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List species and varieties by family |
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| Tharybidae Sars, 1902 ( Clausocalanoidea ) | |
| | | (*) Cenognatha Bradford-Grieve, 2001 | |
| | Rem.: | type: Neoscolecithrix antarctica. 1 sp. (provisionally). Bradford-Grieve, 2004 (p.285) includes in this genus the two species Neoscolecithrix caetanoi and N. farrani. Boxshall & Halsey, 2004 (p.186). Cf. Scolecitrichidae (largo sensu). | | | | Cenognatha antarctica Hulsemann, 1985 (F, Juv.M) | | Syn.: | Neoscolecithrix antarctica Hulsemann, 1985 (p.55, Descr., figs. C5, F & M); Razouls & al., 2000 (p.343, tab. 2, Appendix) | | Ref.: | Bradford-Grieve, 2001 a (p.792, Rem.); Bradford-Grieve, 2004 (p.285) | | Rem.: | Pour Vyshkvartzeva (2000, p.217) cette espèce appartiendrait aux Scolecitrichidae. Bradford-Grieve, 2004 (p.285) |
| | (*) Neoscolecithrix Canu, 1896 (part.) | |
| | Rem.: | type: Neoscolecithrix koehleri Canu,1896. Ohtsuka, Boxshall & Fosshagen (2003, p.61-62) believe that this genus belongs (largo sensu) to Scolecitrichidae. | | | | Neoscolecithrix antarctica Hulsemann, 1985 (F, juv.M) | | Ref.: | Hulsemann, 1985 (p.55, Descr., figs. C5, F & M) | | Rem.: | Cf. Cenognatha antarctica. Cf. Scolecitrichidae (s.l.) | | | | Neoscolecithrix bidentata Farran, 1905 (F) | | Syn.: | Oothrix bidentata Farran, 1905 (p.42, Descr.F, figs.F); Sewell, 1948 (p.501); ? Chen & Zhang, 1965 (p.58, figs.F); Shih & Young, 1995 (p.71) | | Ref.: | Bradford & al., 1983 (p.123); Hulsemann, 1985 (p.55, 60, Rem.) | | Rem.: | Cf. Neoscolecithrix koehleri. Cf. Scolecitrichidae (s.l.) | | | | Neoscolecithrix caetanoi Alvarez, 1985 (F,M) | | Syn.: | Cenognatha caetanoi : Bradford-Grieve, 2004 (p.285) | | Ref.: | Alvarez, 1985 a (p.198, Descr.F,M, figs.F,M); Bradford-Grieve & al., 1999 (p.880, 928, 929, figs.F,M, comme caetanovi) | | Rem.: | Cf. Scolecitrichidae (s.l.) | | | | Neoscolecithrix farrani Smirnov, 1935 (F,M) | | Syn.: | Oothrix bidentata Wirketis, 1926; Bogorov,1932; Oothrix borealis Wiborg, 1949 (p.2, figs.F,M); Cenognatha farrani : Bradford-Grieve, 2004 (p.285) | | Ref.: | Smirnov, 1935 a (p.259, Descr.F, figs.F); Sewell, 1948 (p.496); Fosshagen, 1972 a (p.5, figs.F,M, Rem.); Prygunkova, 1972 (p.251, Descr. M); Shih & Stallard, 1982 (p.59, figs.F, Rem.); Bradford & al., 1983 (p.123); Hulsemann, 1985 (p.60, 61, Rem.) | | Rem.: | Cf. Scolecitrichidae (s.l.) | | | | Neoscolecithrix koehleri Canu, 1896 (F,M) | | Syn.: | Scolecithrix koehleri : Giesbrecht & Schmeil, 1898 (p.47, Rem.F,M); Oothrix bidentata Farran, 1905 (p.42, figs.F, ?Juv); Pearson, 1906 (p.21); Rose, 1933 a (p.140, figs.F); Brodsky & al., 1985 (p.125, 126); ? non Chen & Zhang, 1965 (p.58, figs.F); Shih & Young, 1995 (p.71) | | Ref.: | Canu, 1896 (p.426, figs.F,M); Bradford & al., 