List species and varieties by family
Tharybidae Sars, 1902 ( Clausocalanoidea )
(*) Cenognatha Bradford-Grieve, 2001
Rem.: type: Neoscolecithrix antarctica. 1 sp. (provisionally). Bradford-Grieve, 2004 (p.285) includes in this genus the two species Neoscolecithrix caetanoi and N. farrani. Boxshall & Halsey, 2004 (p.186). Cf. Scolecitrichidae (largo sensu).
Cenognatha antarctica    Hulsemann, 1985   (F, Juv.M)
Syn.: Neoscolecithrix antarctica Hulsemann, 1985 (p.55, Descr., figs. C5, F & M); Razouls & al., 2000 (p.343, tab. 2, Appendix)
Ref.: Bradford-Grieve, 2001 a (p.792, Rem.); Bradford-Grieve, 2004 (p.285)
Rem.: Pour Vyshkvartzeva (2000, p.217) cette espèce appartiendrait aux Scolecitrichidae. Bradford-Grieve, 2004 (p.285)

(*) Neoscolecithrix Canu, 1896 (part.)
Rem.: type: Neoscolecithrix koehleri Canu,1896. Ohtsuka, Boxshall & Fosshagen (2003, p.61-62) believe that this genus belongs (largo sensu) to Scolecitrichidae.
Neoscolecithrix antarctica    Hulsemann, 1985   (F, juv.M)
Ref.: Hulsemann, 1985 (p.55, Descr., figs. C5, F & M)
Rem.: Cf. Cenognatha antarctica. Cf. Scolecitrichidae (s.l.)
Neoscolecithrix bidentata    Farran, 1905   (F)
Syn.: Oothrix bidentata Farran, 1905 (p.42, Descr.F, figs.F); Sewell, 1948 (p.501); ? Chen & Zhang, 1965 (p.58, figs.F); Shih & Young, 1995 (p.71)
Ref.: Bradford & al., 1983 (p.123); Hulsemann, 1985 (p.55, 60, Rem.)
Rem.: Cf. Neoscolecithrix koehleri. Cf. Scolecitrichidae (s.l.)
Neoscolecithrix caetanoi    Alvarez, 1985   (F,M)
Syn.: Cenognatha caetanoi : Bradford-Grieve, 2004 (p.285)
Ref.: Alvarez, 1985 a (p.198, Descr.F,M, figs.F,M); Bradford-Grieve & al., 1999 (p.880, 928, 929, figs.F,M, comme caetanovi)
Rem.: Cf. Scolecitrichidae (s.l.)
Neoscolecithrix farrani    Smirnov, 1935   (F,M)
Syn.: Oothrix bidentata Wirketis, 1926; Bogorov,1932; Oothrix borealis Wiborg, 1949 (p.2, figs.F,M); Cenognatha farrani : Bradford-Grieve, 2004 (p.285)
Ref.: Smirnov, 1935 a (p.259, Descr.F, figs.F); Sewell, 1948 (p.496); Fosshagen, 1972 a (p.5, figs.F,M, Rem.); Prygunkova, 1972 (p.251, Descr. M); Shih & Stallard, 1982 (p.59, figs.F, Rem.); Bradford & al., 1983 (p.123); Hulsemann, 1985 (p.60, 61, Rem.)
Rem.: Cf. Scolecitrichidae (s.l.)
Neoscolecithrix koehleri    Canu, 1896   (F,M)
Syn.: Scolecithrix koehleri : Giesbrecht & Schmeil, 1898 (p.47, Rem.F,M); Oothrix bidentata Farran, 1905 (p.42, figs.F, ?Juv); Pearson, 1906 (p.21); Rose, 1933 a (p.140, figs.F); Brodsky & al., 1985 (p.125, 126); ? non Chen & Zhang, 1965 (p.58, figs.F); Shih & Young, 1995 (p.71)
Ref.: Canu, 1896 (p.426, figs.F,M); Bradford & al., 1983 (p.123); Hulsemann, 1985 (p.60, 61, Rem.F)
Rem.: Cf. Scolecitrichidae (s.l.)
Neoscolecithrix magna    Grice, 1972   (F)
Syn.: Oothrix magna Grice, 1972 a (p.228, figs.F); ? Neoscolecithrix sp Bradford, 1973 (p.147); Bradford-Grieve, 2001 a (p.782, figs.F)
Ref.: Hulsemann, 1985 (p.60, 61, Rem.); Bradford & al., 1983 (p.123)
Rem.: Cf. Scolecitrichidae (s.l.)
Neoscolecithrix ornata    Bradford-Grieve, 2001   (F)
Ref.: Bradford-Grieve, 2001 a (p.887, Descr.F, figs.F)
Rem.: Cf. Scolecitrichidae (s.l.)
Neoscolecithrix watersae    Grice, 1972   (F,M)
Syn.: Xanthocalanus watersae Grice, 1972 a (p.233, figs.F,M)
Ref.: Campaner, 1978 a (p.968, Rem.); Bradford & al., 1983 (p.123); Hulsemann, 1985 (p.60, 61, Rem.)
Rem.: Cf. Scolecitrichidae (s.l.)
Neoscolecithrix sp.    Bradford, 1973   (F, Juv.5 F)
Ref.: Bradford, 1973 (p.147, figs.Fjuv.V); Bradford & al., 1983 (p.124, figs. Juv.5); Hulsemann, 1985 (p.60, 61, Rem.); Bradford-Grieve, 2001 a (p.782, figs.F, Rem.)
Rem.: Cf. Scolecitrichidae (s.l.)
Neoscolecithrix sp.    Chahsavar-Archad & Razouls, 1982   (Juv.F)
Ref.: Chahsavar-Archad & Razouls, 1982 (p.27, fig.Fjuv.); Bradford-Grieve, 2001 a
Rem.: Cf. Scolecitrichidae (s.l.)

