Fiche d'espèce de Copépode
Cyclopoida ( Ordre )
    incertae sedis (Cyclopoida) ( Famille )
        Pachos ( Genre )
Pachos punctatum  (Claus, 1863)   (F,M)
Syn.: Pachysoma punctata Claus, 1863 (p.163, Descr.M, figs.M);
Pachysoma punctatum : ? Brady, 1883 (p.121); Giesbrecht, 1892 (p.612, 614, 774, Descr.M, figs.F,M); A. Scott, 1909 (p.262, Rem.); Ganapati & Shanthakumari, 1962 (p.10, 16); Fish, 1962 (p.23); Humes & Stock, 1973 (p.329); Chen & al., 1974 (p.44, figs.F,M); Zheng & al., 1982 (p.109, figs.F,M); Zheng Zhong & al., 1984 (1989) (p.263, figs.F,M); Guangshan & Honglin, 1984 (p.118, tab.); Shih & Young, 1995 (p.77); Noda & al., 1998 (p.55, Table 3, occurrence); W.-B. Chang & al., 2010 (p.735, Table 2)
Ref.:
Pesta, 1920 (p.653, fig.F); Rose, 1933 a (p.306, figs.F,M); Owre & Foyo, 1967 (p.114, figs.F,M); Vives, 1972 (p.232, figs. juv.F); Boxshall, 1977 a (p.149, figs.F,M); Björnberg & al., 1981 (p.668, figs.F,M); Chihara & Murano, 1997 (p.978, Pl.220: F,M); Boxshall & Halsey, 2004 (p.620, p.621: figs.F,M); Vives & Shmeleva, 2010 (p.398, figs.F,M, Rem.)
Espèce Pachos punctatum - Planche 1 de figures morphologiquesissued from : Q.-c Chen & S.-z. Zhang & C.-s. Zhu in Studia Marina Sinica, 1974, 9. [Pl.9, Figs.1-3].
Male (from China Seas): 1, habitus (dorsal); 2, A1; 3, P4.


Espèce Pachos punctatum - Planche 2 de figures morphologiquesissued from : Q.-c Chen & S.-z. Zhang & C.-s. Zhu in Studia Marina Sinica, 1974, 9. [Pl.8, Figs.11-13].
Female: 11, urosome (dorsal); 12, Mxp; 13, P4.


Espèce Pachos punctatum - Planche 3 de figures morphologiquesissued from : G.A. Boxshall in Bull. Br. Mus. nat. Hist. (Zool.), 1977, 31 (3). [p.150, Fig.24].
Female (from 18°N, 25°W): a, urosome (ventral); b, A1 (ventral; segmentation only); c, A2 (anterior); d, Md (anterior); e, Mx1 (anterior); f, Mx2 (anterior); g, Mxp (antero-ventral); h-k, P1 to P4 (posterior).
Nota: Body globular, with inflated prosome and short urosome; surface of carapace covered with minute denticles; a pair of sensory setules present on carapace and each of remainning 3 prosome somites. Rostrum prominent. caudal rami about 2.2 times longer than broad. Md produced into a slender lash with wide base; armature reduced to a single denticle on the outer margin and a short row of fine bristles near the inner margin. Mx1 bilobed. Mx2 2-segmented; terminal segment forming a curved element distally and with a proximal spine. Mxp 3-segmented, comprising an unarmed basal segment, a robust middle segment with 2 stout denticulate setae, and a claw-like terminal segment bearing a short seta proximally. P5 a short conical process with 2 apical setae and a lateral seta located near its base. P6 represented by 2 long plumose setae situated on the lateral margin of the genital complex.


Espèce Pachos punctatum - Planche 4 de figures morphologiquesissued from : G.A. Boxshall in Bull. Br. Mus. nat. Hist. (Zool.), 1977, 31 (3). [p.152, Fig.25].
Male: a, habitus (dorsal); b, A1 (ventral); c, urosome (ventral); d, Mxp (anterior).
Nota: Urosome with 3 somites posterior to genital complex. Minute denticles and sensory setules present on carapce and prosome as in female. A2 differing from that of female in having only 3 apical setae on the terminal segment instead of 4. Mxp 4-segmented; comprising a broad basal segment, a 2nd segment with an inner proximal swelling armed with 2 setae and covered with fine spinules, a small unarmed 3rd segment and a 4th segment forming a curved claw with a spatulate tip and armed with 2 unequal setae at its base.


Espèce Pachos punctatum - Planche 5 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.28-30, 32, 35]. As Pachysoma punctatum.
Female: 28, Md; 29, Mxp; 30, Mx2; 32, urosome (dorsal); 35, habitus (dorsal).


Espèce Pachos punctatum - Planche 6 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.23-26]. As Pachysoma punctatum.
Female: 23, P4 (posterior view); 24, P1 (anterior view); 25, A2; 26, Mx1.


Espèce Pachos punctatum - Planche 7 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Fig.36]. As Pachysoma punctatum
Female: endopod segment 3 of P3.


Espèce Pachos punctatum - Planche 8 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.22, 27]. As Pachysoma punctatum.
Male: 22, A1 (ventral view); 27, Mxp.