1983 (p.123); Hulsemann, 1985 (p.60, 61, Rem.F) | | Rem.: | Cf. Scolecitrichidae (s.l.) | | | | Neoscolecithrix magna Grice, 1972 (F) | | Syn.: | Oothrix magna Grice, 1972 a (p.228, figs.F); ? Neoscolecithrix sp Bradford, 1973 (p.147); Bradford-Grieve, 2001 a (p.782, figs.F) | | Ref.: | Hulsemann, 1985 (p.60, 61, Rem.); Bradford & al., 1983 (p.123) | | Rem.: | Cf. Scolecitrichidae (s.l.) | | | | Neoscolecithrix ornata Bradford-Grieve, 2001 (F) | | Ref.: | Bradford-Grieve, 2001 a (p.887, Descr.F, figs.F) | | Rem.: | Cf. Scolecitrichidae (s.l.) | | | | Neoscolecithrix watersae Grice, 1972 (F,M) | | Syn.: | Xanthocalanus watersae Grice, 1972 a (p.233, figs.F,M) | | Ref.: | Campaner, 1978 a (p.968, Rem.); Bradford & al., 1983 (p.123); Hulsemann, 1985 (p.60, 61, Rem.) | | Rem.: | Cf. Scolecitrichidae (s.l.) | | | | Neoscolecithrix sp. Bradford, 1973 (F, Juv.5 F) | | Ref.: | Bradford, 1973 (p.147, figs.Fjuv.V); Bradford & al., 1983 (p.124, figs. Juv.5); Hulsemann, 1985 (p.60, 61, Rem.); Bradford-Grieve, 2001 a (p.782, figs.F, Rem.) | | Rem.: | Cf. Scolecitrichidae (s.l.) | | | | Neoscolecithrix sp. Chahsavar-Archad & Razouls, 1982 (Juv.F) | | Ref.: | Chahsavar-Archad & Razouls, 1982 (p.27, fig.Fjuv.); Bradford-Grieve, 2001 a | | Rem.: | Cf. Scolecitrichidae (s.l.) |
| | Paratharybis Tanaka, 1937 | |
| | Rem.: | Cf. Undinella | | | | Paratharybis frontalis Tanaka, 1937 (F) | | Ref.: | Tanaka, 1937 (p.264); Vervoort, 1957 (p.96, Rem.) | | Loc: | Japon (Suruga Bay) | | Rem.: | Cf. Undinella frontalis | | | | (1) Parundinella Fleminger, 1957 | |
| | Rem.: | Total 4 spp.
Diagnosis after Bradford, Haakonssen & Jillett (1983, p.124), after Fleminger 1957 a :
- Head and pediger segment 1 fused, 4th and 5th pediger segments fused.
- Posterolateral corners of metasome produced into lobiform or spiniform process.
- Genital segment symmetrical, swollen laterally, ventrally only weakly produced.
- Rostrum simple, bifurcate, with slender filament on each side.
- A1 23-segmented in female, 21-segmented in male.
- Md blade elongate with 3 to 4 monocuspid, spiniform teeth confined to dorsal-most portion of lobe.
- Mx1 inner lobe 1 moderately sized, not broad or truncate.
Mx2 lobe 5 with 2 sensory filaments, 1 normal seta and 1 enlarged falcate seta; terminal part of endopod with 5 or 6 sensory filaments, some with apical flagellum.
- External exopod spines of P2-P4 ornamented with small denticles along upper and lower margins.
- Female P5 minute, 1-segmented, attached to common basal segment; distal segment terminated by 1 or more pointed processes and 1 robust inner edge spine bearing ventral spinules.
Male P5 biramous, asymmetrical; left leg exopod 3-segmented, segment 1 with inner proximal process, endopod 1- or 2-segmented; right leg exopod 2-segmented, endopod in part fused with basipod segment 2.