Paratharybis Tanaka, 1937
Rem.: Cf. Undinella
Paratharybis frontalis    Tanaka, 1937   (F)
Ref.: Tanaka, 1937 (p.264); Vervoort, 1957 (p.96, Rem.)
Loc: Japon (Suruga Bay)
Rem.: Cf. Undinella frontalis
(1) Parundinella Fleminger, 1957
Rem.: Total 4 spp.

Diagnosis after Bradford, Haakonssen & Jillett (1983, p.124), after Fleminger 1957 a :
- Head and pediger segment 1 fused, 4th and 5th pediger segments fused.
- Posterolateral corners of metasome produced into lobiform or spiniform process.
- Genital segment symmetrical, swollen laterally, ventrally only weakly produced.
- Rostrum simple, bifurcate, with slender filament on each side.
- A1 23-segmented in female, 21-segmented in male.
- Md blade elongate with 3 to 4 monocuspid, spiniform teeth confined to dorsal-most portion of lobe.
- Mx1 inner lobe 1 moderately sized, not broad or truncate.
Mx2 lobe 5 with 2 sensory filaments, 1 normal seta and 1 enlarged falcate seta; terminal part of endopod with 5 or 6 sensory filaments, some with apical flagellum.
- External exopod spines of P2-P4 ornamented with small denticles along upper and lower margins.
- Female P5 minute, 1-segmented, attached to common basal segment; distal segment terminated by 1 or more pointed processes and 1 robust inner edge spine bearing ventral spinules.
Male P5 biramous, asymmetrical; left leg exopod 3-segmented, segment 1 with inner proximal process, endopod 1- or 2-segmented; right leg exopod 2-segmented, endopod in part fused with basipod segment 2.


For Markhaseva, Laakmann & Renz (2014, p.84) this genus must be placed outside Tharybidae, his status remains taxonomically unresolved. Fleminger's description is not detailed enough to clarify the position of Parundinella within the ''Bradfordians''. Several characters exclude this genus apart from the Tharybidae: 1 - A2 endopodal segment 1 with 1 seta (versus 2 setae in Tharybis and exopod armament as 0, 0-1-1-1, 1, 1, 1, 0, 3 (versus 0, 0-0-0, 1, 1, 1, 1, 1, 3 in Tharybis); 2 - Mx1 praecoxal arthrite with 10-12 setal elements (versus 13-14 in Tharybis), exopod with 7 setae (versus 3-5 setae in Tharybis); 3 - Mx2 endopod with 5-6 setal elements (versus 8-9 setae in Tharybis); 4 - Mxp praecoxal lobes with a setal formula as 1, 2 and 2? (versus 1, 2 and 3 setae in Tharybis) (see Table 1-5); 5 - male P5, right and left coxa and basis of nearly same length (versus right and left basipods significantly asymmetrical in Tharybis), P5 right leg biramous (versus uniramous in Tharybis).

[1] Parundinella dakini  Bradford, 1973   (F,M)    [Figs]

[2] Parundinella emarginata  Grice & Hulsemann, 1970   (F,M)    [Figs]

[3] Parundinella manicula  Fleminger, 1957   (F,M)    [Figs]

[4] Parundinella spinodenticula  Fleminger, 1957   (F,M)
Pseudophaenna Sars, 1902
Rem.: Boxshall & Halsey (2004, p.96) transfer this genus to the Diaixidae family.

