Species Card of Copepod
Calanoida ( Order )
    Clausocalanoidea ( Superfamily )
        Scolecitrichidae ( Family )
            Scottocalanus ( Genus )
Scottocalanus securifrons  (T. Scott, 1894)   (F,M)
Syn.: Scolecithrix securifrons T. Scott, 1894 b (p.47, figs.F, no M); Giesbrecht & Schmeil, 1898 (p.49, Rem.F); ? Cleve, 1904 a (p.197); van Breemen, 1908 a (p.76);
Lophothrix securifrons : Wolfenden, 1904 (p.120, figs.F); 1911 (p.268);
Scottocalanus acutus Sars, 1905 c (p.7, Rem.F);
Amallophora securifrons : Wolfenden, 1904 (p.145);
Scolecithrix cuneifrons Willey, 1918 (1919) (p.194, figs.F,M); Tremblay & Anderson, 1984 (p.6);
(?) Scottcalanus securifrons : Tanaka, 1937 (p.259, Rem.F, figs.F);
Scottocalanus securifer : De Decker, 1968 (p.45);
Scottocalanus securiformis: Sameoto & Lewis, 1980
Ref.:
Farran, 1908 b (p.57, Rem.); A. Scott, 1909 (p.104, figs.F,M); With, 1915 (p.220, figs.F,M); Lysholm & Nordgaard, 1921 (p.21); Sars, 1925 (p.160, figs.F,M); Farran, 1926 (p.267, Rem.); 1929 (p.209, 251); Rose, 1929 (p.26); 1933 a (p.144, figs.F,M); Jespersen, 1940 (p.36); Lysholm & al., 1945 (p.26); Sewell, 1947 (p.143); Brodsky, 1950 (1967) (p.243, figs.F,M); Marques, 1953 (p.103); 1959 (p.211); Tanaka, 1961 a (p.141, Rem.F,M, figs.M); Grice, 1962 (p.213, figs.F); Paiva, 1963 (p.45, figs.F); Vervoort, 1965 (p.36, Rem.); Owre & Foyo, 1967 (p.63, figs.F,M, Rem.: p.65); Ramirez, 1969 (p.66, figs.F,M, Rem.); Tanaka, 1969 (p.275); Gopalakrishnan, 1974 (p.274, figs.F,M, Rem.); Björnberg & al., 1981 (p.637, figs.F); Park, 1983 (p.197, 200, Redescr. F, M, figs.F,M); Bradford & al., 1983 (p.116, figs.F,M, Rem.); Roe, 1984 (p.357); Campaner, 1989 (p.229, figs.F,M, Rem.); Schnack, 1989 (p.137, tab.1, fig.6: Md); Nishida & al., 1991 (p.527); He & al., 1992 (p.250); Chihara & Murano, 1997 (p.902, Pl.177: F,M); Mulyadi, 1998 a (p.389, Redescr.M, figs.M); Bradford-Grieve & al., 1999 (p.881, 931, figs.F,M); Boxshall & Halsey, 2004 (p.189, figs.F); Mulyadi, 2004 (p.31, figs.M, Rem.);Vives & Shmeleva, 2007 (p.811, figs.F,M, Rem.); Park & Ferrari, 2009 (p.143, Appendix 1)
Species Scottocalanus securifrons - Plate 1 of morphological figuresissued from : Mulyadi in The Raffles Bull. Zool., 1998, 46 (2). [p.390, Fig.6].
Male (from 07°29'S, 121°05'E): a, habitus (dorsal); b, forehead (left lateral side); c, last thoracic segment and genital somite (left lateral side); d, rostrum (frontal view); e, P2; f, P3; g, left P5; h, distal segment of right P5; i, right P5.


Species Scottocalanus securifrons - Plate 2 of morphological figuresIssued from : J.M. Bradford, L. Haakonssen & J.B. Jillett in Mem. N.Z. oceanogr. Inst., 1983, 90. [p.118, Fig.73].
Female: A, habitus (lateral right side); B, genital segment (dorsal); C, genital segment and P5 (lateral right side); D, rostrum; E, endopod of P2; F, P5.

Male: G, habitus (lateral left side); H, P5.


Species Scottocalanus securifrons - Plate 3 of morphological figuresissued from : T. Park in Antarct. Res. Ser. Washington, 1983, 38 (3). [p.201, Fig.19].
Female: a, habitus (dorsal); b, idem (lateral left side); c, forehead (lateral; d, rostrum (anterior); e, urosome (ventral): f, last thoracic segments and urosome (lateral left side); g, forehead (dorsal); h, A2; i, Md.

Nota: Cephalosome and 1st matasomal segment fused, indicated laterally by a fine line, 4th and 5th separate on dorsal side. Urosome about 20/100 length of prosome. Spermatheca with long, somewhat inflated vesicle.


