Fiche d'espèce de Copépode
Calanoida ( Ordre )
    Arietelloidea ( Superfamille )
        Augaptilidae ( Famille )
            Haloptilus ( Genre )
Haloptilus acutifrons  (Giesbrecht, 1892)   (F,M)
Syn.: Hemicalanus acutifrons Giesbrecht, 1892 (p.384, 398, Descr.F, figs.F);
Hemicalanus spinifrons Sars, 1900 (p.95, figs.F);
Haloptilus spinifrons : Mrazek, 1902 (p.523); Dessier, 1979 (p.206); 1983 (p.89, Tableau 1, Rem., %);
? Haloptilus acutifrons : Grice, 1962 (p.223, figs.F, Rem.)
Ref.:
Giesbrecht & Schmeil, 1898 (p.119, Rem. F, fig.F); Sars, 1902 (1903) (p.122, figs.F); Farran, 1908 b (p.68); T. Scott, 1912 (1913) (p.536); Sars, 1925 (p.250, figs.F); Farran, 1926 (p.285); 1929 (p.209, 267); Rose, 1933 a (p.214, figs.F); Jespersen, 1934 (p.110, fig.28); 1940 (p.56); ? Farran, 1936 a (p.113, juv.5F); Mori, 1937 (1964) (p.77, figs.F); Sewell, 1947 (p.190); Brodsky, 1950 (1967) (p.364, figs.F); Vervoort, 1957 (p.135, Rem.); Chiba & al., 1957 (p.308); 1957 a (p.11); Tanaka, 1964 b (p.42); Owre & Foyo, 1967 (p.81, figs.F); Bradford, 1970 a (p.359: Rem.) Corral Estrada, 1970 (p.193, Rem.); Paiva, 1971 (p.26, figs.F, Rem.); Vidal, 1971 a (p.10, 114, fig.F); Matthews, 1972 (p.47); Björnberg & al., 1981 (p.654, figs.F); Roe, 1984 (p.358); Park, 1988 (p.2); Mazzocchi & al., 1995 (p.31, figs.F, Rem.); Chihara & Murano, 1997 (p.728, Pl.61: F); Bradford-Grieve & al., 1999 (p.882, 941, figs.F,M); Bradford-Grieve,1999 b (p.59, figs.F,M, Rem., figs. 172, 190); Vives & Shmeleva, 2007 (p.236, figs.F,M, Rem.)
Espèce Haloptilus acutifrons - Planche 1 de figures morphologiquesissued from : J.M. Bradford-Grieve in The Marine Fauna of New Zealand: Pelagic Calanoid Copepoda. National Institute of Water and Atmospheric Research (NIWA). NIWA Biodiversity Memoir, 111, 1999. [p.59, Fig.30].
Female: A, habitus (dorsal); B, idem (right lateral side); C, P5.

Male: D, habitus (dorsal); E, P5 (endopods: incomplete).


Espèce Haloptilus acutifrons - Planche 2 de figures morphologiquesIssued from: M.G. Mazzocchi, G. Zagami, A. Ianora, L. Guglielmo & J. Hure in Atlas of Marine Zooplankton Straits of Magellan. Copepods. L. Guglielmo & A. Ianora (Eds.), 1995. [p.32, Fig.3.2.1].
Female: A, habitus (dorsal); B, urosome (dorsal); C, forehead (ventral); D, idem (lateral right side); E, Md (mandibular palp); F, Md (cutting edge of mandibular blade).
Nota: Forehead produced into long, strong spine, slightly bent downward, with subtle filamentous rostrum. A1 exceeds end of body by 5-7 distal segments. Proportional lengths of urosomites and furca 39:11:9:14:27 = 100.


Espèce Haloptilus acutifrons - Planche 3 de figures morphologiquesIssued from : G.O. Sars in Résult. Camp. Scient. Prince Albert I, 69, pls.1-127 (1924). [Pl.LXXIV, figs.1-11].
Female: 1, habitus (dorsal); 2, idem (lateral left side); 3, forehead (lateral); 4, A2; 5, Md; 6, Mx1; 7, Mx2; 8, Mxp; 9, P1; 10, P3; 11, P5.