For Markhaseva, Laakmann & Renz (2014, p.84) this genus must be placed outside Tharybidae, his status remains taxonomically unresolved. Fleminger's description is not detailed enough to clarify the position of Parundinella within the ''Bradfordians''. Several characters exclude this genus apart from the Tharybidae: 1 - A2 endopodal segment 1 with 1 seta (versus 2 setae in Tharybis and exopod armament as 0, 0-1-1-1, 1, 1, 1, 0, 3 (versus 0, 0-0-0, 1, 1, 1, 1, 1, 3 in Tharybis); 2 - Mx1 praecoxal arthrite with 10-12 setal elements (versus 13-14 in Tharybis), exopod with 7 setae (versus 3-5 setae in Tharybis); 3 - Mx2 endopod with 5-6 setal elements (versus 8-9 setae in Tharybis); 4 - Mxp praecoxal lobes with a setal formula as 1, 2 and 2? (versus 1, 2 and 3 setae in Tharybis) (see Table 1-5); 5 - male P5, right and left coxa and basis of nearly same length (versus right and left basipods significantly asymmetrical in Tharybis), P5 right leg biramous (versus uniramous in Tharybis). |
| | [1] Parundinella dakini Bradford, 1973 (F,M) [Figs] | |
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| | [2] Parundinella emarginata Grice & Hulsemann, 1970 (F,M) [Figs] | |
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| | [3] Parundinella manicula Fleminger, 1957 (F,M) [Figs] | |
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| | [4] Parundinella spinodenticula Fleminger, 1957 (F,M) | |
| | | | Rem.: | Boxshall & Halsey (2004, p.96) transfer this genus to the Diaixidae family. |
| | (*) Rythabis Schulz, 1995 | |
| | Rem.: | Cf. Scolecitrichidae | | | | Rythabis atlantica Schulz, 1995 (F) | | Ref.: | in Schulz & Beckmann, 1995 (p.202, Descr.F, figs.F) | | Rem.: | Cf. Scolecitrichidae (lato sensu) | | | | | Rem.: | type: Tharybis macrophthalma Sars, 1903. Total: 22 spp.
Diagnosis from Bradford, Haakonssen & Jillett (1983, p.126), after Schulz, 1981 :
- Head and pediger segment 1 fused or separate, pediger segments 4 and 5 fused.
- Rostrum short and rounded with or without 2 filaments.
- A1 24-segmented in female, 22-segmented on right and 23-segmented on left in male.
- A2 exopod more than twice as long as endopod.
- Md blade with 2 unusually strong bifid teeth followed by dense series of plumose setae; palp with both rami well developed.
- Mx1 with inner lobe 1 enlarged, extending beyond remaider of appendage; exopod reduced with 3 to 5 setae.
- Mx2 endopod with 3 (4?) worm-like and 6 (4,) brush-like sensory filaments; lobes extend over more than half appendage.
-Mxp basipod swollen;
- Posterior surface of swimming legs may bear small spinules,
- Terminal spine of P2-P4 coarsely serrated.
- Female P5 of two segments attached to common basal segment, terminal segment longest; terminal segment with 1 or 2 terminal spines as well as articulated inner edge spine.
- Male P5 asymmetrical, usually biramous on left, uniramous on right; left leg with 2 basal segments, exopod 3-segmented, shorter than the one-segmented endopod, segment 3 inserted terminally; right leg 3-segmented, segment 2 with inner edge distal process.
Remarks: Tharybis and Undinella are difficult to separate when all their species are taken into account. |
| | [1] Tharybis altera (Grice & Hulsemann, 1970) (M) [Figs] | |
| | Rem.: | hyperbenthic. (1750-1822 m).
The change of the genus does not seem to be absolutely assured. |
| | [2] Tharybis angularis Schulz, 1995 (F) [Figs] | |
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| | [3] Tharybis asymmetrica Andronov, 1976 (F) [Figs] | |
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| | [4] Tharybis compacta (Grice & Hulsemann, 1970) (F,M) [Figs] | |
| | Rem.: | hyperbenthic. (1750-1822 m).