Species Scottocalanus securifrons - Plate 4 of morphological figuresissued from : T. Park in Antarct. Res. Ser. Washington, 1983, 38 (3). [p.202, Fig.20].
Female: a, Mx1; b, distal part of Mx2; c, Mxp; d, P1 (posterior); e, P2 (posterior); f, P3 (posterior); g, P4 (posterior); h, P5 (anterior).


Species Scottocalanus securifrons - Plate 5 of morphological figuresissued from : T. Park in Antarct. Res. Ser. Washington, 1983, 38 (3). [p.203, Fig.21].
Male: a, forehead (lateral); b, last thoracic segments and urosome (lateral left side); c, forehead (dorsal); d, last thoracic segments and urosome (dorsal); e, rostrum (anterior); f, A2; g, Md; h, Mx1; i, Mx2.

Nota: Urosome about 30/100 length of prosome. A1 reaching close to distal end of 3rd urosomal segment.


Species Scottocalanus securifrons - Plate 6 of morphological figuresissued from : T. Park in Antarct. Res. Ser. Washington, 1983, 38 (3). [p.204, Fig.22].
Male: a, Mxp; b, P1 (anterior); c, P2 (posterior); d, P3 (posterior); e, P4 (posterior); f, P5 (anterior); g, P5 (posterior); h, distal part of right P5 exopod; i, left P5 (medial); j, distal part of left P5 exopod (posterior).


Species Scottocalanus securifrons - Plate 7 of morphological figuresissued from : F.C. Ramirez in Contr. Inst. Biol. mar., Buenos Aires, 1969, 98. [p.64, Lam. XII, figs.92 ].
Female (from off Mar del Plata): 92, P5
Scale bar in mm: 0.05.


Species Scottocalanus securifrons - Plate 8 of morphological figuresissued from : F.C. Ramirez in Contr. Inst. Biol. mar., Buenos Aires, 1969, 98. [p.62, Lam. XI, figs.79, 81, 82 ].
Female (from off Mar del Plata): 82, habitus (dorsal).

Male: 79, P5; 81, habitus (lateral right side).
Scale bars in mm: 2 ( (81, 82);


Species Scottocalanus securifrons - Plate 9 of morphological figuresIssued from : G.O. Sars in Résult. Camp. Scient. Prince Albert I, 69, pls.1-127 (1924). [Pl.XLV, figs.1-8].
Female: 1, habitus (dorsal); 2, idem (lateral left side); 3, forehead (lateral); 4, rostrum (frontal view); 5, P5; 6, P5 (lateral view).

Male: 7, habitus (dorsal); 8, P5.


Species Scottocalanus securifrons - Plate 10 of morphological figuresissued from : A. Scott in Siboga-Expedition, 1909, XIX a. [Plate XXVIII, Figs.1-9].
Male (from Indonesia-Malaysia): 1, habitus (dorsal); 2, forehead (lateral); 3, last thoracic and genital segments (left side); 4, rostrum; 5, A1; 6, Mx2 (distal portion); 7, P2; 8, P4; 9, P5.


Species Scottocalanus securifrons - Plate 11 of morphological figuresissued from : A. Scott in Siboga-Expedition, 1909, XIX a. [Plate XXV, Figs.1-9].
Female: 1, habitus (dorsal); 2, forehead (lateral); 3, last thoracic and genital segments (left side); 4, rostrum; 5, A1; 6, Mx2 (distal portion); 7, P2; 8, P4; 9, P5.


Species Scottocalanus securifrons - Plate 12 of morphological figuresissued from : S. Ohtsuka & R. Huys in Hydrobiologia, 2001, 453/454. [p.452, Fig.7, F].
Male (from Japan): F, terminal exopodal segment of left P5.
Scale bar = 0.2 mm.


Species Scottocalanus securifrons - Plate 13 of morphological figuresissued from : O. Tanaka in Publ. Seto Mar. Biol. Lab., 1961, IX (1). [p.142, Fig.106].
Male: a, last thoracic segment (lateral, right side); b, rostrum (frontal view); c, P2 (posterior view); d, P5; e, distal segments of right P5.
Urosme 5-segmented; segments and caudal rami in the proportional lengths 15:29:21:19:6:10 = 100; 2nd to 4th segments stiated with fine teeth on the distal margin. A1 extends to the distal end of the 3rd urosomal segment (segments 8-9, 20-21 fused on the left side. Mouth parts and swimming legs as those of the female.


Species Scottocalanus securifrons - Plate 14 of morphological figuresissued from : G.D. Grice in Fish. Bull. Fish and Wildl. Ser., 1962, 61. [p.212, Pl.19, Figs.12-15].
Female (from 00°03'S, 157°00'E): 12, forehead (lateral); 13, posterior part of thorax and urosome (dorsal); 14, same (lateral, left side); 15, P5.

