Species Card of Copepod
Calanoida ( Order )
    Diaptomoidea ( Superfamily )
        Tortanidae ( Family )
            Tortanus ( Genus )
                Atortus ( Sub-Genus )
Tortanus (Atortus) scaphus  Bowman, 1971   (F,M)
Syn.: Tortanus murrayi : C.B. Wilson,1 950 (p.346, figs.F,M)
Bowman, 1971 a (p.521, Descr.F,M, figs.F,M, carte); Ohtsuka, 1992 a (p.265); McKinnon, 1993 (p.1, figs.F); Chen & al., 2004 (p.134, tab.I: Rem.)
Species Tortanus (Atortus) scaphus - Plate 1 of morphological figuresissued from : T.E. Bowman in Pacif. Sc., 1971, XXV (4). [p.522, Fig.1].
Female: a, posterior prosome and urosome (dorsal); b, idem (ventral); c, idem (lateral); d, A2 (only four of the six apical setae of endopod shown); e, right Md (masticatory edge; outer surface); f, Md (mandibular palp); g, Mx1 (medial); h, Mx2 (medial); i, Mxp (medial).

Nota: Genital segment symmetrical, without processes. Anal segment fused with caudal rami; excavation on left side at level of anal operculum. left caudal ramus slightly wider than right. A1 reacing slightly beyond caudal rami. Two patches of fine surface hairs on anterior surface of 2nd endopod segment of legs 2-4.

Male: j, right A1; k, A1 (juncture of segments 17 and 18); l, A1 (juncture of segments 18 and 19-21); m, A1 (spinous process of segment 19-21).

Species Tortanus (Atortus) scaphus - Plate 2 of morphological figuresissued from : T.E. Bowman in Pacif. Sc., 1971, XXV (4). [p.523, Fig.2, a-h].
Female: b, P1 (anterior); c, P2 (anterior); d, P3 (anterior).

Male: a, posterior prosome and urosome (dorsal); e, P4 (posterior); f, P5 (anterior); g, right P5 (anterior); idem (posterior).

Species Tortanus (Atortus) scaphus - Plate 3 of morphological figuresissued from : T.E. Bowman in Pacif. Sc., 1971, XXV (4). [p.524, Fig.3, a].
Female: a, P5.

Species Tortanus (Atortus) scaphus - Plate 4 of morphological figuresissued from : A.D. McKinnon in Proc. R. Soc. Qd, 1993, 103. [p.2, Fig.1].
Female (from Malakal Harbour, Koror, Palau Islands): a urosome (dorsal; b, idem (lateral); c, oblque view of left caudal ramus.
Nota: Anal somite fused with the caudal rami; asymmetrical caudal rami broad, spatulate.Left caudal ramus with large excavation on outer margin at about mid-length. Ratio of length : width (at base) of each caudal ramus 3.4.
variant Female (from Davies Reef lagoon, Great Barrier Reef): d, urosome (dorsal); e, idem (lateral).
Nota: Caudal rami fused with anal somite, asymmetrical, that on the left slightly larger than that on the right, and with a more spatulate terminal portion; ratio of length : width (at base) of left caudal ramus 5.4, of right caudal ramus 6.0.

Species Tortanus (Atortus) scaphus - Plate 5 of morphological figuresissued from : C.B. Wilson in Bull. U.S. natn Mus., 1950, 100, 14 (4). [Pl.18, Figs.235-242].
As Tortanus murrayi.
Female (from Philippines): 235, habitus (dorsal); 237, A2; 238, Md; 239, Mx1; 240, Mx2; 241, P5.

Male: 236, habitus (dorsal); 242, P5.

Compl. Ref.:
Chen Q., 1983 (p.133); Shih & Young, 1995 (p.74)
NZ: 3

Distribution map of Tortanus (Atortus) scaphus by geographical zones
Species Tortanus (Atortus) scaphus - Distribution map 2issued from : S. Ohtsuka & K. Kimoto in J. Crustacean Biol., 1989, 9 (3). [p.405, Fig.11].
Distribution of several species of the subgenus Atortus. Localties: 1, Kuchinoerabu Island (South Japan); 2, Ryukyu Islands; 3, Hong Kong; 4, South China Sea between the Zhongsha Is. and Xisha Is.); 5, Malakal Harbor (Koror, Palau); 6, Makin Lagoon (Gilbert Islans); 7, Madang Harbor (New Guinea); 8, Viet-Nam; 9, Pulau Tiga (Sabah); 10, Celebes Sea; 11, 12, Banda Sea; 13, Bay of Kankamaraan (Kangeang); 14, Kuwait; 15, Ruweis (R.U.A.); 16, Assab (South Red Sea); 17, Swartkops estuary (South Africa); 18, Langebaan Lagoon (South Africa); 19, South Yemen.
Species Tortanus (Atortus) scaphus - Distribution map 3issued from : T.E. Bowman in Pacif. Sc., 1971, XXV (4). [p.528, Fig.5, a].
Locality records of three species of the subgenus Atortus
W Pacif. tropical ( Gilbert Is., Palau Is.), Australia (Great Barrier), China Seas (South China Sea, off S Hong Kong)
N: 6
(137) F: 2,25; M: 2,15; (647) F: 2,8-2,6; M: 2,4-2,2; (860) F: 2,56; {F: 2,25-2,80; M: 2,15-2,40}
Rem.: Tortanus (Atortus) scaphus variant
Ref.: McKinnon, 1993 (p.1, figs.F, Rem.)
Loc.: Great Barrier Reef (Davies Reef)
Lg: (860) F: 2,62; {F: 2,62}
Nota: Frequently seen associated with aggregations of Acartia australis attracted to torchlihgt when diving at night in the lagoon of Davies Reef.
Last update : 01/02/2015
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