The change of the genus does not seem to be absolutely assured.Undinella compacta approaches the genus Tharybis in the relatively large 1st inner lobe (= arthrite) of Mx1, the swollen 2nd basipodal segment of Mxp, and the elongate terminal segment of P5. However, this present species is placed in the genus Undinella because the Head and 1st thoracic segment and4th and 5th thoracic segments partially separatd, the exopod of Mx1 with 3 setae, and the lobes of Mx2 crowded within the distal half of the appendage. The absence of a rostrum and rostral filaments is unique in the family Tharybidae. |
| | Tharybis crenata Schulz, 1995 (F) | | Ref.: | in Schulz & Beckmann, 1995 (p.207, figs.F); Andronov, 2002 (p.37); Ferrari & Markhaseva, 2005 (Rem.: p.33; p.46: tab.1) | | Loc: | Islande S | | N: | 1 | | Lg.: | (594) F: 1,2-1,13; {F: 1,13-1,20} | | Rem.: | Cf. Tharybis groenlandica |
| | [5] Tharybis fultoni Park, 1967 (F,M) [Figs] | |
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| | [6] Tharybis groenlandicus (Tupitzky, 1982) (F,M) [Figs] | |
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| | [7] Tharybis inaequalis Bradford-Grieve, 2001 (F,M) [Figs] | |
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| | [8] Tharybis inflata Andronov, 2002 (F,M) [Figs] | |
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| | [9] Tharybis juhlae Ferrari & Markhaseva, 2005 (F,M) [Figs] | |
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| | [10] Tharybis lauta Andronov, 2002 (F,M) [Figs] | |
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| | Tharybis macrocephalon Grice & Hulsemann,1970 (F) | | Rem.: | Cf. Byrathis macrocephalon |
| | [11] Tharybis macrophthalma Sars, 1902 (F,M) [Figs] | |
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| | [12] Tharybis macrophthalmoida Andronov, 2002 (F,M) [Figs] | |
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| | [13] Tharybis magna Bradford & Wells, 1983 (F,M) [Figs] | |
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| | [14] Tharybis megalodactyla Andronov, 1976 (F,M) [Figs] | |
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| | Tharybis minor Schulz, 1981 (F,M) | | Ref.: | Schulz, 1981 (p.171, figs.F,M); Bradford & al., 1983 (p.126); Andronov, 2002 (p.34); Ferrari & Markhaseva, 2005 (p.46: tab.1; Rem.) | | Loc: | Mauritanie (off Cap Blanc N) | | N: | 1 | | Lg.: | (597) F: 0,96-0,88; M: 0,91-0,83; {F: 0,88-0,96; M: 0,83-0,91} | | Rem.: | Cf. Tharybis asymmetrica |
| | [15] Tharybis neptuni (Cleve, 1904) (F,M) [Figs] | |
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| | [16] Tharybis paraincertus (Grice & Hulsemann, 1965) (F) [Figs] | |
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| | [17] Tharybis pseudomegalodactyla Ferrari & Markhaseva, 2005 (F,M) [Figs] | |
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| | [18] Tharybis sagamiensis Tanaka, 1960 (F) [Figs] | |
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| | [19] Tharybis scaura Andronov, 2002 (F,M) [Figs] | |
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| | [20] Tharybis shuheiella Ferrari & Markhaseva, 2005 (F) [Figs] | |
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| | [21] Tharybis tuberosa Andronov, 2002 (F,M) [Figs] | |
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| | [22] Tharybis tumidula Andronov, 2002 (F) [Figs] | |
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| | Tharybis sp. | | Syn.: | Stephus neptuni : Wolfenden, 1911 (p.206, Descr.F, figs.F) | | Loc: | Antarct. (continent) | | Lg.: | (10) F: 1,8-1,75; {F: 1,75-1,80} | | Rem.: | Wolfenden émet des doutes concernant l’attribution de sa forme à l’espèce décrite par Cleve comme Scolcithricella neptuni (1904 a, p.206) | | | | Rem.: | type: Undinella oblonga Sars,1900. Total: 9 spp. + 1 undet.