Species Scottocalanus securifrons - Plate 15 of morphological figuresissued from : C. With in The Danish Ingolf-Expedition, Copepoda I, 1915, III, 4. [p.220, Text-fig. 71].
Female (from 51°32'N, 12°03'W): a, last thoracic segment and urosome (dorsal); b, same (left lateral); c, genital segment (half right); d, segments 8-10 of A1.


Species Scottocalanus securifrons - Plate 16 of morphological figuresissued from : C. With in The Danish Ingolf-Expedition, Copepoda I, 1915, III, 4. [p.221, Text-fig. 72].
Male: a, last thoracic segment and urosome (dorsal); b, segments VII-XIV of A1; c, right P5; exopodal segments 2, 3 of right P5 (right view); e, two first segments of left P5 (dorsal view).


Species Scottocalanus securifrons - Plate 17 of morphological figuresissued from : C. With in The Danish Ingolf-Expedition, Copepoda I, 1915, III, 4. [Pl. VIII, Fig. 13].
Male: last two thoracic segments with P5 and urosome (left lateral); b, left P5; c, same (distal segments).


Species Scottocalanus securifrons - Plate 18 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.62, Figs.400-402].
Female (from 23°31'-25°40'N, 82°32'-79°43'W): 400, habitus (dorsal); 401, last thoracic segment and genital segment (left lateral); 402, P5.


Species Scottocalanus securifrons - Plate 19 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.63, Figs.409-410].
Female: 409, rostrum.

Male: 410, rostrum.
Nota: Terminal projections of the bifurcated rostrum minute.


Species Scottocalanus securifrons - Plate 20 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.65].
Female: Proportional lengths of the segments on A1.


Species Scottocalanus securifrons - Plate 21 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.62, Fig.403].
Male: habitus (dorsal).


Species Scottocalanus securifrons - Plate 22 of morphological figuresissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. 1967. [p.23, Fig.98].
Male: 98, P5.


Species Scottocalanus securifrons - Plate 23 of morphological figuresissued from : S.B. Schnack in Crustacean Issue, 1989, 6. [p.143, Fig.6: 8].
8, Scottocalanus securifrons (from off NW Africa, upwelling region): Cutting edge of Md.

Compl. Ref.:
Sewell, 1948 (p.329, 502, 516, 519, 521, 530, 532, 546); C.B. Wilson, 1950 (p.340); King & Hida, 1955 (p.11); Østvedt, 1955 (p.15: Table 3, p.69); Grice, 1963 a (p.495); Unterüberbacher, 1964 (p.24); De Decker & Mombeck, 1964 (p.14); Fleminger, 1967 a (tabl.1); Vinogradov, 1968 (1970) (p.258); Morris, 1970 (p.2301); Park, 1970 (p.476); Roe, 1972 (p.277, tabl.1, tabl.2); 1972 b (p.530); Vives & al., 1975 (p.43, tab.II, XII); Deevey & Brooks, 1977 (p.256, tab.2, Station "S"); Carter, 1977 (1978) (p.35); Dessier, 1979 (p.205); Vives, 1982 (p.292); Cummings, 1984 (p.163, Table 2); Guangshan & Honglin, 1984 (p.118, tab.); Brenning, 1984 (p.4, Rem.); Tremblay & Anderson, 1984 (p.6, Rem.); Brenning, 1985 a (p.28, Table 2); Madhupratap & Haridas, 1986 (p.105, tab.1); Lozano Soldevilla & al., 1988 (p.59); Heinrich, 1990 (p.17); Suarez & Gasca, 1991 (tab.2); Hattori, 1991 (tab.1, Appendix, comme secrifrons); Hernandez-Trujillo, 1991 (1993) (tab.I); Suarez, 1992 (App.1); Gopalakrishnan & Balachandran, 1992 (p.167, figs.1, 6, Table 1, 2); Yen & al., 1992 (p.495, tab.1, mechanoreception); Webber & Roff, 1995 (tab.1); Shih & Young, 1995 (p.74); Kotani & al., 1996 (tab.2); Suarez-Morales & Gasca, 1998 a (p.111); Neumann-Leitao & al., 1999 (p.153, tab.2); Lapernat, 2000 (tabl.3, 4); Razouls & al., 2000 (p.343, Appendix); Lopez-Salgado & al., 2000 (tab.1); d'Elbée, 2001(tabl. 1); Holmes, 2001 (p.60); Yamaguchi & al., 2002 (p.1007, tab.1); Sameoto & al., 2002 (p.13); Hsiao & al., 2004 (p.326, tab.1); Lo & al., 2004 (p.89, tab.1); Hwang & al., 2007 (p.25); Kuriyama & Nishida, 2006 (p.300: Tab.II, fig.7); Ikeda & al., 2006 (p.1791, Table 2); Dur & al., 2007 (p.197, Table IV); Neumann-Leitao & al., 2008 (p.799: Tab.II, fig.6); Galbraith, 2009 (pers. comm.); Dias & al., 2010 (p.230, Table 1); Medellin-Mora & Navas S., 2010 (p.265, Tab. 2); Hsiao S.H. & al., 2011 (p.475, Appendix I)
NZ: 20