Espèce Pachos punctatum - Planche 9 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.31, 33-34]. As Pachysoma punctatum.
Male: 31, genital segment (ventral); 33, urosome (dorsal); 34, habitus (dorsal).


Espèce Pachos punctatum - Planche 10 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. - Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.48, Figs.38, 39]. As Pachysoma punctatum
Female: 38, habitus (lateral); 39, endopod segment 3 of P2.


Espèce Pachos punctatum - Planche 11 de figures morphologiquesissued from : Z. Zheng, S. Li, S.J. Li & B. Chen in Marine planktonic copepods in Chinese waters. Shanghai Sc. Techn. Press, 1982 [p.110, Fig.67-1]. As Pachysoma punctatum.
Female; a, habitus (dorsal); b, urosome (dorsal); c, A1; d, A2; e, Mxp; f, P1; g, P2; h, endopod of P3; i, P4.
Scale bar in mm.


Espèce Pachos punctatum - Planche 12 de figures morphologiquesissued from : Z. Zheng, S. Li, S.J. Li & B. Chen in Marine planktonic copepods in Chinese waters. Shanghai Sc. Techn. Press, 1982 [p.111, Fig.67-2]. As Pachysoma punctatum.
Male: j, habitus (dorsal); k, urosome (dorsal); l, A2; m, distal part of A2; n, Mxp; o, P4.
Scale bar in mm.

Ref. compl.:
Rose & Vaissiere, 1952 (1953 a) (p.174); Wilson, 1942 a (p.200); 1950 (p.274); Fagetti, 1962 (p.46); Grice & Hart, 1962 (p.287, table 3, p.293: Rem.); V.N. Greze, 1963 a (tabl.2); Mazza, 1966 (p.73); Deevey, 1971 (p.224); Apostolopoulou, 1972 (p.329, 380); Boxshall, 1977 b (p.552); Deevey & Brooks, 1977 (p.156, tab.2, Station "S"); Dessier, 1979 (p.207); Vaissière & Séguin, 1980 (p.23, tab.2); Vives, 1982 (p.296); Kovalev & Shmeleva, 1982 (p.86); Scotto di Carlo & al., 1984 (p.1045); Regner, 1985 (p.11, Rem.: p.42); Hure & Krsinic, 1998 (p.83, 104); Gilabert & Moreno, 1998 (tab.1, 2); Suarez-Morales & Gasca, 1998 a (p.112); Fernandez-Alamo & al., 2000 (p.1139, Appendix); El-Sherif & Aboul Ezz, 2000 (p.61, Table 3: occurrence); Seridji & Hafferssas, 2000 (tab.1); Lopez-Salgado & al., 2000 (tab.1); Osore & al., 2003 (p.69); Vukanic, 2003 (p.139, tab.1); Khelifi-Touhami & al., 2007 (p.327, Table 1); Mazzocchi & Di Capua, 2010 (p.429); Tutasi & al., 2011 (p.791, Table 2); Andersen N.G. & al., 2011 (p.71, Fig.3: abundance); Hsiao S.H. & al., 2011 (p.475, Appendix I); Tseng L.-C. & al., 2011 (p.47, Table 2, occurrences vs mesh sizes); Uysal & Shmeleva, 2012 (p.909, Table I); in CalCOFI regional list (MDO, Nov. 2013; M. Ohman, comm. pers.); Zakaria & al., 2016 (p.1, Table 1) ; Benedetti & al., 2016 (p.159, Table I, fig.1, functional characters); Marques-Rojas & Zoppi de Roa, 2017 (p.495, Table 1).
NZ: 14

Carte de distribution de Pachos punctatum par zones géographiques
Loc:
Congo, Ghana, Cape Verde Is., off Madeira, S Brazil, off Amazon, Barbados Is., Caribbean Sea, Bahia de Mochima (Venezuela), G. of Mexico, Cuba, Florida, Chesapeake Bay, Sargasso Sea, off Bermuda: Station ‘’ S’’ (32°10’N, 64°30’W), Medit. (Algiers Bay, Gulf of Annaba, Baleares, W Basin, Ligurian Sea, Tyrrhenian Sea, Messina, N & S Adriatic Sea, Ionian Sea, Aegean Sea, W Egyptian coast, Lebanon Basin), Safaga, Red Sea, Kennya, Madagascar (Nosy Bé), India (Lawson's Bay), Indonesia-Malaysia, Philippines, China Seas (East China Sea, South China Sea), Taiwan (S, E: Kuroshio Current), S Japan (Kuchonoerabu Is.), Japan (Honshu), Aleutian Is. (in C.B. Wilson, 1950), W Baja California, G. of Tehuantepec, Central America, Galapagos-Ecuador, off Peru, off Chile
N: 65
Lg.:
(46) F: 2,2-1,9; M: 2-1,87; (109) F: 2,3-2; M: 2,1-1,9; (139) F: 2,25; M: 2,2 ; (449) F: 2,2-1,9; (785) F: 2,64-2,4; (1023) F: 1,83; M: 1,84; {F: 1,83-2,64; M: 1,87-2,20}
Rem.: épipélagique.
Voir aussi les remarques en anglais
Dernière mise à jour : 20/02/2018

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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2020. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Sorbonne Université, CNRS. Disponible sur http://copepodes.obs-banyuls.fr [Accédé le 28 septembre 2020]

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