Diagnosis from Bradford, Haakonssen & Jillett (1983, p.127), after Sars, 1900 and Fleminger, 1957 :
- Head and pediger segment 1 partly separate, pediger segments 4 and 5 fused or separate.
- Posterior corners of metasome prolonged posteriorly in female.
- Rostrum with basal portion bifid, both parts terminated by 1 filament.
- A1 24-segmented in female; 23-segmented in male.
- A2 exopod more than twice as long as endopod.
- Md blade with 2 strong bidentate teeth; palp with endopod larger than exopod.
- Mx1 inner lobe 1 moderately enlarged, not extending beyond endopod; exopod reduced, with 2 or 3 setae.
- Mx2 endopod with sensory filaments; lobes crowded on distal half of appendage.
- Mxp slender.
- Posterior surfaces of swimming legs may have some small spinules.
- Terminal spine of P2-P4 with fine shallow serrations.
- Female P5 symmetrical or asymmetrical, 2-segmented, attached to common basal segment; segments of approximately equal length; distal segment with 2 to 4 short terminal spines and sometimes outer edge spine.
- Male P5 asymmetrical; uniramous on right, 3-segmented, segment 2 usually with inner edge distal process; biramous on left, with on, endopod 1-segmented, longer than 3-segmented exopod, exopod segment 3 inserted subterminally on segment 2 to make pincers. basal segment attached to common basal segment.
Remarks: Tharybis and Undinella are difficult to separate when all their species are taken into account. |
| | [1] Undinella aculeata Andronov, 2002 (F,M) [Figs] | |
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| | [2] Undinella acuta Vaupel-Klein, 1970 (F) [Figs] | |
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| | Undinella altera Grice & Hulsemann, 1970 (M) | | Ref.: | Grice & Hulsemann, 1970 (p.194); Schulz, 1981 (p.169, 171, Rem.); Bradford & al., 1983 (p.127) | | Rem.: | Probable dans le genre Tharybis. Cf. Tharybis altera |
| | Undinella brevipes Farra, 1908 (F,M) | | Syn.: | Xanthocalanus simplex Wolfenden, 1906 (p.30, figs.F); Undinella simplex : Farran, 1926 (p.268); Rose, 1933 a (p.142, figs.F); Vives & al., 1975 (p.43, tab.II); Deevey & Brooks, 1977 (tab.2); Vives, 1982 (p.292); Greze & al., 1985 (p.7); Undinella simplex (M) : Grice & Hulsemann, 1965 (p.224, 242, figs.M) | | Ref.: | Farran, 1908 b (p.50, Descr.F, figs.F); Lysholm & Nordgaard, 1921 (p.19); Lysholm & al., 1945 (p.29) Vervoort, 1957 (p.95, Rem.); Tanaka, 1960 a (p.131, figs. juv.M); Minoda, 1971 (p.33, figs.F); Bradford & al., 1983 (p.127, figs.F,M, Rem.); Boxshall & Halsey, 2004 (p.211, fig.F) | | Rem.: | Cf. Undinelle simplex. |
| | Undinella compacta Grice & Hulsemann, 1970 (F,M) | | Ref.: | Grice & Hulsemann, 1970 (p.195); Schulz, 1981 (p.171, Rem.) | | Rem.: | Cf. Tharybis compacta |
| | [3] Undinella frontalis (Tanaka, 1937) (F,M) [Figs] | |
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| | [4] Undinella gricei Wheeler, 1970 (F,M) [Figs] | |
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| | [5] Undinella hampsoni Grice & Hulsemann, 1970 (F,M) [Figs] | |
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| | [6] Undinella oblonga Sars, 1900 (F,M) [Figs] | |
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| | [7] Undinella simplex (Wolfenden, 1906) (F,M) [Figs] | |
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| | [8] Undinella spinifera Tanaka, 1960 (F) [Figs] | |
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| | [9] Undinella stirni Grice, 1971 (F,M) [Figs] | |
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| | [10] Undinella sp.1 Roe, 1975 (F) [Figs] | |
| | Rem.: | Probably in this genus. | | | (4) incertae sedis (Tharybidae) | |
| | Rem.: | The following species would belong to this family:
[1] "Amallothrix" robustipes Grice & Hulsemann,1965 (F)
Syn.: ? Xanthocalanus paululus Park, 1970 (p.497); Roe, 1975 (p.364)
Ref.: Grice & Hulsemann, 1965 (p.239, Descr.F, figs.F); 1967 (p.16); Bradford, 1973 (p.147, Rem.); Roe, 1975 (p.365, Rem.); Bradford & al., 1983 (p.123, Rem.)
