Species Card of Copepod
Calanoida ( Order )
    Diaptomoidea ( Superfamily )
        Tortanidae ( Family )
            Tortanus ( Genus )
                Eutortanus ( Sub-Genus )
Tortanus (Eutortanus) dextrilobatus  Chen & Zhang, 1965   (F,M)
Ref.:
Chen & Zhang, 1965 (p.118, 128, figs.F,M); Ohtsuka & al., 1992 (p.112, figs.F,M, Rem.); Ferrari & Ueda, 2005 (p.346, figs.juv,F,M)
Species Tortanus (Eutortanus) dextrilobatus - Plate 1 of morphological figuresissued from: Q.-c Chen & S.-z. Zhang in Studia Marina Sinica, 1965, 7. [Pl.53, 6-11].
Female (from Aoy, Formosa Strait): 6, habitus (dorsal); 7, idem (lateral right side); 8, P5 (anterior).
Nota: Head and 1st thoracic segment separate, 4th and 5th .fused. Postero-lateral angles of last thoracic segment protuberant and symmetrical. Urosome 3-segmented. Anal segment asymmetrical, its upper right side with a triangular process. Furcal rami large, 4 times as long as wide, right furcal seta longer than the left. P5 not very symmetrical (left longer than right), inner margin of both legs with bristles.

Male: 9, habitus (dorsal); 10, geniculated portion of right A1; 11, P5 (posterior).
Nota: Urosome 5-segmented, anal segment very short. Furcal rami like those of female. Right A1 geniculate, 17-segmented, anterior border of 13th-15th armed with denticles. right P5 chelate


Species Tortanus (Eutortanus) dextrilobatus - Plate 2 of morphological figuresissued from : S. Ohtsuka, Y.H. Yoon & Y. Endo in The J. Oceanol. Soc. Korea, 1992, 27 (2). [p.115, Fig.2].
Female (from Somjin River, S Korea): A, habitus (dorsal); B, idem (lateral left side); C-E, 5th pedigerous somite and urosome (dorsal); F, idem (lateral right side); G, idem (lateral left side); H, genital operculum; I, A2; J, Md.
Nota: Cephalosome with transverse groove medially and dorsal median protuberance posteriorly. Head and 1st thoracic segment separate, 4th and 5th fused (suture line slifghtly visible in lateral view). Eye large, reddish. A1 17-segmented. A2 with coxa and basis partly fused, coxa with small medial seta, basis with 2 medial setae of unequal lengths; basis and endopod completely fused; endopod 2-segmented, proximal with 1 seta and row of spinules subterminally, distal with 6 setae terminally and row of spinules subterminally; exopod 3-segmented, proximal unarmed, middle with 3 setae, distal with2 terminal setae. Endopod of Md: 2-segmented, proximal segmnt without seta, distal with 5 setae; exopod 5-segmented incompletely fused with 3 rows of minute spinules. Urosome 3-segmented, 2nd urosomal segment with posterior dorsal protuberance on right side, 3rd segment having large irregular dorsolateral process on anterior right side. Anal segment incompletely fused with left caudal ramus and almost completely with right one.
variation: The dorsolateral process on the right side of the 3rd urosomal somite is slightly variable in shape


Species Tortanus (Eutortanus) dextrilobatus - Plate 3 of morphological figuresissued from : S. Ohtsuka, Y.H. Yoon & Y. Endo in The J. Oceanol. Soc. Korea, 1992, 27 (2). [p.116, Fig.3].
Female: A, A1; B, Mx1; C, Mx2; D, Mxp; E, P1.
Nota: Mx1 with basis, endopod and exopod completely absent; praecoxal arthrite stout, with 12 plumose setae and 1 minute setule (indicated by arrowhead); coxal endite with 3 stout, spinulose setae terminally. Mx2 with proximal and distal endites of praecoxa and coxa having 1, 2, 1 and 3 setae, respectively; basal endite with 1 developed and 2 minute setae; endopod with setal formula 1, 2, 2, 2 (1 seta on 2nd segment rudimentary). Mxp with setal formula on praecoxal and coxal endites: 0, 2, 2, 1; basis unarmed; endopod 1-segmented with 3 inner plumose setae and 1 outer seta.


