Species Card of Copepod
Cyclopoida ( Order )
    incertae sedis (Cyclopoida) ( Family )
        Pachos ( Genus )
Pachos punctatum  (Claus, 1863)   (F,M)
Syn.: Pachysoma punctata Claus, 1863 (p.163, Descr.M, figs.M);
Pachysoma punctatum : ? Brady, 1883 (p.121); Giesbrecht, 1892 (p.612, 614, 774, Descr.M, figs.F,M); A. Scott, 1909 (p.262, Rem.); Ganapati & Shanthakumari, 1962 (p.10, 16); Fish, 1962 (p.23); Humes & Stock, 1973 (p.329); Chen & al., 1974 (p.44, figs.F,M); Zheng & al., 1982 (p.109, figs.F,M); Zheng Zhong & al., 1984 (1989) (p.263, figs.F,M); Guangshan & Honglin, 1984 (p.118, tab.); Shih & Young, 1995 (p.77); Noda & al., 1998 (p.55, Table 3, occurrence); W.-B. Chang & al., 2010 (p.735, Table 2)
Ref.:
Pesta, 1920 (p.653, fig.F); Rose, 1933 a (p.306, figs.F,M); Owre & Foyo, 1967 (p.114, figs.F,M); Vives, 1972 (p.232, figs. juv.F); Boxshall, 1977 a (p.149, figs.F,M); Björnberg & al., 1981 (p.668, figs.F,M); Chihara & Murano, 1997 (p.978, Pl.220: F,M); Boxshall & Halsey, 2004 (p.620, p.621: figs.F,M); Vives & Shmeleva, 2010 (p.398, figs.F,M, Rem.)
Species Pachos punctatum - Plate 1 of morphological figuresissued from : Q.-c Chen & S.-z. Zhang & C.-s. Zhu in Studia Marina Sinica, 1974, 9. [Pl.9, Figs.1-3].
Male (from China Seas): 1, habitus (dorsal); 2, A1; 3, P4.


Species Pachos punctatum - Plate 2 of morphological figuresissued from : Q.-c Chen & S.-z. Zhang & C.-s. Zhu in Studia Marina Sinica, 1974, 9. [Pl.8, Figs.11-13].
Female: 11, urosome (dorsal); 12, Mxp; 13, P4.


Species Pachos punctatum - Plate 3 of morphological figuresissued from : G.A. Boxshall in Bull. Br. Mus. nat. Hist. (Zool.), 1977, 31 (3). [p.150, Fig.24].
Female (from 18°N, 25°W): a, urosome (ventral); b, A1 (ventral; segmentation only); c, A2 (anterior); d, Md (anterior); e, Mx1 (anterior); f, Mx2 (anterior); g, Mxp (antero-ventral); h-k, P1 to P4 (posterior).
Nota: Body globular, with inflated prosome and short urosome; surface of carapace covered with minute denticles; a pair of sensory setules present on carapace and each of remainning 3 prosome somites. Rostrum prominent. caudal rami about 2.2 times longer than broad. Md produced into a slender lash with wide base; armature reduced to a single denticle on the outer margin and a short row of fine bristles near the inner margin. Mx1 bilobed. Mx2 2-segmented; terminal segment forming a curved element distally and with a proximal spine. Mxp 3-segmented, comprising an unarmed basal segment, a robust middle segment with 2 stout denticulate setae, and a claw-like terminal segment bearing a short seta proximally. P5 a short conical process with 2 apical setae and a lateral seta located near its base. P6 represented by 2 long plumose setae situated on the lateral margin of the genital complex.


Species Pachos punctatum - Plate 4 of morphological figuresissued from : G.A. Boxshall in Bull. Br. Mus. nat. Hist. (Zool.), 1977, 31 (3). [p.152, Fig.25].
Male: a, habitus (dorsal); b, A1 (ventral); c, urosome (ventral); d, Mxp (anterior).
Nota: Urosome with 3 somites posterior to genital complex. Minute denticles and sensory setules present on carapce and prosome as in female. A2 differing from that of female in having only 3 apical setae on the terminal segment instead of 4. Mxp 4-segmented; comprising a broad basal segment, a 2nd segment with an inner proximal swelling armed with 2 setae and covered with fine spinules, a small unarmed 3rd segment and a 4th segment forming a curved claw with a spatulate tip and armed with 2 unequal setae at its base.


Species Pachos punctatum - Plate 5 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.28-30, 32, 35]. As Pachysoma punctatum.
Female: 28, Md; 29, Mxp; 30, Mx2; 32, urosome (dorsal); 35, habitus (dorsal).


Species Pachos punctatum - Plate 6 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.23-26]. As Pachysoma punctatum.
Female: 23, P4 (posterior view); 24, P1 (anterior view); 25, A2; 26, Mx1.


Species Pachos punctatum - Plate 7 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Fig.36]. As Pachysoma punctatum
Female: endopod segment 3 of P3.


Species Pachos punctatum - Plate 8 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.22, 27]. As Pachysoma punctatum.
Male: 22, A1 (ventral view); 27, Mxp.


Species Pachos punctatum - Plate 9 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.31, 33-34]. As Pachysoma punctatum.
Male: 31, genital segment (ventral); 33, urosome (dorsal); 34, habitus (dorsal).


Species Pachos punctatum - Plate 10 of morphological figuresIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. - Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.38, 39]. As Pachysoma punctatum
Female: 38, habitus (lateral); 39, endopod segment 3 of P2.


Species Pachos punctatum - Plate 11 of morphological figuresissued from : Z. Zheng, S. Li, S.J. Li & B. Chen in Marine planktonic copepods in Chinese waters. Shanghai Sc. Techn. Press, 1982 [p.110, Fig.67-1]. As Pachysoma punctatum.
Female; a, habitus (dorsal); b, urosome (dorsal); c, A1; d, A2; e, Mxp; f, P1; g, P2; h, endopod of P3; i, P4.
Scale bar in mm.