Distribution map of Scottocalanus securifrons by geographical zones
Species Scottocalanus securifrons - Distribution map 3issued from : T.C. Gopalakrishnan & T. Balachandran in Oceanogr. Indian Ocean, 1992. [p.171, fig.6, C].
Distribution of Scottocalanus securifrons in the Indian Ocean.
Nota: Main occurnces between 10° N - 20° S.
Loc:
Antarct. (Indien), sub-Antarct. (Indien, Pacif. SW), Afr. S (E & W), off Tristan da Cunha E, Namibie, Angola, G. de Guinée, Argentine, Brésil (off Macaé, off Natal), Is. du Cap Vert, off Mauritanie-Is. du Cap Vert NW, off Mauritanie, Canaries, off Madère, off Cap Finisterre W, Baie Ibéro-marocaine, Brésil S, off Amazone, Mer des Antilles, Caribbean Colombia, Jamaïque, G. du Mexique, Floride, Mer des Sargasses, off Bermuda: Station ‘’ S’’ (32°10’N, 64°30’W), Açores, G. de Gascogne, off Irlande W, off Ecosse W, off Nova Scotia E, Flemish Cap, Islande S, Féroé, Mer de Norvège, Mer du Nord N, Mer Arabe, Natal, Madagascar (Nosy Bé), Indien, Indonesia-Malaysia, Flores Sea, Philippines, mers de Chine (East China Sea, South China Sea), Taiwan (S, E, Mienhua Canyon), Japon (Izu, Onagawa, off Sanriku), Station Knot, Pacif. W (équatorial), Nouvelle-Zélande, Pacif. équatorial central, off Hawaii N, Oregon (rare), Californie, Basse Californie W, off Marquises NE, off Tuamotu NE, Galapagos, Equateur, Pérou, Pacif. SE (tropic.)
N: 95
Lg.:
(1) F: 4,5; (5) F: 4,3; M: 4,75; (6) F: 4,08-3,66; M: 5,33-4,75; (7) F: 4,49; M: 4,98; (9) F: 4,6-3,38; M: 5,3-4,5; (16) F: 4,4-4,05; M: 4,9-4,55; (22) F: 4,9-4,3; M: 4,9-4,7; (35) F: 4,5-3,88; (38) M: 4,9-4,8; (47) F: ± 4; (54) F: 4,01; (73) F: 4,57-4,31; M: 4,99; (75) (101) F: 4,08; (105) F: 4,07; M: 3,79; (108) F: 4,29; M: 4,81; (128) F: 4; (199) F: 4,53-4,1; M: 4,96-4,48; (254) F: F: ±4,3; M: 5; (432) F: 4; (539) F: 4,2; M: 4,57; (777) M: 4,55; (800) F: 4,69-4,53; (1000) F: 4,5 ± 0,1; M: 4,8 ± 0,1; (1111) F: 4,2; 4,7; M: 5,09; (1122) M: 4,55; {F: 3,38-4,90; M: 3,79-5,33}
Rem.: Meso-bathypelagic, but can also migrate to the surface (Roe, 1972; 1972 b, p.530). Sargasso Sea: 0-1000 m (Deevey & Brooks, 1977, Station "S");
For Østvedt (1955) it is probably only an exceptional circumstance that this species penetrates as far into the Norwegian Sea as latitude 66° N.
Vervoort (1965, p.36, 37) does not share the opinion of Brodsky (1950) concerning the identification of the female of Tanaka (1937). For Vervoort (1965, p.37), the reference to this species in Canu (1886, Norman (1903), Cleve (1904 a), Pearson (1906), Farran (1908), Paulsen (1909), Stebbing (1910) and Wolfenden (1911) must be taken with some reserve as no descriptions are added to their records, so that other species may have been included. S. securifrons must be considered as a species from deep and intermediate water layers. For Owre & Foyo (1967, p.65) S. securifrons is grossly similar to S. corystes but larger.
Park (1983, p.205) notes the occurence of the species in the north of the Antarctic convergence, the southermost was between 46°40'S, 170°03'W.
For Schnack (1989) the Itoh's index value of the mandibular gnathobase = 552.

See remarks and figures in Macandrewella chelipes.
Last update : 26/04/2013
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