Loc.: Atlant. NE (off Canaries-Açores-), ? Is. du Cap Vert, ? G. du Mexique, Indien W
N: 2
Lg.: (226) F: 1,13; {F: 1,13}
Xanthocalanus groenlandicus Tupitzky,1982 (F,M)
Ref.: Tupitzky, 1982 (p.297, figs.F,M); Kosobokova & al., 1955 (p.194)
Loc.: Arct., mer de Laptev
Lg.: (453) F: 1,5-1,4; M: 1,4; {F: 1,40-1,50; M: 1,40}
Rem.: Campaner, 1983 (comm. pers.): Cf. ? Tharybis. Voir spp. inc. sedis
[2] "Xanthocalanus" hispidus Grice & Hulsemann, 1967 (F) [Figs] details 
Xanthocalanus macrocephalon Grice & Hulsemann,1970 (F)
Ref.: Grice & Hulsemann, 1970 (p.191, Descr.F, figs.F); Bradford, 1973 (p.139, 147, Rem.); Bradford & al., 1983 (p.127); Schulz & Beckmann, 1995 (p.210, Rem.)
Loc.: off Woods Hole
N: 1
Lg.: (200) F: 1,12-1,06; {F: 1,06-1,12}
Rem.: Cf. Tharybis
Xanthocalanus paraincertus Grice & Hulsemann,1965 (F)
Ref.: Grice & Hulsemann, 1965 (part., p.235, Descr.F, figs.F); Bradford, 1973 (p.139, 147, Rem.); Bradford & al., 1983 (p.127, Rem.); Schulz & Beckmann, 1995 (p.210, Rem.)
Loc.: Atlant. NE (locality not specified)
N: 1
Lg.: (226) F: 1,61-1,15; {F: 1,15-1,61}
Rem.: abyssopelagic. Cf. Tharybis
“Xanthocalanus” paululus Park,1970 (F,M)
Ref.: Park, 1970 (p.497, Descr.F, figs.F); Bradford, 1973 (p.139, 147, Rem.); Roe, 1975 (p.364, Descr.F,M, figs.F,M, Rem.); Bradford, 1983 (p.127, Rem.)
Ref. compl.: Suarez-Morales & Gasca, 1998 a (p.110)
Loc.: Is. du Cap Vert, Mer des Caraïbes, G. du Mexique
N: 3
Lg.: (8) F: 1,29-1,14; M: 1,37-1,29; (88) F: 1,28-1,22; {F: 1,14-1,29; M: 1,29-1,37}
Rem.: No “Xanthocalanus” hispidus. ? Cf. Undinella. Cf.: Brodskius paululus
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| | [1] Amallothrix robustipes Grice & Hulsemann, 1965 (F) [Figs] | |
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| | Xanthocalanus groenlandicus Tupitzky, 1982 (F,M) | | Ref.: | Tupitzky, 1982 (p.297, figs.F,M); Kosobokova & al., 1955 (p.194) | | Loc: | Arct., mer de Laptev | | Lg.: | (453) F: 1,5-1,4; M: 1,4; {F: 1,40-1,50; M: 1,40} | | Rem.: | Campaner, 1983 (comm. pers.): Cf. ? Tharybis. Voir spp. inc. sedis | |
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Razouls C., Desreumaux N., Kouwenberg J. and de Bovée F., 2005-2025. - Biodiversity of Marine Planktonic Copepods (morphology, geographical distribution and biological data). Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed April 02, 2025] © copyright 2005-2025 Sorbonne University, CNRS
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