(*) Rythabis Schulz, 1995
Rem.: Cf. Scolecitrichidae
Rythabis atlantica    Schulz, 1995   (F)
Ref.: in Schulz & Beckmann, 1995 (p.202, Descr.F, figs.F)
Rem.: Cf. Scolecitrichidae (lato sensu)
(2) Tharybis Sars, 1902
Rem.: type: Tharybis macrophthalma Sars, 1903. Total: 22 spp.

Diagnosis from Bradford, Haakonssen & Jillett (1983, p.126), after Schulz, 1981 :
- Head and pediger segment 1 fused or separate, pediger segments 4 and 5 fused.
- Rostrum short and rounded with or without 2 filaments.
- A1 24-segmented in female, 22-segmented on right and 23-segmented on left in male.
- A2 exopod more than twice as long as endopod.
- Md blade with 2 unusually strong bifid teeth followed by dense series of plumose setae; palp with both rami well developed.
- Mx1 with inner lobe 1 enlarged, extending beyond remaider of appendage; exopod reduced with 3 to 5 setae.
- Mx2 endopod with 3 (4?) worm-like and 6 (4,) brush-like sensory filaments; lobes extend over more than half appendage.
-Mxp basipod swollen;
- Posterior surface of swimming legs may bear small spinules,
- Terminal spine of P2-P4 coarsely serrated.
- Female P5 of two segments attached to common basal segment, terminal segment longest; terminal segment with 1 or 2 terminal spines as well as articulated inner edge spine.
- Male P5 asymmetrical, usually biramous on left, uniramous on right; left leg with 2 basal segments, exopod 3-segmented, shorter than the one-segmented endopod, segment 3 inserted terminally; right leg 3-segmented, segment 2 with inner edge distal process.

Remarks: Tharybis and Undinella are difficult to separate when all their species are taken into account.

[1] Tharybis altera  (Grice & Hulsemann, 1970)   (M)    [Figs]
Rem.: hyperbenthic. (1750-1822 m).
The change of the genus does not seem to be absolutely assured.

[2] Tharybis angularis  Schulz, 1995   (F)    [Figs]

[3] Tharybis asymmetrica  Andronov, 1976   (F)    [Figs]

[4] Tharybis compacta  (Grice & Hulsemann, 1970)   (F,M)    [Figs]
Rem.: hyperbenthic. (1750-1822 m).
The change of the genus does not seem to be absolutely assured.Undinella compacta approaches the genus Tharybis in the relatively large 1st inner lobe (= arthrite) of Mx1, the swollen 2nd basipodal segment of Mxp, and the elongate terminal segment of P5. However, this present species is placed in the genus Undinella because the Head and 1st thoracic segment and4th and 5th thoracic segments partially separatd, the exopod of Mx1 with 3 setae, and the lobes of Mx2 crowded within the distal half of the appendage. The absence of a rostrum and rostral filaments is unique in the family Tharybidae.

Tharybis crenata    Schulz, 1995   (F)
Ref.: in Schulz & Beckmann, 1995 (p.207, figs.F); Andronov, 2002 (p.37); Ferrari & Markhaseva, 2005 (Rem.: p.33; p.46: tab.1)
Loc: Islande S
N: 1
Lg.: (594) F: 1,2-1,13; {F: 1,13-1,20}
Rem.: Cf. Tharybis groenlandica

[5] Tharybis fultoni  Park, 1967   (F,M)    [Figs]

[6] Tharybis groenlandicus  (Tupitzky, 1982)   (F,M)    [Figs]

[7] Tharybis inaequalis  Bradford-Grieve, 2001   (F,M)    [Figs]

[8] Tharybis inflata  Andronov, 2002   (F,M)    [Figs]

[9] Tharybis juhlae  Ferrari & Markhaseva, 2005   (F,M)    [Figs]

[10] Tharybis lauta  Andronov, 2002   (F,M)    [Figs]

Tharybis macrocephalon    Grice & Hulsemann,1970   (F)
Rem.: Cf. Byrathis macrocephalon

[11] Tharybis macrophthalma  Sars, 1902   (F,M)    [Figs]

[12] Tharybis macrophthalmoida  Andronov, 2002   (F,M)    [Figs]

[13] Tharybis magna  Bradford & Wells, 1983   (F,M)    [Figs]

[14] Tharybis megalodactyla  Andronov, 1976   (F,M)    [Figs]