Species Tortanus (Eutortanus) dextrilobatus - Plate 4 of morphological figuresissued from : S. Ohtsuka, Y.H. Yoon & Y. Endo in The J. Oceanol. Soc. Korea, 1992, 27 (2). [p.118, Fig.4].
Female: A, P2 (anterior); B, P3 (anterior); C, P4 (anterior); D, P5 (anterior).
Nota: Right and left coxal segments of P5 fused

Male: E, habitus (dorsal); F, 5th pedigerous somite and urosome (lateral right side, relatively stout sensilla indicated by arrows); G, idem (dorsal).
Nota: 4th and 5th pedigerous somites incompletely fused. Urosome 5-segmented. Genital somite with opening on left lateral side. 1st abdominal somite with 2 relatively stout sensilla ventrolaterally (indicated by arrows in fig.4 F. Anal somite separate from caudal rami. Right caudal ramus slightly longer than left.


Species Tortanus (Eutortanus) dextrilobatus - Plate 5 of morphological figuresissued from : S. Ohtsuka, Y.H. Yoon & Y. Endo in The J. Oceanol. Soc. Korea, 1992, 27 (2). [p.119, Fig.5 A-B].
Male: A, right A1; B, P5 (anterior).
Right A1 geniculate, indistinctly 16-segmented but consisting of 14 fully separated segments (geniculation between 14th and 15th). Right and left coxal segments of P5 fused; basis and 1st exopod segment of right leg fused; basis of left leg with outer seta subterminally, proximal exopod segment with short, thick spine subterminally, distal exopod segment with tuft of hairs and small seta on proximal inner margin, tuft of spinules along distal inner margin and 1 medial inner and 2 terminal short spines, terminal outer portion with 13 transverse and 3 longitudinal ridges.
Variation: The number of the ridges along the inner margin of right P5 is 24 or 25 (number of individuals examined: 3).

Compl. Ref.:
Chen Q., 1983 (p.133); Dai & al., 1991 (tab.1); Myung & al., 1994 (tab.1); Shih & Young, 1995 (p.74); Ohtsuka & al., 1995 (p.159); Cordell & al., 1997 (On line pdf.); Orsi & Ohtsuka, 1999 (p.128); Bollens & al., 2002 (p.87); Hooff & Bollens, 2004 (p.167, predatory imact); Ohtsuka & al., 2008 (p.115, Table 6); Cordell & al., 2008 (p.753); Bollens & al., 2011 (p.1358, Table III, fig.8); Soh & al., 2013 (p.723: Rem.)
NZ: 3

Distribution map of Tortanus (Eutortanus) dextrilobatus by geographical zones
Species Tortanus (Eutortanus) dextrilobatus - Distribution map 2issued from : S. Ohtsuka, Y.H. Yoon & Y. Endo in The J. Oceanol. Soc. Korea, 1992, 27 (2). [p.120, Fig.6].
Schematic illustration of distribution patterns of five sepecies of the subgenus Eutortanus.
Loc:
China Seas (East China Sea, South China Sea, Xiamen Harbour), Amoy, S Korea (Seomjin River estuary, Asan Bay), Ariake Sea, NE Pacif. (San Francisco Estuary)
N: 12
Lg.:
(290) F: 2,25-2; M: 2,05-2; (641) F: 1,79-1,74; M: 1,76-1,53; {F: 1,74-2,25; M: 1,53-2,05}
Rem.: Introduced in San Francisco by ship's ballast waters.
According to Cordell & al., 2008 (p.753) the 1st record of this species in San Francisco Estuary is 1993.
Last update : 19/02/2014
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