Espèce Haloptilus acutifrons - Planche 4 de figures morphologiquesissued from : T. Mori in The pelagic Copepoda from the neighbouring waters of Japan, 1937 (2nd edit., 1964). [Pl.39, Figs.1-4].
Female: 1, P5; 2, habitus (dorsal); 3, Mx1; 4, Mx2.


Espèce Haloptilus acutifrons - Planche 5 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.27 , Fig.4 ].
Hemicalanus acutifrons.
Female: 4, rostrum and sements 1 to 14 of A1 (ventral view).
Aes = aesthetasc


Espèce Haloptilus acutifrons - Planche 6 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.27 , Fig.4 ].
Hemicalanus acutifrons.
Female: 4, segments 14 to 25 of A1 (ventral view).:


Espèce Haloptilus acutifrons - Planche 7 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.27 , Figs.12, 18].
Hemicalanus acutifrons.
Female: 12, Mx1 (posterior view); 18, Md (biting edge of mandibular blade).
Li = inner lobe; Le = outer lobe; B = basipodite; Re = exopodite; Ri = endopodite


Espèce Haloptilus acutifrons - Planche 8 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.27 , Fig.26].
Hemicalanus acutifrons.
Female: 26, exopodite 3 of P2.


Espèce Haloptilus acutifrons - Planche 9 de figures morphologiquesissued from : G.D. Grice in Fish. Bull. Fish and Wildl. Ser., 1962, 61. [p.224, Pl.25, Figs.1-8].
Female (from equatorial Pacific): 1habitus (dorsal); 2, Md (mandibular palpus); 3, Md (biting edge); 4, P1; 5, P2; 6, P3; 7, P4; 8, P5.


Espèce Haloptilus acutifrons - Planche 10 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.42, Fig.12, 20]. As Hemicalanus oxycephalus.
Female: 12, forehead (lateral); 20, urosome (ventral).


Espèce Haloptilus acutifrons - Planche 11 de figures morphologiquesissued from : J.M. Bradford-Grieve, E.L. Markhaseva & C.E.F. Rocha & B. Abiahy in South Atlantic Zooplankton, Edit. D. Boltovskoy. 1999. Vol.2. Copepoda. [p.1041, Fig. 7.265].
Female: Mx1.

Exopodite of Mx1 with 11 setae; outer lobe 1 with 10 setae.