Tharybis minor    Schulz, 1981   (F,M)
Ref.: Schulz, 1981 (p.171, figs.F,M); Bradford & al., 1983 (p.126); Andronov, 2002 (p.34); Ferrari & Markhaseva, 2005 (p.46: tab.1; Rem.)
Loc: Mauritanie (off Cap Blanc N)
N: 1
Lg.: (597) F: 0,96-0,88; M: 0,91-0,83; {F: 0,88-0,96; M: 0,83-0,91}
Rem.: Cf. Tharybis asymmetrica

[15] Tharybis neptuni  (Cleve, 1904)   (F,M)    [Figs]

[16] Tharybis paraincertus  (Grice & Hulsemann, 1965)   (F)    [Figs]

[17] Tharybis pseudomegalodactyla  Ferrari & Markhaseva, 2005   (F,M)    [Figs]

[18] Tharybis sagamiensis  Tanaka, 1960   (F)    [Figs]

[19] Tharybis scaura  Andronov, 2002   (F,M)    [Figs]

[20] Tharybis shuheiella  Ferrari & Markhaseva, 2005   (F)    [Figs]

[21] Tharybis tuberosa  Andronov, 2002   (F,M)    [Figs]

[22] Tharybis tumidula  Andronov, 2002   (F)    [Figs]

Tharybis sp.       
Syn.: Stephus neptuni : Wolfenden, 1911 (p.206, Descr.F, figs.F)
Loc: Antarct. (continent)
Lg.: (10) F: 1,8-1,75; {F: 1,75-1,80}
Rem.: Wolfenden émet des doutes concernant l’attribution de sa forme à l’espèce décrite par Cleve comme Scolcithricella neptuni (1904 a, p.206)
(3) Undinella Sars, 1900
Rem.: type: Undinella oblonga Sars,1900. Total: 9 spp. + 1 undet.

Diagnosis from Bradford, Haakonssen & Jillett (1983, p.127), after Sars, 1900 and Fleminger, 1957 :
- Head and pediger segment 1 partly separate, pediger segments 4 and 5 fused or separate.
- Posterior corners of metasome prolonged posteriorly in female.
- Rostrum with basal portion bifid, both parts terminated by 1 filament.
- A1 24-segmented in female; 23-segmented in male.
- A2 exopod more than twice as long as endopod.
- Md blade with 2 strong bidentate teeth; palp with endopod larger than exopod.
- Mx1 inner lobe 1 moderately enlarged, not extending beyond endopod; exopod reduced, with 2 or 3 setae.
- Mx2 endopod with sensory filaments; lobes crowded on distal half of appendage.
- Mxp slender.
- Posterior surfaces of swimming legs may have some small spinules.
- Terminal spine of P2-P4 with fine shallow serrations.
- Female P5 symmetrical or asymmetrical, 2-segmented, attached to common basal segment; segments of approximately equal length; distal segment with 2 to 4 short terminal spines and sometimes outer edge spine.
- Male P5 asymmetrical; uniramous on right, 3-segmented, segment 2 usually with inner edge distal process; biramous on left, with on, endopod 1-segmented, longer than 3-segmented exopod, exopod segment 3 inserted subterminally on segment 2 to make pincers. basal segment attached to common basal segment.

Remarks: Tharybis and Undinella are difficult to separate when all their species are taken into account.

[1] Undinella aculeata  Andronov, 2002   (F,M)    [Figs]

[2] Undinella acuta  Vaupel-Klein, 1970   (F)    [Figs]

Undinella altera    Grice & Hulsemann, 1970   (M)
Ref.: Grice & Hulsemann, 1970 (p.194); Schulz, 1981 (p.169, 171, Rem.); Bradford & al., 1983 (p.127)
Rem.: Probable dans le genre Tharybis. Cf. Tharybis altera

Undinella brevipes    Farra, 1908   (F,M)
Syn.: Xanthocalanus simplex Wolfenden, 1906 (p.30, figs.F); Undinella simplex : Farran, 1926 (p.268); Rose, 1933 a (p.142, figs.F); Vives & al., 1975 (p.43, tab.II); Deevey & Brooks, 1977 (tab.2); Vives, 1982 (p.292); Greze & al., 1985 (p.7); Undinella simplex (M) : Grice & Hulsemann, 1965 (p.224, 242, figs.M)
Ref.: Farran, 1908 b (p.50, Descr.F, figs.F); Lysholm & Nordgaard, 1921 (p.19); Lysholm & al., 1945 (p.29) Vervoort, 1957 (p.95, Rem.); Tanaka, 1960 a (p.131, figs. juv.M); Minoda, 1971 (p.33, figs.F); Bradford & al., 1983 (p.127, figs.F,M, Rem.); Boxshall & Halsey, 2004 (p.211, fig.F)
Rem.: Cf. Undinelle simplex.

