Species Card of Copepod
Cyclopoida ( Order )
    Corycaeidae ( Family )
        Corycaeus ( Genus )
            Ditrichocorycaeus ( Sub-Genus )
Corycaeus (Ditrichocorycaeus) asiaticus  F. Dahl, 1894   (F,M)
Syn.: Corycaeus murrayi Farran, 1911 a (p.294, figs.F);
Ditrichocorycaeus asiaticus : Dessier, 1983 (p.89, Tableau 1, Rem., %); 1988 (tabl.1); Boxshall & Halsey, 2004 (p.495); McKinnon & al., 2008 (p:843: Tab.1); Zakaria & al., 2016 (p.1, Table 1, Rem.)
Ref.:
F. Dahl, 1894 (p.71); M. Dahl, 1912 (p.74, figs.F,M); Früchtl, 1924 b (p.101, Rem.); Gurney, 1927 (p.163, figs.F, Rem.); Farran, 1936 a (p.137); Mori, 1937 (1964) (p.136, figs.F); Dakin & Colefax, 1940 (p.115, figs.M); Sewell, 1947 (p.281, fig.F); Tanaka, 1957 (p.87, figs.F,M); 1960 (p.79, figs.M); Motoda, 1963 (p.237, figs.M); Tanaka, 1964 (p.15); Chen & al., 1974 (p.60, figs.F,M, p.76: Rem.); Bradford, 1978 (p.138: Rem.); Zheng & al., 1982 (p.146, figs.F); Chihara & Murano, 1997 (p.963, Pl.212: F,M); Conway & al., 2003 (p.253, figs.F,M, Rem.);
Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 1 of morphological figuresissued from : O. Tanaka in J. Fac. Agricult. Kyushu Univ., 1957, 11 (1). [Pl.6, Figs.13-19].
Female (from Japanese waters): 13, habitus (dorsal); 14, urosome (lateral right side); 15, A2; 16, apical spine of exopod of P2; 17, P4
Nota: Anterior segment 1.6 times as long as wide. The abdominal segments and furca in the proportional lengths 42:29:29. Genital segment about as long as wide (20:28), without ventral hook. Anal segment about as long as it is wide at the proximal (12:8). Furcal rami 5 times as long as wide (14:3).

Male: 18, habitus (dorsal); 19, urosome (lateral right side).
Nota: Wing-like expansions of 3rd thoracic segment extend posteriorly about to the middle of genital segment. The abdominal segments and furca in the proportional lengths 49:24:27. Genital segment 1.3 times as long as wide (23:18). Furcal rami about 6 times as long as it is wide at the middle of the ramus (25:4). Ventral hook of genital segment shows some individual difference.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 2 of morphological figuresissued from : R.B.S. Sewell in The John Murray Expedition, 1933-34, Scientific Reports, VIII (1), 1947. [p.280, Fig.70, D].
Male (from S Arabian coast): D, A2.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 3 of morphological figuresissued from : Q.-c Chen & S.-z. Zhang & C.-s. Zhu in Studia Marina Sinica, 1974, 9. [Pl.19, Figs.8-15].
Female (from E China Sea): 8, habitus (dorsal); 9, urosome (lateral left side); 10, A2; 11, terminal spines of exopod of P2; 12, P4.

Male: 13, habitus (dorsal); 14, urosome (lateral left side); 15, A2.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 4 of morphological figuresissued from: M. Dahl in Die Copepoden der Plankton-Expedition. I. Die Corycaeinen. Ergebnisse der Plankton-Expedition der Humboldt-Stiftung, 1912; [Taf.XI, Figs.1, 2].
Female: 1, habitus (dorsal); 2, idem (lateral).