Ref. compl.:
Pearson, 1906 (p.28); Rose, 1924 d (p.481); Wilson, 1942 a (p.188); Massuti Alzamora, 1942 (p.111); Lysholm & al., 1945 (p.37); Sewell, 1948 (p.330, 496, 504, 509, 522, 534, 538, 547, 558, 559, 567); C.B. Wilson, 1950 (p.235); Hoenigman, 1955 (p.49); Yamazi, 1958 (p.150, Rem.); Fagetti, 1962 (p.31); Grice & Hart, 1962 (p.287, 293: Rem.); V.N. Greze, 1963 a (tabl.2); Duran, 1963 (p.23); Grice, 1963 a (p.496); Ahlstrom & Thrailkill, 1963 (p.57, Table 5, abundance); Gaudy, 1963 (p.27, Rem.); M.W. Johnson, 1963 (p.89, Table 1, 2); Björnberg, 1963 (p.54, Rem.); De Decker & Mombeck, 1964 (p.12); Grice & Hulsemann, 1965 (p.224); Mazza, 1966 (p.72); Pavlova, 1966 (p.44); Harding, 1966 (p.17, 65, 71); Fleminger, 1967 a (tabl.1); Grice & Hulsemann, 1967 (p.19); Dunbar & Harding, 1968 (p.319); Delalo, 1968 (p.138); Park, 1970 (p.478); Deevey, 1971 (p.224); Roe, 1972 (p.277, tabl.1, tabl.2); 1972 c (p.1033); Apostolopoulou, 1972 (p.328, 362); Bainbridge, 1972 (p.61, Appendix Table II, vertical distribution %); Björnberg, 1973 (p.347, 386); Corral Estrada & Pereiro Muñoz, 1974 (tab.I); Vives & al., 1975 (p.50, tab.II, III); Deevey & Brooks, 1977 (p.156, tab.2, Station "S"); Carter, 1977 (1978) (p.36); Vaissière & Séguin, 1980 (p.23, tab.1); Buchanan & Sekerak, 1982 (p.41, vertical distribution); Vives, 1982 (p.293); Kovalev & Shmeleva, 1982 (p.84); Greze & al., 1983 (p.17); Scotto di Carlo & al., 1984 (p.1044); Cummings, 1984 (p.163, Table 2); Longhurst, 1985 (tab.2); Brenning, 1985 a (p.28, Table 2); 1986 (p.14, Rem.); Brinton & al., 1986 (p.228, Table 1); Madhupratap & Haridas, 1986 (p.105, tab.1); Lozano Soldevilla & al., 1988 (p.60); Kosobokova, 1989 (p.26); Heinrich, 1990 (p.18); Suarez & al., 1990 (tab.2); Hattori, 1991 (tab.1, Appendix); Scotto di Carlo & al., 1991 (p.271); Suarez & Gasca, 1991 (tab.2); Suarez, 1992 (App.1); Mumm, 1993 (tab.1, fig.2); Seguin & al., 1993 (p.23); Richter, 1994 (tab.4.1a); Shih & Young,1995 (p.68); Hanssen, 1997 (tab.3.1); Hure & Krsinic, 1998 (p.102); Padmavati & al., 1998 (p.347); Gilabert & Moreno, 1998 (tab.1, 2); Suarez-Morales & Gasca, 1998 a (p108); Kosobokova & al., 1998 (tab.2); Voronina & Kolosova, 1999 (p.71); Razouls & al., 2000 (p.343, tab. 5, Appendix); Lopez-Salgado & al., 2000 (tab.1); Fernandez-Alamo & al., 2000 (p.1139, Appendix); Kosobokova & Hirche, 2000 (p. 2029, tab. 2); Holmes, 2001 (p.12, Rem.); Auel & Hagen, 2002 (p.1013, tab.2); Vukanic, 2003 (139, tab.1); Hsiao & al., 2004 (p.325, tab.1); Isari & al., 2006 (p.241, tab.II); Dur & al., 2007 (p.197, Table IV); Khelifi-Touhami & al., 2007 (p.327, Table 1); McKinnon & al., 2008 (p.843: Tab.1); Ayon & al., 2008 (p.238, Table 4: Peruvian samples); Neumann-Leitao & al., 2008 (p.799: Tab.II, fig.6); Morales-Ramirez & Suarez-Morales, 2008 (p.517); Fernandes, 2008 (p.465, Tabl.2); Raybaud & al., 2008 (p.1765, Table A1); Galbraith, 2009 (pers. comm.); C.E. Morales & al., 2010 (p.158, Table 1); Kosobokova & Hopcroft, 2010 (p.96, Table 1, fig.7); Mazzocchi & Di Capua, 2010 (p.424); Medellin-Mora & Navas S., 2010 (p.265, Tab. 2); Dvoretsky & Dvoretsky, 2010 (p.991, Table 2); Kosobokova & al., 2011 (p.29, Table 2, Rem.: Arctic Basins); Hsiao S.H. & al., 2011 (p.475, Appendix I); Tutasi & al., 2011 (p.791, Table 2, abundance distribution vs La Niña event); Pillai H.U.K. & al., 2011 (p.239, Table 3, vertical distribution); Andersen N.G. & al., 2011 (p.71, Fig.3: abundance); Uysal & Shmeleva, 2012 (p.909, Table I); Miloslavic & al., 2012 (p.165, Table 2, transect distribution); in CalCOFI regional list (MDO, Nov. 2013; M. Ohman, comm. pers.); Lidvanov & al., 2013 (p.290, Table 2, % composition); Mazzocchi & al., 2014 (p.64, Table 4, abundance); Benedetti & al., 2016 (p.159, Table I, fig.1, functional characters); Benedetti & al., 2018 (p.1, Fig.2: ecological functional group); Belmonte, 2018 (p.273, Table I: Italian zones); Acha & al., 2020 (p.p.1, Table 3: occurrence % vs ecoregions).
NZ: 22