Species Pachos punctatum - Plate 12 of morphological figuresissued from : Z. Zheng, S. Li, S.J. Li & B. Chen in Marine planktonic copepods in Chinese waters. Shanghai Sc. Techn. Press, 1982 [p.111, Fig.67-2]. As Pachysoma punctatum.
Male: j, habitus (dorsal); k, urosome (dorsal); l, A2; m, distal part of A2; n, Mxp; o, P4.
Scale bar in mm.

Compl. Ref.:
Rose & Vaissiere, 1952 (1953 a) (p.174); Wilson, 1942 a (p.200); 1950 (p.274); Fagetti, 1962 (p.46); Grice & Hart, 1962 (p.287, table 3, p.293: Rem.); V.N. Greze, 1963 a (tabl.2); Mazza, 1966 (p.73); Deevey, 1971 (p.224); Apostolopoulou, 1972 (p.329, 380); Boxshall, 1977 b (p.552); Deevey & Brooks, 1977 (p.156, tab.2, Station "S"); Dessier, 1979 (p.207); Vaissière & Séguin, 1980 (p.23, tab.2); Vives, 1982 (p.296); Kovalev & Shmeleva, 1982 (p.86); Scotto di Carlo & al., 1984 (p.1045); Regner, 1985 (p.11, Rem.: p.42); Hure & Krsinic, 1998 (p.83, 104); Gilabert & Moreno, 1998 (tab.1, 2); Suarez-Morales & Gasca, 1998 a (p.112); Fernandez-Alamo & al., 2000 (p.1139, Appendix); El-Sherif & Aboul Ezz, 2000 (p.61, Table 3: occurrence); Seridji & Hafferssas, 2000 (tab.1); Lopez-Salgado & al., 2000 (tab.1); Osore & al., 2003 (p.69); Vukanic, 2003 (p.139, tab.1); Khelifi-Touhami & al., 2007 (p.327, Table 1); Mazzocchi & Di Capua, 2010 (p.429); Tutasi & al., 2011 (p.791, Table 2); Andersen N.G. & al., 2011 (p.71, Fig.3: abundance); Hsiao S.H. & al., 2011 (p.475, Appendix I); Tseng L.-C. & al., 2011 (p.47, Table 2, occurrences vs mesh sizes); Uysal & Shmeleva, 2012 (p.909, Table I); in CalCOFI regional list (MDO, Nov. 2013; M. Ohman, comm. pers.); Zakaria & al., 2016 (p.1, Table 1) ; Benedetti & al., 2016 (p.159, Table I, fig.1, functional characters); Marques-Rojas & Zoppi de Roa, 2017 (p.495, Table 1).
NZ: 14

Distribution map of Pachos punctatum by geographical zones
Loc:
Congo, Ghana, Cape Verde Is., off Madeira, S Brazil, off Amazon, Barbados Is., Caribbean Sea, Bahia de Mochima (Venezuela), G. of Mexico, Cuba, Florida, Chesapeake Bay, Sargasso Sea, off Bermuda: Station ‘’ S’’ (32°10’N, 64°30’W), Medit. (Algiers Bay, Gulf of Annaba, Baleares, W Basin, Ligurian Sea, Tyrrhenian Sea, Messina, N & S Adriatic Sea, Ionian Sea, Aegean Sea, W Egyptian coast, Lebanon Basin), Safaga, Red Sea, Kennya, Madagascar (Nosy Bé), India (Lawson's Bay), Indonesia-Malaysia, Philippines, China Seas (East China Sea, South China Sea), Taiwan (S, E: Kuroshio Current), S Japan (Kuchonoerabu Is.), Japan (Honshu), Aleutian Is. (in C.B. Wilson, 1950), W Baja California, G. of Tehuantepec, Central America, Galapagos-Ecuador, off Peru, off Chile
N: 65
Lg.:
(46) F: 2,2-1,9; M: 2-1,87; (109) F: 2,3-2; M: 2,1-1,9; (139) F: 2,25; M: 2,2 ; (449) F: 2,2-1,9; (785) F: 2,64-2,4; (1023) F: 1,83; M: 1,84; {F: 1,83-2,64; M: 1,87-2,20}
Rem.: epiplagic. Overall Depth Range in Sargasso Sea: 0-1000 m (Deevey & Brooks, 1977, Station "S");
Boxshall (1977 a, p.153) points to the segmentation of A1 appears to be rather variable. Claus (1863) recorded 8 segments in the female whereas Giesbrecht (1892) found 5 in the female and 7 in the male. This apparent variation is probably due to the difficulty of determining the precise segmentation of the densely setate proximal portion of A1. Seven distinct segments were observed in both sexes although some degree of fusion was observed between segments 3 and 4 in the female.
Last update : 20/02/2018
    to add your remark
  

 Any use of this site for a publication will be mentioned with the following reference :

Razouls C., de Bovée F., Kouwenberg J. and Desreumaux N., 2005-2020. - Diversity and Geographic Distribution of Marine Planktonic Copepods. Sorbonne University, CNRS. Available at http://copepodes.obs-banyuls.fr/en [Accessed October 20, 2020]

© copyright 2005-2020 Sorbonne University, CNRS

Webmaster
CNRS   Observatoire Océanologique de Banyuls sur Mer - Laboratoire Arago
Sorbonne Université

 

Version française
English version

 

Search

On the WEB of CNRS

 


Marine Planktonic Copepods

Marine Planktonic Copepods

 

Imprimer Contact Accueil Plan du site Accès restreint Retour Une du Labo Imprimer Contact Plan du site Crédits Téléchargez les Plug-Ins