Undinella compacta    Grice & Hulsemann, 1970   (F,M)
Ref.: Grice & Hulsemann, 1970 (p.195); Schulz, 1981 (p.171, Rem.)
Rem.: Cf. Tharybis compacta

[3] Undinella frontalis  (Tanaka, 1937)   (F,M)    [Figs]

[4] Undinella gricei  Wheeler, 1970   (F,M)    [Figs]

[5] Undinella hampsoni  Grice & Hulsemann, 1970   (F,M)    [Figs]

[6] Undinella oblonga  Sars, 1900   (F,M)    [Figs]

[7] Undinella simplex  (Wolfenden, 1906)   (F,M)    [Figs]

[8] Undinella spinifera  Tanaka, 1960   (F)    [Figs]

[9] Undinella stirni  Grice, 1971   (F,M)    [Figs]

[10] Undinella sp.1  Roe, 1975   (F)    [Figs]
Rem.: Probably in this genus.
(4) incertae sedis (Tharybidae)
Rem.: The following species would belong to this family:
[1] "Amallothrix" robustipes Grice & Hulsemann,1965 (F)
Syn.: ? Xanthocalanus paululus Park, 1970 (p.497); Roe, 1975 (p.364)
Ref.: Grice & Hulsemann, 1965 (p.239, Descr.F, figs.F); 1967 (p.16); Bradford, 1973 (p.147, Rem.); Roe, 1975 (p.365, Rem.); Bradford & al., 1983 (p.123, Rem.)
Loc.: Atlant. NE (off Canaries-Açores-), ? Is. du Cap Vert, ? G. du Mexique, Indien W
N: 2
Lg.: (226) F: 1,13; {F: 1,13}

Xanthocalanus groenlandicus Tupitzky,1982 (F,M)
Ref.: Tupitzky, 1982 (p.297, figs.F,M); Kosobokova & al., 1955 (p.194)
Loc.: Arct., mer de Laptev
Lg.: (453) F: 1,5-1,4; M: 1,4; {F: 1,40-1,50; M: 1,40}
Rem.: Campaner, 1983 (comm. pers.): Cf. ? Tharybis. Voir spp. inc. sedis

[2] "Xanthocalanus" hispidus Grice & Hulsemann, 1967 (F)  [Figs]  details

Xanthocalanus macrocephalon Grice & Hulsemann,1970 (F)
Ref.: Grice & Hulsemann, 1970 (p.191, Descr.F, figs.F); Bradford, 1973 (p.139, 147, Rem.); Bradford & al., 1983 (p.127); Schulz & Beckmann, 1995 (p.210, Rem.)
Loc.: off Woods Hole
N: 1
Lg.: (200) F: 1,12-1,06; {F: 1,06-1,12}
Rem.: Cf. Tharybis

Xanthocalanus paraincertus Grice & Hulsemann,1965 (F)
Ref.: Grice & Hulsemann, 1965 (part., p.235, Descr.F, figs.F); Bradford, 1973 (p.139, 147, Rem.); Bradford & al., 1983 (p.127, Rem.); Schulz & Beckmann, 1995 (p.210, Rem.)
Loc.: Atlant. NE (locality not specified)
N: 1
Lg.: (226) F: 1,61-1,15; {F: 1,15-1,61} Rem.: abyssopelagic. Cf. Tharybis

“Xanthocalanus” paululus Park,1970 (F,M)
Ref.: Park, 1970 (p.497, Descr.F, figs.F); Bradford, 1973 (p.139, 147, Rem.); Roe, 1975 (p.364, Descr.F,M, figs.F,M, Rem.); Bradford, 1983 (p.127, Rem.)
Ref. compl.: Suarez-Morales & Gasca, 1998 a (p.110)
Loc.: Is. du Cap Vert, Mer des Caraïbes, G. du Mexique
N: 3
Lg.: (8) F: 1,29-1,14; M: 1,37-1,29; (88) F: 1,28-1,22; {F: 1,14-1,29; M: 1,29-1,37}
Rem.: No “Xanthocalanus” hispidus. ? Cf. Undinella. Cf.: Brodskius paululus

[1] Amallothrix robustipes  Grice & Hulsemann, 1965   (F)    [Figs]

Xanthocalanus groenlandicus    Tupitzky, 1982   (F,M)
Ref.: Tupitzky, 1982 (p.297, figs.F,M); Kosobokova & al., 1955 (p.194)
Loc: Arct., mer de Laptev
Lg.: (453) F: 1,5-1,4; M: 1,4; {F: 1,40-1,50; M: 1,40}
Rem.: Campaner, 1983 (comm. pers.): Cf. ? Tharybis. Voir spp. inc. sedis

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