Carte de distribution de Haloptilus acutifrons par zones géographiques
Loc:
Antarct. (Weddell Sea, SE Atlant.), sub-Antarct. (S Tasmania, SE Pacif., Straits of Magellan: Pacific area), South Africa (E), Congo, off Lagos, Cape Verde Is., off Morocco-Mauritania, Canary Is., off Madeira, off W Cape Finisterre, Azores, S Brazil, off Amazon, Caribbean Colombia, Caribbean Sea, G. of Mexico, Cuba, Florida, Sargasso Sea, off Bermuda: Station ‘’ S’’ (32°10’N, 64°30’W), Baffin Bay, W Greenland, Greenland Sea, S Iceland, Arctic (all polar Basins), Arctic. (central, Fletcher's Ice Is.), Nansen Basin, Lomonosov Ridge, Barents Sea, Laptev Sea, Canadian abyssal plain, Canada Basin, Chukchi Sea, E Beaufort Sea, off W Ireland, Porcupine bank, Norway, North Sea, Bay of Biscay, Baie Ibéro-marocaine, Medit. (Alboran Sea, Gulf of Annaba, Castellon, Baleares, Ligurian Sea, Tyrrhenian Sea, Strait of Messina, Adriatic Sea, Mljet Is., Ionian Sea, Aegean Sea, Johnston Bank, Lebanon Basin), Red Sea, G. of Aden, G. of Oman, Arabian Sea, Natal, Indian, Bay of Bengal, Andaman Sea (Batten Island); Australia (North West Cape), Philippines, China Seas (East China Sea, South China Sea), Taiwan (SW, E), Japan (Izu, Tanabe Bay, off Sanriku), Bikini Is., Pacif. (tropical), ? Australia (Great Barrier), New Zealand (North Island: NE & NW, SE South Island), N Tasman Sea, Hawaii, Vancouver Is., California, W Baja California, Gulf of California, G. of Tehuantepec, off W Guatemala, W Costa Rica, off W Costa Rica, off SW Panama, Pacif. (SE tropical), SE Easter Is., Galapagos-Ecuador, Peru, Chile (., Concepcion)
N: 158
Lg.:
(1) F: 3,3; (22) F: 3,2-2,6; (35) F: 3,9-3,5; (36) F: 4,66-4,35; (38) F: 3,5-3,2; (46) F: 2,9-2,6; (65) F: 3,2; (69) F: 2,99; (91) F: 3-2,5; ? (101) F: 3,8; (134) F: ± 4; (199) F: 3,95-3,12; M: 3,19; (237) F: 2,0; (310) F: 3,86-3,64; (765) F: 3-2,6; M: 2,4-2,1; (1108) F: 3,8; {F: 2,00-4,66; M: 2,1-3,19}
Rem.: épi-bathypélagique.
Sampling depth (Antarct., sub-Antarct.): 100-250 m.
La description originale du mâle ne nous est pas connue, mais indiquée in Bradford-Grieve & al. (1999) et Bradford-Grieve (1999 b, p.59, Fig.30 et remarques p.60).
Voir aussi les remarques en anglais
Dernière mise à jour : 03/04/2020

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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2020. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Sorbonne Université, CNRS. Disponible sur http://copepodes.obs-banyuls.fr [Accédé le 28 septembre 2020]

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