U.S. natn. Mus., 1950, 100, 14 (4). [Pl. 23, Figs.340-341]. As Gaetanus recticornis.
Female (from Pacific): 340, habitus (dorsal); 341, forehead (lateral).


Espèce Gaetanus pileatus - Planche 24 de figures morphologiquesissued from : R.N. Wolfenden in Plankton Studies Part I. Copepoda. 1905 [Pl.III, 2]. As Gaetanus caudani
Female: 2, habitus (lateral).


Espèce Gaetanus pileatus - Planche 25 de figures morphologiquesGaetanus pileatus female:
1 - Anterior part of cephalon with frontal spine.
2 - Exopodal segment 1 of A2 without seta; exopodal segment 2 with 1 seta (near distal margin). 2nd internal lobe of Mx1 with 4 setae. Exopod of P1 with 2 external spines.
3 - Mxp protopodite with lateral plate.
4 - Exopodal segment 2 of A2 with 1 seta.
5 - Spines of last thoracic segment exceeding at least the midlength of genital segment. Frontal spine large.
6 -, Frontal spine straight, not curved to rostrum by its top.
7 - Spines of last thoracic segment reaching third of genital segment length. External spine of exopodal segment 2 of P1 not reaching the midlength of the next segment.


Espèce Gaetanus pileatus - Planche 26 de figures morphologiquesGaetanus pileatus1 - Cephalon with frontal spine.
2 - Exopodal segment 3 of left P5 prolonged, stylet-like, never bilobated. It is always longer than exopodal segment 2.
3 - Frontal spine is in anterior part of cephalon dorsally (lateral view), well visible.
4 - Spines on last thoracic segment significantly shorter than in G. simplex.
5 - Endopod of left P5 shorter than half length of exopodal segment 1.

Ref. compl.:
Lysholm & al., 1945 (p.17); Sewell, 1948 (p.329, 500, 520, 529, 531, 539, 545, 556); C.B. Wilson, 1950 (p.233); Østvedt, 1955 (p.15: Table 3, p.63); Wickstead, 1962 (p.549, food & feeding); Grice & Hart, 1962 (p.287, table 3); Grice, 1963 a (p.495); De Decker & Mombeck, 1964 (p.12); Grice Hulsemann, 1965 (p.223); 1967 (p.15); 1968 (tab.2); Park, 1970 (p.476); Roe, 1972 (p.277, tabl.1, tabl.2); 1972 a (p.338); Bainbridge, 1972 (p.61, Appendix Table III: occurrence); Björnberg, 1973 (p.324, 386); Carter, 1977 (1978) (p.35); Deevey & Brooks, 1977 (p.256, tab.2, Station "S"); Vives, 1982 (p.291); Roe, 1984 (p.357); Guangshan & Honglin, 1984 (p.118, tab.); Madhupratap & Haridas, 1986 (p.105, tab.1); Lozano Soldevilla & al., 1988 (p.58); Heinrich, 1990 (p.17); Baessa De Aguiar, 1991 (1993) (p.101); Lampitt & al., 1993 (p.689, faecal pellets v.s. ingestion); Shih & Young, 1995 (p.67); Errhif & al., 1997 (p.422); Padmavati & al., 1998 (p.347); Suarez-Morales & Gasca, 1998 a (p108); Lapernat, 2000 (tabl. 3, 4); Razouls & al., 2000 (p.343, Appendix); Madhupratap & al., 2001 (p. 1345, vertical distribution vs. O2, figs.4, 5: clusters, p.1353); d'Elbée, 2001(tabl. 1); Holmes, 2001 (p.49); Park, W & al., 2004 (p.464, tab.1); Gaard & al., 2008 (p.59, Table 1, N Mid-Atlantic Ridge); Galbraith, 2009 (pers. comm.); Park & Ferrari, 2009 (p.143, Table 5, Appendix 1, biogeography); Teuber & al., 2013 (p.1, Table 1, 2, abundance vs oxygen minimum zone, respiration rates); in CalCOFI regional list (MDO, Nov. 2013; M. Ohman, comm. pers.); Bode M. & al., 2013 (p.1, Table 1, respiration rate & ETS activity); Bonecker & a., 2014 (p.445, Table II: frequency, horizontal & vertical distributions, Table IV: new geographical record)
NZ: 19

Carte de distribution de Gaetanus pileatus par zones géographiques
Espèce Gaetanus pileatus - Carte de distribution 3Issued from : J.M. Bradford & J.B. Jillett in New Zealand Ocean. Inst. Memoir, 86, 1980. [p.87, Figs.63].
Distribution of several species of Gaetanus in the Tasman Sea and around New Zealand.
Loc:
sub-Antarct. (Indian, SE Pacif.), South Africa (E), off E Tristan da Cunha, Angola, Congo, off E Sao Tomé Is. Brazil (off Rio de Janeiro), G. of Guinea, off Lagos, Cape Verde Is., off Mauritania-NW Cape Verde Is., off Morocco-Mauritania, Canary Is., off Madeira, off Portugal, off W Cabo Finisterre, Azores, Barbada Is., Caribbean Sea, G. of Mexico, Florida, off Bermuda: Station "S" (32°10'N, 64°30'W), Sargasso Sea, W Ireland, Greenland, Iceland, Faroe Is., Norwegian Sea, Bay of Biscay, Arabian Sea, Laccadive Is., Natal, off SW Sri Lanka, Indian, S Indian (subtropical convergence), Indo-Malaysia, Philippines, China Seas (East China Sea, South China Sea), Japan (Izu), Pacif. (W equatorial), Australia (Great Barrier), New Zealand, Hawaii, Vancouver Is., California, off Galapagos, off Peru, Pacif. (SE tropical), off Juan Fernandez Is., Chile.

Type locality: NE Atlantic (SW Ireland Slope).
N: 72
Lg.:
(1) F: 6,2; M: 4,7; (5) F: 6; (7) F: 5,74; (10) F: 5-4,6; (14) F: 6,1-5,9; M: 5,3-4,8; (17) F: 6,5; M: 5,3; (29) F: 4,1; (37) F: 6,7-4,9; M: 5,08-4,44; (38) F: 5,4-5,3; (40) F: 6,41-4,92; M: 5,08-4,44; (56) F: 6,14; (73) F: 5,82; (89) F: 5,93; (113) F: 6-5,5; (199) F: 6,56-5,55; M: 4,96-4,64; (201) F: 6,65-5,5; M: 4,85; 4,8; (249) F: 6,1-6; {F: 4,10-6,70; M: 4,44-5,30}
Rem.: méso-bathypélagique.
Sampling depth (sub-Antarct.) : 0-1000 m.
Signalé par Park, W & al. (2004) dans le Golfe d'Alaska (Icy Strait).
Voir aussi les remarques en anglais
Dernière mise à jour : 22/06/2020

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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2020. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Sorbonne Université, CNRS. Disponible sur http://copepodes.obs-banyuls.fr [Accédé le 27 septembre 2020]

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