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 5 of morphological figuresissued from: M. Dahl in Die Copepoden der Plankton-Expedition. I. Die Corycaeinen. Ergebnisse der Plankton-Expedition der Humboldt-Stiftung, 1912; [Taf.XI, Fig.5].
Female: 5, A2.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 6 of morphological figuresissued from: M. Dahl in Die Copepoden der Plankton-Expedition. I. Die Corycaeinen. Ergebnisse der Plankton-Expedition der Humboldt-Stiftung, 1912; [Taf.XI, Fig.8].
Female: 8, P2.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 7 of morphological figuresissued from: M. Dahl in Die Copepoden der Plankton-Expedition. I. Die Corycaeinen. Ergebnisse der Plankton-Expedition der Humboldt-Stiftung, 1912; [Taf.XI, Figs.3, 4].
Male: 3, habitus (dorsal); 4, idem (lateral).


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 8 of morphological figuresissued from: M. Dahl in Die Copepoden der Plankton-Expedition. I. Die Corycaeinen. Ergebnisse der Plankton-Expedition der Humboldt-Stiftung, 1912; [Taf.XI, Fig.6].
Male: 6, A2.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 9 of morphological figuresissued from: M. Dahl in Die Copepoden der Plankton-Expedition. I. Die Corycaeinen. Ergebnisse der Plankton-Expedition der Humboldt-Stiftung, 1912; [Taf.XI, Figs.7, 9].
Male: 7, P1; 9, P4.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 10 of morphological figuresissued from : G.P. Farran in Proc. zool. Soc. Lond., 1911. [Plate XIII, Figs.1-6]. As Corycaeus murrayi.
Female (from Christmas Island, Indian): 1-2, habitus (dorsal and lateral, respectively); 3, 3rd and 4th thoracic segments (mounted); 4, A2; 5, P4; 6, P2.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 11 of morphological figuresissued from : Z. Zheng, S. Li, S.J. Li & B. Chen inMarine planktonic copepods in Chinese waters. Shanghai Sc. Techn. Press, 1982 [p.147, Fig.92].
Female: a, habitus (dorsal); b, urosome (lateral, left side); c, A2; d, distal spines of exopod of P2; e, P4.

Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 12 of morphological figuresissued from : T. Mori in The Pelagic copepoda from the neighbouring waters of Japan, 1937 (1964). [Pl.75, Figs.6-8]. .
Female: 6, habitus (dorsal); 7, P4; 8, A2.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 13 of morphological figuresIssued from : O. Tanaka in Spec. Publs. Seto mar. biol. Lab., 10, 1960 [Pl. XXXIV, 3-4].
Male (from South China Sea and Indian Ocean): 3-4, last thoracic segment and abdomen (dorsal and lateral, respectively).

Nota: Proportional lengths of the cephalothorax and abdomen 60 to 40.
4th toracic margin produced posteriorly into a very acute process on each side.
Abdominal segments and caudal rami in the proportional lengths 50 : 23 : 27 = 100.
Genital segment broadish oval, 1.3 times as long as broad. The cylindrical part short, about 1/4 the length of the posterior border of the segment (3 : 13). Ventral hook slender and long, directs downwards.
Anal segment 1.6 times as long as broad; lateral margins parallel.
Caudal rami 4.8 times as long as it is broad at the proximal.


Species Corycaeus (Ditrichocorycaeus) asiaticus - Plate 14 of morphological figuresIssued from : R. Gurney in Trans. zool. Soc. London, 1927, 22. [p.162, Fig.23].
Female (Port Taufiq): A, habitus (dorsal); B, last thoracic segment and abdomen (dorsal); C, exopodite 3 of P2.

Proportion in male: furca, anal and genital somite 18: 14: 36, respectively. Genital somite with ventral hook. endopod of P4 with 2 setae. Terminal spine of P2 straight.

Compl. Ref.:
Sewell, 1948 (p.420, 434); Yamazi, 1958 (p.155, Rem.); De Decker, 1964 (p.15, 21, 28); De Decker & Mombeck, 1964 (p.12); Furuhashi, 1966 a (p.295, vertical distribution in Kuroshio region, Table 10); Palet, 1975 (p.660); Marques, 1982 (p.776); Guangshan & Honglin, 1984 (p.118, tab.); Othman & al., 1990 (p.561, 564, Table 1); Yoo, 1991 (tab.1); Kim al., 1993 (p.270); Shih & Young, 1995 (p.76); Go & al., 1997 (tab.1); Hsieh & Chiu, 1998 (tab.2); Noda & al., 1998 (p.55, Table 3, occurrence); Lo & al., 2001 (1139, tab.I); Hwang & al., 2003 (p.193, tab.2); Rezai & al., 2004 (p.490, tab.2); Lan & al., 2004 (p.332, tab.1); Lan & al., 2004 (p.332, tab.1, tab.2); Lo & al.*, 2004 (p.218, fig.6); Lo & al., 2004 (p.89, tab.1); Hwang & al., 2006 (p.943, tabl. I); Jitlang & al., 2008 (p.65, Table 1); Fernandes, 2008 (p.465, Tabl.2); Lan Y.-C. & al., 2008 (p.61, Table 1, % vs stations); C.-Y. Lee & al., 2009 (p.151, Tab.2); Fazeli & al., 2010 (p.153, Table 1); Hsiao S.H. & al., 2011 (p.475, Appendix I); Hsiao & al., 2011 (p.317, Table 2, indicator of seasonal change); Maiphae & Sa-ardrit, 2011 (p.641, Table 2, 3, Rem.); Tseng & al., 2012 (p.621, Table 3: abundance); Rojas-Herrera & al., 2016 (p.40, Table 2: temporal abundance)
NZ: 11

Distribution map of Corycaeus (Ditrichocorycaeus) asiaticus by geographical zones
Loc:
South Africa (off Cape of Good Hope, SE), E Medit. (W Egyptian coast), Suez Canal (KM. 152), Port Taufiq, Suez Bay, Red Sea, Gulf of Oman, Arabian coast, off Zanzibar, Arabian Sea, Madagascar (Nosy Bé), Rodrigues Is., Seychelles Is., Indian, NW India (Saurashtra coast), Bay of Bengal, Christmas Is., Straits of Malacca, G. of Thailand, Indonesia-Malaysia, Banda Straits, China Seas (Yellow Sea, East China Sea, South China Sea),Taiwan Strait, Taiwan (E, W, SW, NW, N, NE, Mienhua Canyon), Okinawa, S Korea, Japan Sea, Japan, Kuchinoerabu Is., Tanabe Bay, Pacif. (W equatorial), Australia (Great Barrier, New South Wales, North West Cape, G. of Carpentaria), New Caledonia, off N Hawaii, Pacif. (E equatorial), Acapulco Bay (rare)
N: 60
Lg.:
(34) F: 1,38-1,26; M: 1,26-1,16; (66) M: 1,22-1,02; (91) F: ± 1,2; (104) M: 1,1; (107) F: 1,19-1,15; M: 1,12-1,04; (109) F: 1,22-1,17; M: 1,1-1,07; (114) F: 1,4; (178) M: 0,98; (332) F: 1,05; M: 0,96; (666) F: 1,2-1,14; M: 1,01-0,97; (667) M: 1,1; (669) F: 1,2; (786) M: 1,15-1,07; (991) F: 1,15-1,2; M: 1,04-1,12; (1023) F: 1,17-1,26; {F: 1,05-1,40; M: 0,96-1,26}
Rem.: Epipelagic.
Corycaeus ( Ditrichocorycaeus) asiaticus mertoni Früchtl,1923 (F)
Ref.: Früchtl, 1923 (p.450, 453, fig.F); 1924 b (p.100, fig.F); Sewell, 1948 (p.377)
Loc.: Is. Aru
Lg.: (625) F: 1,12

. Corycaeus (Ditrichocorycaeus) asiaticus murrayi Farran, 1911 (F)
Syn.: Corycaeus murrayi Farran, 1911 a (p.294, figs.F)
Ref.: Früchtl, 1924 b (p.101, Rem.)
Loc.: Indian (I. Christmas)
Lg.: (669) F: 1,2.

See in DVP Conway & al., 2003 (version 1)
Zakaria & al. (2016) note for the first time this species in the East Mediterranean Sea, probably a lesseptian species.

For Boxshall & Halsey (2004, p.491) the subgenus (Dahl, 1912) is considered as full generic status pending phylogenetic revision of the whole family at the generic level.
Last update : 18/04/2020
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